# Urocyclidae

Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea.
Some species within this family make and use love darts before mating.

## Distribution

The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.

## Taxonomy

The family Urocyclidae consists of 3 subfamilies:
* subfamily Rhysotininae Schileyko, 2002
* subfamily Sheldoniinae Connolly, 1925 (1912) - synonyms: Peltatinae Godwin-Austen, 1912; Trochonanininae Connolly, 1912; Trochozonitinae Iredale, 1914; Ledoulxiinae Pilsbry, 1919; Gymnarionidae Von Mol, 1970; Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
* subfamily Urocyclinae Simroth, 1889
** tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
** tribe Dendrolimacini Van Goethem, 1977
** tribe Leptichnini Van Goethem, 1977
** tribe Upembellini Van Goethem, 1977

### Genera

The following genera are recognised in the family Urocyclidae:

#### unsorted

* † Koruella Pickford, 2009

#### Subfamily Rhysotininae

* Rhysotina Ancey, 1887

#### Subfamily Sheldoniinae

* Acantharion Binder & Tillier, 1985
* Africarion Godwin-Austen, 1883
* Amatarion Van Mol, 1970
* Angustivestis Pilsbry, 1919
* Aspidelus Morelet, 1884
* Aspidotomium Degner, 1932
* Belonarion Pilsbry, 1919
* Bloyetia Bourguignat, 1890
* Camerunarion Van Mol, 1970
* Carinazingis van Bruggen & de Winter, 1990
* Centrafricarion Van Mol, 1970
* Chlamydarion Van Mol, 1968
* Colparion Laidlaw, 1938
* Degneria Verdcourt, 1956
* Entagaricus Pilsbry, 1919
* Estria Poirier, 1887
* Falloonella Preston, 1914
* Fuellebornia Verdcourt, 1998
* Granularion Germain, 1912
* Gymnarion Pilsbry, 1919
* †Hamya Bourguignat, 1885
* Kerkophorus Godwin-Austen, 1912
* Lacrimarion Connolly, 1939
* Malagarion Tillier, 1979
* Mesafricarion Pilsbry, 1919
* Microkerkus Godwin-Austen, 1912
* Montanobloyetia Verdcourt, 1961
* Morrumbalia Verdcourt, 1998
* Paragranularion Van Mol, 1970
* Percivalia Preston, 1914
* Plicatonanina Verdcourt, 1961
* Principicochlea D. Holyoak & G. Holyoak, 2020
* Principitrochoidea D. Holyoak & G. Holyoak, 2020
* Ptilototheca Herbert, 2016
* Rhopalogonium Degner, 1932
* Selatodryas Herbert, 2017
* Senegalarion Van Mol, 1970
* Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae
* Sjostedtina Verdcourt, 1961
* Sylvarion Van Mol, 1970
* Thapsia Albers, 1860
* Thielarion Van Mol, 1970
* Thomithapsia D. Holyoak & G. Holyoak, 2020
* Thomitrochoidea D. Holyoak & G. Holyoak, 2020
* Tresia Van Goethem, 1975
* Trochonanina Mousson, 1869
* Trochozonites Pfeffer, 1883
* Tropidocochlion Verdcourt, 1998
* Verdcourtia Van Mol, 1970
* Verrucarion Van Mol, 1970
* Zingis E. von Martens, 1878
* Zonitarion Pfeffer, 1883

#### Subfamily Urocyclinae

Tribe Dendrolimacini
* Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini
Tribe Leptichnini
* Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini
Tribe Upembellini
* Leptichnoides Van Goethem, 1975
* Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini
tribe Urocyclini
* Anisotoxon Van Goethem, 1975
* Atoxon Simroth, 1888
* Atoxonoides Van Goethem, 1973
* Atrichotoxon Simroth, 1910
* Buettneria Simroth, 1910
* Bukobia Simroth, 1896
* Elisolimax Cockerell, 1893
* Emphysetes Verdcourt, 2003
* Microcyclus Simroth, 1896
* Nupnus Van Goethem, 1975
* Pembatoxon Van Goethem, 1975
* Phaneroporus Simroth, 1888
* Polytoxon Simroth, 1897
* Pseudatoxon Van Goethem, 1975
* Tanzalimax Rowson, Paustian & Van Goethem, 2017
* Trichotoxon Simroth, 1888
* Udzungwalimax Rowson, Paustian & Van Goethem, 2017
* Urocyclus Gray, 1864 - the type genus of the family Urocyclidae

## Cladogram

The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade: