# Sphenophorus

Sphenophorus is a genus of weevils, often known as billbugs, in the family Curculionidae, and tribe Sphenophorini. Eleven species of billbugs infest managed turfgrass in North America.

## Species

These 278 species belong to the genus Sphenophorus.
* Sphenophorus abbreviatus (J.C. Fabricius, 1787)
* Sphenophorus adspersus Gyllenhal, 1838
* Sphenophorus aduncus Erichson, 1847
* Sphenophorus aequalis Gyllenhal, 1838 (clay-coloured billbug)
* Sphenophorus albicollis Schoenherr, 1838
* Sphenophorus alfurus Heller, 1914
* Sphenophorus ambiguus Gyllenhal, 1838
* Sphenophorus anceps Gyllenhal, 1838
* Sphenophorus angustus Boheman, 1845
* Sphenophorus apicalis LeConte, 1878
* Sphenophorus ardesius Schoenherr, 1839
* Sphenophorus arizonensis Horn, 1873
* Sphenophorus asper Vaurie, 1978
* Sphenophorus aterrimus Champion, 1910
* Sphenophorus atratus Gyllenhal, 1838
* Sphenophorus atrivittata Chittenden, 1924
* Sphenophorus aurofasciatus Brême, 1844
* Sphenophorus austerus Gyllenhal, 1838
* Sphenophorus australis Chittenden, 1905
* Sphenophorus bartramiae Chittenden, 1924
* Sphenophorus basilanus Heller, 1922
* Sphenophorus bifasciatus Gyllenhal, 1838
* Sphenophorus bisbisignatus Gyllenhal, 1838
* Sphenophorus blanchardi Chittenden, 1905
* Sphenophorus blatchleyi Chittenden, 1924
* Sphenophorus brasiliensis Hustache, 1936
* Sphenophorus bruchi Hustache, 1936
* Sphenophorus brunnipennis (E.F. Germar, 1823)
* Sphenophorus brutus Gyllenhal, 1838
* Sphenophorus callizona L.A.A. Chevrolat, 1882
* Sphenophorus callosipennis Chittenden, 1919
* Sphenophorus callosus (Olivier, 1807) (southern corn billbug)
* Sphenophorus canaliculatus Boheman, 1845
* Sphenophorus canalipes Gyllenhal, 1838
* Sphenophorus carbonarius Schoenherr, 1838
* Sphenophorus carinicollis Gyllenhal, 1838
* Sphenophorus cariosus (Olivier, 1807) (nutgrass billbug)
* Sphenophorus caroli Vaurie, 1967
* Sphenophorus castaneipennis Boheman, 1838
* Sphenophorus castanipes Boheman, 1845
* Sphenophorus cazieri Vaurie, 1951
* Sphenophorus chittendeni Blatchley, 1916
* Sphenophorus cicatricosus Schoenherr, 1838
* Sphenophorus cicatripennis Fåhraeus, 1845
* Sphenophorus cicatristriatus Fahraeus, 1838 (Rocky Mountain billbug)
* Sphenophorus cincticollis Gyllenhal, 1838
* Sphenophorus cinctus Gyllenhal, 1838
* Sphenophorus cinerascens V. de Motschulsky, 1858
* Sphenophorus coesifrons Gyllenhal, 1838
* Sphenophorus compressirostris (Say, 1824)
* Sphenophorus confluens J. Prena, 2018
* Sphenophorus contractus Gyllenhal, 1838
* Sphenophorus costatus O.C. Waterhouse, 1879
* Sphenophorus costicollis Chittenden, 1919
* Sphenophorus costifer Chittenden, 1924
* Sphenophorus costipennis Horn, 1873
* Sphenophorus cribricollis F. Walker, 1859
* Sphenophorus cristulatus Hustache, 1936
* Sphenophorus cruciata G. Quedenfeldt, 1888
* Sphenophorus cruciger V. de Motschulsky, 1858
* Sphenophorus crudus Erichson, 1847
* Sphenophorus cultellatus Horn, 1873
* Sphenophorus cultrirostris Gyllenhal, 1838
* Sphenophorus decoratus Gyllenhal, 1838
* Sphenophorus deficiens Chittenden, 1920
* Sphenophorus defrictus Boheman, 1845
* Sphenophorus dehaanii Gyllenhal, 1838
* Sphenophorus delumbatus Boheman, 1845
* Sphenophorus destructor Chittenden, 1905
* Sphenophorus dietrichi (Satterthwait, 1933)
* Sphenophorus dimidiatipennis H. Jekel, 1858
* Sphenophorus discolor Mannerheim, 1843
* Sphenophorus dispar Gyllenhal, 1838
* Sphenophorus diversus Chittenden, 1905
* Sphenophorus dolosus Vaurie, 1978
* Sphenophorus elephantulus Chittenden, 1924
* Sphenophorus ensirostris Schoenherr, 1838
* Sphenophorus erythrurus L.A.A. Chevrolat, 1880
* Sphenophorus exquisita F. Walker, 1859
* Sphenophorus fahraei Gyllenhal, 1838
* Sphenophorus fasciatus Gemminger & Harold, 1871
* Sphenophorus ferrugineus Boheman, 1845
* Sphenophorus flexuosus Gyllenhal, 1838
* Sphenophorus fossor Gyllenhal, 1838
* Sphenophorus foveatus Vaurie, 1978
* Sphenophorus gagatinus Gyllenhal, 1838
* Sphenophorus geminatus Hustache, 1936
* Sphenophorus gentilis LeConte, 1857
* Sphenophorus germari Horn, 1873
* Sphenophorus glabricollis Gyllenhal, 1838
* Sphenophorus glabridiscus F. Walker, 1859
* Sphenophorus glyceriae J. Prena, 2018
* Sphenophorus graminis Chittenden, 1905
* Sphenophorus guttatus Fåhraeus, 1845
* Sphenophorus haneti G. Quedenfeldt, 1888
* Sphenophorus hebetatus Gyllenhal, 1838
* Sphenophorus hemipterus Schoenherr, 1825
* Sphenophorus hoegbergii Boheman, 1845
* Sphenophorus holosericus Chittenden, 1924
* Sphenophorus hypocrita Gyllenhal, 1838
* Sphenophorus immunis (T. Say, 1832)
* Sphenophorus implicatus Gyllenhal, 1838
* Sphenophorus imus Gyllenhal, 1838
* Sphenophorus inaequalis (Say, 1831)
* Sphenophorus incongruus Chittenden, 1905
* Sphenophorus incurrens Gyllenhal, 1838
* Sphenophorus indus (C. Linnaeus, 1758)
* Sphenophorus infelix Gistel, 1848
* Sphenophorus inscripta Gyllenhal, 1838
* Sphenophorus insculptus Gyllenhal, 1838
* Sphenophorus insularis C.H. Boheman, 1859
* Sphenophorus interruptecostata Schaufuss, 1885
* Sphenophorus interstitialis (Say, 1831)
* Sphenophorus intervallatus Chittenden, 1924
* Sphenophorus javanensis Günther, 1937
* Sphenophorus jugosus Chittenden, 1924
* Sphenophorus kuatunensis Voss, 1958
* Sphenophorus laetus Erichson, 1847
* Sphenophorus larvalis Schoenherr, 1838
* Sphenophorus lateritius Gyllenhal, 1838
* Sphenophorus latinasus Horn, 1873
* Sphenophorus latiscapus T. Kirsch, 1869
* Sphenophorus lebasii Gyllenhal, 1838
* Sphenophorus leprosus Fåhraeus, 1845
* Sphenophorus letestui Hustache, 1936
* Sphenophorus leucographus Fåhraeus, 1845
* Sphenophorus levis Vaurie, 1978
* Sphenophorus lineatocollis Gyllenhal, 1838
* Sphenophorus lineatus Champion, 1910
* Sphenophorus liratus Gyllenhal, 1838
* Sphenophorus longicollis Schoenherr, 1838
* Sphenophorus lutulentus Champion, 1910
* Sphenophorus maculata Gyllenhal, 1838
* Sphenophorus maculifer Fåhraeus, 1845
* Sphenophorus maderensis Hoffmann, 1938
* Sphenophorus maidis Chittenden, 1905 (maize billbug)
* Sphenophorus marinus Chittenden, 1905
* Sphenophorus maurus Gyllenhal, 1838
* Sphenophorus melancholicus Gyllenhal, 1838
* Sphenophorus melanocardius Schoenherr, 1838
* Sphenophorus melanocephalus (Fabricius, 1801)
* Sphenophorus melanurus T. Kirsch, 1875
* Sphenophorus memnonius Gyllenhal, 1838
* Sphenophorus meridionalis Gyllenhal, 1838
* Sphenophorus mimelus Vaurie, 1978
* Sphenophorus minimus Hart, 1890 (lesser billbug)
* Sphenophorus missouriensis Chittenden, 1919
* Sphenophorus monilis Gyllenhal, 1838
* Sphenophorus mormon Chittenden, 1904
* Sphenophorus multipunctatus Champion, 1910
* Sphenophorus mundus Champion, 1910
* Sphenophorus musaecola L. Fairmaire, 1898
* Sphenophorus mutilatus Schoenherr, 1838
* Sphenophorus napoanus Hustache, 1936
* Sphenophorus nasutus Chittenden, 1924
* Sphenophorus nawradi T. Kirsch, 1869
* Sphenophorus nebulosus G.C. Champion, 1910
* Sphenophorus necydaloides (Fabricius, 1801)
* Sphenophorus neomexicanus Chittenden, 1904
* Sphenophorus nevadensis Chittenden, 1905
* Sphenophorus nigerrimus Gyllenhal, 1838
* Sphenophorus nigroplagiata G. Quedenfeldt, 1888
* Sphenophorus nubilus Gyllenhal, 1838
* Sphenophorus nudicollis T. Kirsch, 1877
* Sphenophorus obscuripennis Chittenden, 1924
* Sphenophorus obsoletus Gyllenhal, 1838
* Sphenophorus octocostatus Champion, 1910
* Sphenophorus octomaculatus Heller, 1934
* Sphenophorus omissus Blatchley, 1920
* Sphenophorus opacus Gyllenhal, 1838
* Sphenophorus orizabensis L.A.A. Chevrolat, 1882
* Sphenophorus paganus (J.C. Fabricius, 1775)
* Sphenophorus panops Walk. in Mshl., 1920
* Sphenophorus parumpunctatus Gyllenhal, 1838
* Sphenophorus perforatus Fåhraeus, 1845
* Sphenophorus pertinax (Olivier, 1807)
* Sphenophorus phoeniciensis Chittenden, 1904 (phoenix billbug)
* Sphenophorus piceus (P.S. Pallas, 1771)
* Sphenophorus picirostris L. Fairmaire, 1897
* Sphenophorus pinguis Chittenden, 1924
* Sphenophorus placidus Schoenherr, 1838
* Sphenophorus planipennis Gyllenhal, 1838
* Sphenophorus polygrammus Gyllenhal, 1838
* Sphenophorus pontederiae Chittenden, 1905
* Sphenophorus procerus J.L. LeConte, 1858
* Sphenophorus promissa F.P. Pascoe, 1885
* Sphenophorus pulchellus Gyllenhal, 1838
* Sphenophorus pumilus Gyllenhal, 1838
* Sphenophorus punctatostriatus Gyllenhal, 1838
* Sphenophorus punctatus Gyllenhal, 1838
* Sphenophorus pustulosus Gyllenhal, 1838
* Sphenophorus quadrimaculata Gyllenhal, 1838
* Sphenophorus quadrisignatus Gyllenhal, 1838
* Sphenophorus quadrivittatus Gyllenhal, 1838
* Sphenophorus quadrivulnerata J. Thomson, 1858
* Sphenophorus quinquepunctatus Gyllenhal, 1838
* Sphenophorus rectistriatus Chittenden, 1924
* Sphenophorus rectus (Say, 1831)
* Sphenophorus reticulaticollis J. Prena, 2018
* Sphenophorus reticulatus Chittenden, 1924
* Sphenophorus retusus Gyllenhal, 1838
* Sphenophorus rimoratus Gyllenhal, 1838
* Sphenophorus robustior Chittenden, 1905
* Sphenophorus robustus Horn, 1873
* Sphenophorus roelofsi L.A.A. Chevrolat, 1882
* Sphenophorus rubellus Gyllenhal, 1838
* Sphenophorus rufofasciatus Gyllenhal, 1838
* Sphenophorus rufus Szalay Marzsó, 1957
* Sphenophorus rusticus Gyllenhal, 1838
* Sphenophorus sacchari Schoenherr, 1838
* Sphenophorus sanguineus Gyllenhal, 1845
* Sphenophorus sanguinolentus L.A.A. Chevrolat, 1882
* Sphenophorus sationis Heller, 1925
* Sphenophorus saucius Gyllenhal, 1838
* Sphenophorus sayi Gyllenhal, 1838
* Sphenophorus schoenherrii Gyllenhal, 1838
* Sphenophorus schwarzi Chittenden, 1924
* Sphenophorus scirpi Chittenden, 1924
* Sphenophorus scoparius Horn, 1873
* Sphenophorus semicalvus Hustache, 1936
* Sphenophorus senegalensis Gyllenhal, 1838
* Sphenophorus sericans C.H. Boheman, 1845
* Sphenophorus sericeus Gyllenhal, 1838
* Sphenophorus seriepunctatus Gyllenhal, 1838
* Sphenophorus serratipes Chittenden, 1924
* Sphenophorus sierrakowskyi Gyllenhal, 1838
* Sphenophorus signaticollis Gyllenhal, 1838
* Sphenophorus simplex LeConte, 1860
* Sphenophorus soltauii Chittenden, 1905
* Sphenophorus sordidus Schoenherr, 1838
* Sphenophorus spadiceus Gyllenhal, 1838
* Sphenophorus spangleri Anderson, 2014
* Sphenophorus spinolae Gyllenhal, 1838
* Sphenophorus squamosus Boheman, 1845
* Sphenophorus sriatoforatus Gyllenhal, 1838
* Sphenophorus stigmaticus Fåhraeus, 1845
* Sphenophorus strangulatus Gyllenhal, 1838
* Sphenophorus striatipennis Chittenden, 1905
* Sphenophorus striatoforatus Gyllenhal, 1838
* Sphenophorus striatopunctatus (J.A.E. Goeze, 1777)
* Sphenophorus striatus Fåhraeus, 1845
* Sphenophorus strigosus Erichson, 1847
* Sphenophorus subcarinatus Mannerheim, 1843
* Sphenophorus subcostatus L. Fairmaire, 1903
* Sphenophorus sublaevis F.H. Chittenden, 1906
* Sphenophorus subulatus Chittenden, 1905
* Sphenophorus suturalis Gyllenhal, 1838
* Sphenophorus taboa Vanin, 1990
* Sphenophorus tardus Fall, 1901
* Sphenophorus tenuis Vaurie, 1978
* Sphenophorus terebrans Schoenherr, 1838
* Sphenophorus terricola Champion, 1910
* Sphenophorus testardi X. Montrouzier, 1860
* Sphenophorus tetricus Gyllenhal, 1838
* Sphenophorus thoracicus Gyllenhal, 1838
* Sphenophorus tincturata F.P. Pascoe, 1885
* Sphenophorus tomentosus Vaurie, 1978
* Sphenophorus tornowii Brèthes, 1910
* Sphenophorus tredecimpunctatus Schoenherr, 1838
* Sphenophorus tremolerasi Hustache, 1937
* Sphenophorus truncatus Schoenherr, 1838
* Sphenophorus validirostris Gyllenhal, 1838
* Sphenophorus validus J.L. LeConte, 1858
* Sphenophorus variabilis Gyllenhal, 1838
* Sphenophorus variegatus Schoenherr, 1838
* Sphenophorus velutinus LeConte, 1876
* Sphenophorus venatus (Say, 1831) (hunting billbug)
* Sphenophorus vestitus J. Prena, 2018
* Sphenophorus vilis Hustache, 1936
* Sphenophorus villosiventris Chittenden, 1905
* Sphenophorus vittata Chittenden, 1924
* Sphenophorus vomerinus LeConte, 1858
* Sphenophorus zamiae Gyllenhal, 1838
* Sphenophorus zeae Walsh, 1867 (Timothy billbug)
* † Sphenophorus elegans Theobald, 1935
* † Sphenophorus naegelianus O. Heer, 1847
* † Sphenophorus proluviosus Heyden & Heyden, 1866
* † Sphenophorus regelianus O. Heer, 1847