# Gomphonema

Gomphonema is a genus of diatoms belonging to the family Gomphonemataceae.
The genus has cosmopolitan distribution.

## Species

As accepted by WoRMS;
* Gomphonema acceptatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema acuminatum Ehrenberg, 1832
* Gomphonema adhikarii C.Radhakrishnan, Sudipta K.Das, Kociolek et B.Karthick in Radhakrishnan et al., 2020
* Gomphonema affine Kützing, 1844
* Gomphonema africanum G.S.West, 1907
* Gomphonema alavariense C. Delgado, S. Blanco, L. Gonzalez-Paz & S.F.P. Almeida in Gonzalez-Paz et al., 2020
* Gomphonema angkoricum Cejudo-Figueiras, S.Blanco & Ãlvarez-Blanco, 2012
* Gomphonema angusticapitatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema angusticlavatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema angustissimum F.Hustedt
* Gomphonema angustum C.Agardh, 1831
* Gomphonema anjae Lange-Bertalot & Reichardt
* Gomphonema asiaticum Liu & Kociolek, 2013
* Gomphonema augur Ehrenberg, 1840
* Gomphonema auguriforme Levkov, Mitic-Kopanja, Wetzel & Ector in Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema auritum A.Braun ex Kützing, 1849
* Gomphonema balatonis Pantocsek
* Gomphonema berggreni
* Gomphonema berggrenii Cleve
* Gomphonema bergii Skvortzov & Meyer
* Gomphonema bicepiformis Wei Zhang & Kociolek, 2018
* Gomphonema bigutianchnensis Y.Li in Liao & Y.Li, 2018
* Gomphonema bohemicum Reichelt & Fricke, 1902
* Gomphonema boreale E.V.Østrup
* Gomphonema brasiliensoide Metzeltin, Lange-Bertalot & García-Rodríguez, 2005
* Gomphonema bukycanyona L.N.Bukhtiyarova, 2013
* Gomphonema californicum Stancheva & Kociolek, 2016
* Gomphonema candelariae J.Frenguelli
* Gomphonema cantalicum J.-J.Brun & Héribaud-Joseph
* Gomphonema capitatum Ehrenberg, 1838
* Gomphonema capitoccultum Reichardt
* Gomphonema capsulare J.A.Ress, E.W.Thomas & Kociolek, 2016
* Gomphonema carolinense Hagelstein, 1939
* Gomphonema catarinensis G.Krasske, 1948
* Gomphonema changyangicum Li, Shi & Lei, 1999
* Gomphonema charcotii M.Peragallo
* Gomphonema chemeron Jeff.R.Stone, Westover & Kociolek
* Gomphonema chinense Liu & Kociolek, 2013
* Gomphonema clavatoides H.P.Gandhi, 1960
* Gomphonema clavatuloides Rimet, D.G. Mann, Trobajo et N. Abarca in Rimet et al., 2018
* Gomphonema clavatum Ehrenberg, 1832
* Gomphonema clerici Frenguelli, 1923
* Gomphonema clevei Fricke, 1902
* Gomphonema confusum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema correntinum Frenguelli, 1933
* Gomphonema dalatica Glushchenko, Kulikovskiy & Kociolek, 2017
* Gomphonema darwinii L.L.Bahls, 2014
* Gomphonema delicatula Skvortzov, 1937
* Gomphonema dharwarensis H.P.Gandhi, 1956
* Gomphonema dichotiforme Z.X.Shi, 2004
* Gomphonema difformum B.Karthick & J.P.Kociolek, 2011
* Gomphonema diminutum B.Karthick & J.P.Kociolek, 2011
* Gomphonema dirangense S.K.Das, C.Radhakrishnan, Kociolek & B.Karthick, 2018
* Gomphonema distincte-striatum Kociolek, J. P. & Woodward, J.
* Gomphonema dojranense Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema doonense B.Karthick, R.Nautiyal & Kociolek, 2015
* Gomphonema dubravicense Pantocsek
* Gomphonema eileencoxiae (Bahls, 2017) Bahls, C.E.Wetzel & Ector, 2018
* Gomphonema elegantissimum E.Reichardt & H.Lange-Bertalot, 2011
* Gomphonema elongatum W.Smith, 1855
* Gomphonema eriensioides Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
* Gomphonema evolutionense L.L.Bahls, 2014
* Gomphonema frequentiformis (Metzeltin & Krammer) C.E.Wetzel & F.F.Almeida in Almeida et al., 2020
* Gomphonema frickei O.Müller, 1905
* Gomphonema gallaudi Héribaud-Joseph
* Gomphonema gandhii B.Karthick & J.P.Kociolek, 2011
* Gomphonema gautieriforme D.Mitic-Kopanja, C.E.Wetzel, L.Ector & Z. Levkov, 2014
* Gomphonema genestoermeri Liu & Kociolek, 2013
* Gomphonema graciledictum E.Reichardt, 2015
* Gomphonema grande B. Karthick, Kociolek, J.C. Taylor & Cocquyt, 2016
* Gomphonema gregarium J.P Kociolek & Jeff Woodward, 2016
* Gomphonema hedinii Hustedt, 1922
* Gomphonema heideni F.Fricke
* Gomphonema helveticum Brun, 1895
* Gomphonema himalayaense Jüttner in Jüttner et al., 2018
* Gomphonema hinziae Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema hristovskii Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema hubeicum Li, Shi & Lei, 1999
* Gomphonema idsbense Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema indistinctum Kociolek, J. P. & Woodward, J.
* Gomphonema innata Skvortzov, 1937
* Gomphonema insigne Gregory, 1856
* Gomphonema insuetum Kociolek in Kociolek et al., 2014
* Gomphonema insularum Kociolek, J. P. & Woodward, J.
* Gomphonema interruptum J.YChen & H.Z.Zhu, 1994
* Gomphonema intricatoides You & Kociolek, 2015
* Gomphonema intricatum Kützing, 1844
* Gomphonema italicum Kützing, 1844
* Gomphonema jablanicense Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema jahniana Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
* Gomphonema johnsonii L.Bahls, 2013
* Gomphonema juettnerii B.Karthick, R.Nautiyal & Kociolek, 2015
* Gomphonema juriljii Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema kalahariense Jeff.R.Stone & Kociolek
* Gomphonema kallarense Pardhi et al., 2020
* Gomphonema kaznakowi K.S.Mereschkowsky
* Gomphonema kerguelenensis E.E.Manguin, 1954
* Gomphonema kezlyae Pardhi et al., 2020
* Gomphonema kociolekii R.Jahn & N.Abarca
* Gomphonema kozufense Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema lacifrigidum M.Kulikovskiy, J.P.Kociolek & Solak in Kociolek & Guiry, 2018
* Gomphonema lacusrankala Gandhi, 1958
* Gomphonema lacusrankaloides B.Karthick & J.P.Kociolek, 2012
* Gomphonema lacus-victoriense E.Reichardt, 2007
* Gomphonema lagerheimei A.Cleve, 1895
* Gomphonema lagerheimii A.Cleve, 1895
* Gomphonema lapponicum (A.Cleve) Cleve-Euler
* Gomphonema latelanceolatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema lateripunctatum Reichardt & Lange-Bertalot, 1991
* Gomphonema liebscheri E.Reichardt, 2012
* Gomphonema longiceps Ehrenberg, 1854
* Gomphonema lujanensis E.Reichardt & N.Maidana, 2008
* Gomphonema lychnidum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema magniaqua M.Kulikovskiy, J.P.Kociolek & Solak in Kociolek & Guiry, 2018
* Gomphonema magnifica H.P.Gandhi
* Gomphonema mariovense Levkov & Tofilovska in Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema martini F.Fricke
* Gomphonema matanensis D. Kapustin, Kociolek & Kulikovskiy in Kociolek et al., 2018
* Gomphonema mayamae S.K.Das, C.Radhakrishnan, Kociolek & B.Karthick, 2018
* Gomphonema megalobrebissonii D.A.Chudaev, J.P.Kociolek & M.A.Golobova, 2014
* Gomphonema meridionale J.P.Kociolek & E.W.Thomas, 2009
* Gomphonema metzeltinii You & Kociolek, 2015
* Gomphonema microcapitatum Kulikovskiy, Kociolek & Solak, 2018
* Gomphonema microlanceolatum You & Kociolek, 2015
* Gomphonema microlaticollum Kulikovskiy, Kociolek & Solak, 2018
* Gomphonema micropus Kützing, 1844
* Gomphonema minusculum Krasske
* Gomphonema minutum (C.Agardh) C.Agardh, 1831
* Gomphonema modicum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema moniliforme Gandhi, 1960
* Gomphonema mutunensis Ruwer, Ludwig & Rodrigues, 2019
* Gomphonema nagpurense Sarode & Kamat, 1984
* Gomphonema navicelloides Frenguelli, 1942
* Gomphonema naviculoides W. Smith, 1856
* Gomphonema neotropicum N.Abarca & D.Mora
* Gomphonema nesophilum Kociolek, J. P. & Woodward, J.
* Gomphonema obesoccultum Reichardt
* Gomphonema occidentale
* Gomphonema olivaceum (Lyngbye) Desmazières, 1825
* Gomphonema oregonicum Ehrenberg
* Gomphonema oxycephalum Cleve
* Gomphonema paracapitatum Kulikovskiy, Kociolek & Solak, 2018
* Gomphonema paradaphnoides Cejudo-Figueiras, S.Blanco & Ãlvarez-Blanco, 2012
* Gomphonema paraexilissimum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema parapygmaeum Jüttner & Kociolek in Jüttner et al., 2018
* Gomphonema parasundaense Jüttner in Jüttner et al., 2018
* Gomphonema paratergestinum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema parvicapitatum Kociolek in Kociolek et al., 2014
* Gomphonema parvuliforme Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema parvulum (Kützing) Kützing, 1849
* Gomphonema pelisteriense Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema preliciae Levkov, Mitic-Kopanja & E. Reichardt, 2016
* Gomphonema prespanense Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema prowsei Glushchenko, Kulikovskiy & Kociolek, 2017
* Gomphonema pseudacuminatum Kulikovskiy, Kociolek & Solak, 2015
* Gomphonema pseudaffine Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema pseudoparvulum Kociolek, J. P. & Woodward, J.
* Gomphonema puiggarianum
* Gomphonema pusillum (Grunow, 1880) Kulikovskiy & Kociolek, 2018
* Gomphonema pygmaeoides You & Kociolek, 2015
* Gomphonema qingyiensis L-X. Zhang, P. Yu & Q-M. You in Zhang et al., 2020
* Gomphonema raraense Jüttner & S.Gurung in Jüttner et al., 2018
* Gomphonema reediae Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema resendei A.R.Moura
* Gomphonema rexlowei Liu & Kociolek, 2013
* Gomphonema rhombicum Fricke, 1904
* Gomphonema rimetiana Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
* Gomphonema ristovskae Levkov & Tofilovska in Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema rostellatum (Kützing) G.L.Rabenhorst
* Gomphonema russicum Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
* Gomphonema saparense S.K.Das, C.Radhakrishnan, Kociolek & B.Karthick, 2018
* Gomphonema saravathense Gandhi, 1970
* Gomphonema scardicum D.Mitic-Kopanja, C.E.Wetzel, L.Ector & Z.Levkov, 2014
* Gomphonema semiapertum Grunow, 1880
* Gomphonema shanghaiensis Zhang & Kociolek, 2016
* Gomphonema shiwania Y.Liu, Kulikovskiy & Kociolek in Zhang et al., 2020
* Gomphonema sichuanenesis Y.Li & J.P.Kociolek, 2010
* Gomphonema sichuanensis Li & Kociolek in Li et al., 2010
* Gomphonema sierrianum Stancheva & Kociolek, 2016
* Gomphonema sphaerophorum Ehrenberg, 1845
* Gomphonema spicula Gandhi, 1958
* Gomphonema spiculoides H.P.Gandhi
* Gomphonema spirkovskae Levkov & Mitic-Kopanja in Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema stauroneiforme Grunow
* Gomphonema staurophorum (Pantocsek) Cleve-Euler, 1955
* Gomphonema subapicatum F.E.Fritsch & M.F.Rich
* Gomphonema subclavatum (Grunow) Grunow, 1884
* Gomphonema subinsigniforme L.Ge, Y.Liu & J.P.Kociolek, 2014
* Gomphonema sublaticollum Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema submalayense Gandhi, 1970
* Gomphonema subnaviculoides Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema subodhii Jüttner & S.Gurung in Jüttner et al., 2018
* Gomphonema subramosum C.A.Agardh
* Gomphonema subtile Ehrenberg, 1843
* Gomphonema svalbardense E.Pinseel, K.Kopalová & Van der Vijver, 2014
* Gomphonema tamilense B.Karthick & J.P.Kociolek, 2012
* Gomphonema tenoccultum Reichardt
* Gomphonema tenuis Gandhi, 1960
* Gomphonema tergestinum (Grunow in Van Heurck) Fricke, 1902
* Gomphonema truncatum Ehrenberg, 1832
* Gomphonema tumens Kociolek & Stoermer, 1991
* Gomphonema turgidum Ehrenberg, 1854
* Gomphonema undulans I.Ãlvarez-Blanco, C.Cejudo-Figueiras & S.Blanco, 2013
* Gomphonema validum Cleve, 1894
* Gomphonema vardarense Levkov, Mitic-Kopanja & E.Reichardt, 2016
* Gomphonema variabilis A.Jurilj, 1957
* Gomphonema varioreduncum Jüttner, Ector, E. Reichardt, Van de Vijver & E.J.Cox, 2013
* Gomphonema vastum F.Hustedt
* Gomphonema ventricosum Gregory, 1856
* Gomphonema vibrio Ehrenberg, 1843
* Gomphonema vidarbhense Sarode & Kamat, 1984
* Gomphonema williamsii Kociolek & Liu, 2013
* Gomphonema witkowskii Kociolek & Liu, 2013
* Gomphonema wuyiensis W.Zhang & Kociolek in Zhang et al., 2018
* Gomphonema xiantaoicum Z.X.Shi & N.Li in Z.X.Shi, 2014
* Gomphonema xinjiangianum You & Kociolek, 2015
* Gomphonema yangtzensis Li in Li, Gong, Xie & Shen, 2006
* Gomphonema yaominae Li, 2012
* Gomphonema yberiense Frenguelli, 1933
* Gomphonema yunnaniana Y.Liu et Kociolek in Liu et al., 2020

## Fossils

Several species are fossils.
* Gomphonema jianghanense Y.L.Li, Z.J.Gong, P.Xie & J.Shen, 2007 †
* Gomphonema jianghanensis Li, Gong, Xie & Shen, 2007 †
* Gomphonema jilinense Huang in Huang et al., 1983 †