# Scutiger (frog)

Scutiger is a genus of toads in the family Megophryidae. Common name lazy toads has been coined for them. They occur in China, Burma, Nepal, and northern India in high-altitude habitats. Most are endemic to China.

## Evolution

A 2017 molecular phylogenetic study found that Scutiger originated in Paleo-Tibet during the Oligocene.

## Species

The following species are recognised in the genus Scutiger:
* Scutiger adungensis Dubois, 1979 – Adung lazy toad
* Scutiger bhutanensis Delorme and Dubois, 2001
* Scutiger boulengeri (Bedriaga, 1898) – Boulenger's lazy toad, Xizang alpine toad, Himalayan stream frog
* Scutiger brevipes (Liu, 1950)
* Scutiger chintingensis Liu and Hu, 1960 – Chinting lazy toad, Chinting alpine toad
* Scutiger ghunsa Khatiwada, Shu, Subedi, Wang, Ohler, Cannatella, Xie, and Jiang, 2019 – Ghunsa high altitude toad
* Scutiger glandulatus (Liu, 1950) – Hopachai lazy toad, chest gland cat-eyed toad
* Scutiger gongshanensis Yang and Su, 1979 – Gongshan lazy toad, Gongshan cat-eyed toad
* Scutiger jiulongensis Fei, Ye, and Jiang, 1995 – Jiulong cat-eyed toad
* Scutiger liupanensis Huang, 1985 – Liupan lazy toad, Liupan alpine toad
* Scutiger maculatus (Liu, 1950) – spotted lazy toad, piebald alpine toad
* Scutiger mammatus (Günther, 1896) – Tungsolo lazy toad
* Scutiger muliensis Fei and Ye, 1986 – Muli cat-eyed toad
* Scutiger nepalensis Dubois, 1974 – Nepal lazy toad, mountain pelobatid toad, Asiatic spadefoot toad
* Scutiger ningshanensis Fang, 1985 – Ningshan lazy toad, Ningshan alpine toad
* Scutiger nyingchiensis Fei, 1977 – Nyingchi lazy toad, Ladakh pelobatid toad, Nyingshi alpine toad
* Scutiger occidentalis Dubois, 1978 – Ladakh high altitude toad
* Scutiger pingwuensis Liu and Tian, 1978 – Pingwu lazy toad, Pingwu alpine toad
* Scutiger sikimmensis (Blyth, 1855) – Sikkim lazy toad, Sikkim high altitude toad, Blyth's short-limbed frog
* Scutiger spinosus Jiang, Wang, Li, and Che, 2016 – spiny lazy toad
* Scutiger tengchongensis Yang & Huang, 2019
* Scutiger tuberculatus Liu and Fei, 1979 – bumpy lazy toad, round-tubercled cat-eyed toad
* Scutiger wanglangensis Ye and Fei, 2007 – Wanglang alpine toad
* Scutiger wuguanfui Jiang, Rao, Yuan, Wang, Li, Hou, Che, and Che, 2012 – Wu's lazy toad, Medog lazy toad

## Endemic ranges

Many Scutiger species are endemic to highly restricted geographical areas in the Eastern Himalayas. The ranges often overlap with those of Oreolalax species.
* Ningxia, China
** Scutiger liupanensis: Liupanshan National Nature Reserve, Jingyuan County, Ningxia Hui Autonomous Region, China
* Shaanxi, China
** Scutiger ningshanensis: Ningshan County, Shaanxi
* Sichuan, China
** Scutiger brevipes: Dawu County, Sichuan
** Scutiger chintingensis: Mount Emei area, Sichuan
** Scutiger jiulongensis: Jiulong County, Sichuan
** Scutiger maculatus: Garze, Sichuan and Jiangda, Tibet
** Scutiger muliensis: Muli County, Sichuan
** Scutiger pingwuensis: Pingwu County, Sichuan
** Scutiger wanglangensis: Sichuan
* Yunnan, China
** Scutiger gongshanensis: Gongshan County, Yunnan
* Tibet
** Scutiger maculatus: Garze, Sichuan and Jiangda, Tibet
** Scutiger nyingchiensis: Nyingchi Prefecture, Tibet
** Scutiger spinosus: Medog County, Tibet and Tawang District, Arunachal Pradesh
** Scutiger wuguanfui: Medog County, Tibet
* Bhutan
** Scutiger bhutanensis: Bhutan
* Sikkim, India
** Scutiger sikimmensis: Sikkim, India