# List of ground beetle genera

This page lists the genera in the family Carabidae, according to Catalogue of Life and Carabcat Database as of March 2023, excluding Tiger Beetles (which are considered to be in the family Cicindelidae).

## Family Carabidae
### Subfamily Anthiinae Bonelli, 1813

    Tribe Anthiini Bonelli, 1813
       Anthia Weber, 1801
       Atractonotus Perroud, 1847
       Baeoglossa Chaudoir, 1850
       Cycloloba Chaudoir, 1850
       Cypholoba Chaudoir, 1850
       Eccoptoptera Chaudoir, 1878
       Gonogenia Chaudoir, 1844
       Netrodera Chaudoir, 1850
    Tribe Helluonini Hope, 1838
       Subtribe Helluonina Hope, 1838
          Aenigma Newman, 1836
          Ametroglossus Sloane, 1914
          Dicranoglossus Chaudoir, 1872
          Epimicodema Sloane, 1914
          Gigadema J.Thomson, 1859
          Helluapterus Sloane, 1914
          Helluarchus Sloane, 1914
          Helluo Bonelli, 1813
          Helluodema Laporte, 1867
          Helluonidius Chaudoir, 1872
          Helluopapua Darlington, 1968
          Helluosoma Laporte, 1867
          Neohelluo Sloane, 1914
          Platyhelluo Baehr, 2005
       Subtribe Omphrina Jedlicka, 1941
          Colfax Andrewes, 1920
          Creagris Nietner, 1857
          Dailodontus Reiche, 1843
          Erephognathus Alluaud, 1932
          Helluobrochus Reichardt, 1974
          Helluomorpha Laporte, 1834
          Helluomorphoides Ball, 1951
          Macrocheilus Hope, 1838
          Meladroma Motschulsky, 1855
          Omphra Dejean, 1825
          Pleuracanthus Gray, 1832
          Triaenogenius Chaudoir, 1877
    Tribe Physocrotaphini Chaudoir, 1863
       Anguloderus Anichtchenko & Sciaky, 2017
       Foveocrotaphus Anichtchenko, 2014
       Helluodes Westwood, 1847
       Holoponerus Fairmaire, 1883
       Physocrotaphus Parry, 1849
       Physoglossus Akhil & Sabu, 2020
       Pogonoglossus Chaudoir, 1863
       Schuelea Baehr, 2004

### Subfamily Apotominae LeConte, 1853

    Apotomus Illiger, 1807

### Subfamily Brachininae Bonelli, 1810

    Tribe Brachinini Bonelli, 1810
       Aptinoderus Hubenthal, 1919
       Aptinus Bonelli, 1810
       Brachinulus Basilewsky, 1958
       Brachinus Weber, 1801
       Mastax Fischer von Waldheim, 1828
       Pheropsophus Solier, 1833
       Styphlodromus Basilewsky, 1959
       Styphlomerus Chaudoir in Putzeys, 1875
    Tribe Crepidogastrini Jeannel, 1949
       Brachynillus Reitter, 1904
       Crepidogaster Boheman, 1848
       Crepidogastrillus Basilewsky, 1959
       Crepidogastrinus Basilewsky, 1957
       Crepidolomus Basilewsky, 1959
       Crepidonellus Basilewsky, 1959

### Subfamily Broscinae Hope, 1838

    Tribe Broscini Hope, 1838
       Subtribe Axonyina Roig-Juñent, 2000
          Broscodes Bolivar y Pieltain, 1914
          Rawlinsius Davidson & Ball, 1998
       Subtribe Baripodina Jeannel, 1941
          Baripus Dejean, 1828
          Bembidiomorphum Champion, 1918
       Subtribe Broscina Hope, 1838
          Broscodera Lindroth, 1961
          Broscosoma Rosenhauer, 1846
          Broscus Panzer, 1813
          Chaetobroscus Semenov, 1900
          Craspedonotus Schaum, 1863
          Eobroscus Kryzhanovskij, 1951
          Kashmirobroscus J.Schmidt; Wrase & Sciaky, 2013
          Miscodera Eschscholtz, 1830
          Zacotus LeConte, 1869
       Subtribe Creobiina Jeannel, 1941
          Acallistus Sharp, 1886
          Adotela Laporte, 1867
          Anheterus Putzeys, 1868
          Bountya Townsend, 1971
          Brithysternum W.J.MacLeay, 1873
          Cascellius Curtis, 1838
          Cerotalis Laporte, 1867
          Creobius Guérin-Méneville, 1838
          Gnathoxys Westwood, 1842
          Nothocascellius Roig-Juñent, 1995
          Promecoderus Dejean, 1829
       Subtribe Nothobroscina Roig-Juñent, 2000
          Chylnus Sloane, 1920
          Diglymma Sharp, 1886
          Eurylychnus Bates, 1891
          Mecodema Blanchard, 1843
          Monteremita Seldon & Holwell, 2019
          Nothobroscus Roig-Juñent & Ball, 1995
          Oregus Putzeys, 1868
          Orthoglymma Liebherr; Marris; Emberson; Syrett & Roig-Juñent, 2011
          Percolestus Sloane, 1892
          Percosoma Schaum, 1858

### Subfamily Carabinae Linnaeus, 1802

    Tribe Carabini Linnaeus, 1802
       Subtribe Carabina Linnaeus, 1802
          Aplothorax G.R.Waterhouse, 1842
          Calosoma Weber, 1801
          Carabus Linnaeus, 1758
       Subtribe Ceroglossina Lapouge, 1927
          Ceroglossus Solier, 1848
    Tribe Cychrini Perty, 1830
       Subtribe Cychrina Perty, 1830
          Cychropsis Boileau, 1901
          Cychrus Fabricius, 1794
          Scaphinotus Dejean, 1826
          Sphaeroderus Dejean, 1826
       Subtribe Pamborina Hope, 1838
          Maoripamborus Brookes, 1944
          Pamborus Latreille, 1812

### Subfamily Ctenodactylinae Laporte, 1834

    Tribe Ctenodactylini Laporte, 1834
       Alachnothorax Liebke, 1929
       Amblycoleus Chaudoir, 1872
       Antipionycha Liebke, 1928
       Askalaphium Liebke, 1938
       Ctenodactyla Dejean, 1825
       Leptotrachelon Liebke, 1928
       Leptotrachelus Latreille, 1829
       Oilea Liebke, 1931
       Parapionycha Liebke, 1929
       Pionycha Chaudoir, 1848
       Plagiotelum Solier, 1849
       Propionycha Liebke, 1928
       Pseudometabletus Liebke, 1930
       Schidonychus Klug, 1834
       Teukrus Liebke, 1931
       Wate Liebke, 1928
    Tribe Hexagoniini G.Horn, 1881
       Dinopelma Bates, 1889
       Hexagonia Kirby, 1825
       Omphreoides Fairmaire, 1896

### Subfamily Dryptinae Bonelli, 1810

    Tribe Dryptini Bonelli, 1810
       Desera Hope, 1831
       Drypta Latreille, 1797
       Megadrypta Sciaky & Anichtchenko, 2020
       Neodrypta Basilewsky, 1960
       Nesiodrypta Jeannel, 1949
       Prionodrypta Jeannel, 1949
    Tribe Galeritini Kirby, 1825
       Ancystroglossus Chaudoir, 1863
       Eunostus Laporte, 1835
       Galerita Fabricius, 1801
       Trichognatha Latreille, 1829
    Tribe Zuphiini Bonelli, 1810
       Subtribe Dicrodontina Machado, 1992
          Dicrodontus Chaudoir, 1872
       Subtribe Leleupidiina Basilewsky, 1951
          Colasidia Basilewsky, 1954
          Gunvorita Landin, 1955
          Leleupidia Basilewsky, 1951
          Neoleleupidia Basilewsky, 1953
          Paraleleupidia Basilewsky, 1951
       Subtribe Planetina Jedlicka, 1941
          Planetes W.S.MacLeay, 1825
       Subtribe Zuphiina Bonelli, 1810
          Acrogenys W.J.MacLeay, 1864
          Agastus Schmidt-Goebel, 1846
          Chaudoirella Mateu, 1982
          Coarazuphium Gnaspini; Vanin & Godoy, 1998
          Ildobates Español, 1966
          Metaxidius Chaudoir, 1852
          Metazuphium Mateu, 1992
          Mischocephalus Chaudoir, 1863
          Parazuphium Jeannel, 1942
          Polistichus Bonelli, 1810
          Pseudaptinus Laporte, 1834
          Speothalpius B.Moore, 1995
          Speozuphium B.Moore, 1995
          Typhlozuphium Baehr, 2014
          Zuphioides Ball & Shpeley, 2013
          Zuphium Latreille, 1805

### Subfamily Elaphrinae Latreille, 1802

    Blethisa Bonelli, 1810
    Diacheila Motschulsky, 1844
    Elaphrus Fabricius, 1775
    †Elaphrotites Haupt, 1856

### Subfamily Gineminae Ball & Shpeley, 2002

    Ginema Ball & Shpeley, 2002

### Subfamily Harpalinae Bonelli, 1810

    Tribe Anisodactylini Lacordaire, 1854
       Allendia Noonan, 1974
       Allocinopus Broun, 1903
       Amphasia Newman, 1838
       Anisodactylus Dejean, 1829
       Anisostichus Emden, 1953
       Cenogmus Sloane, 1898
       Chydaeus Chaudoir, 1854
       Crasodactylus Guérin-Méneville, 1847
       Criniventer Emden, 1953
       Diachromus Erichson, 1837
       Dicheirus Mannerheim, 1843
       Gaioxenus Broun, 1910
       Geopinus LeConte, 1847
       Gnathaphanus W.S.MacLeay, 1825
       Gynandromorphus Dejean, 1829
       Haplaner Chaudoir, 1878
       Harpalomimetes Schauberger in Csiki, 1932
       Hypharpax W.S.MacLeay, 1825
       Maoriharpalus Larochelle & Larivière, 2005
       Nornalupia Kataev, 2002
       Notiobia Perty, 1830
       Parabaris Broun, 1881
       Progonochaetus G.Müller, 1938
       Pseudanisotarsus Noonan, 1973
       Pseudognathaphanus Schauberger, 1932
       Pseudorhysopus Kataev & Wrase, 2001
       Rhysopus Andrewes, 1929
       Scybalicus Schaum, 1862
       Triplosarus Bates, 1874
       Tuiharpalus Larochelle & Larivière, 2005
       Xestonotus LeConte, 1853
    Tribe Harpalini Bonelli, 1810
       Subtribe Amblystomina Fauvel, 1889
          Amblystomus Erichson, 1837
          Anomostomus LaFerté-Sénectère, 1853
          Barysomus Dejean, 1829
          Oosoma Nietner, 1857
       Subtribe Ditomina Bonelli, 1810
          Bronislavia Semenov, 1891
          Carenochyrus Solsky, 1874
          Carterus Dejean, 1830
          Chilotomus Chaudoir, 1842
          Ditomus Bonelli, 1810
          Dixus Billberg, 1820
          Eocarterus Stichel, 1923
          Eucarterus Reitter, 1900
          Graniger Motschulsky, 1864
          Indocarterus Kataev & Wrase, 2018
          Liochirus Tschitscherine, 1897
          Machozetus Chaudoir, 1850
          Odotoncarus Solier, 1835
          Oedesis Motschulsky, 1850
          Pachycarus Solier, 1835
          Parapenthus Kataev & Wrase, 2018
          Penthus Chaudoir, 1843
          Phorticosomus Schaum, 1863
          Proditomus Schauberger, 1934
          Pseudaristus Reitter, 1900
          Tschitscherinellus Csiki, 1906
       Subtribe Harpalina Bonelli, 1810
          Acinopus Dejean, 1821
          Afromizonus Basilewsky, 1947
          Allosiopelus N.Ito, 1995
          Amblygnathus Dejean, 1829
          Anisocnemus Chaudoir, 1843
          Athrostictus Bates, 1878
          Axinotoma Dejean, 1829
          Aztecarpalus Ball, 1970
          Bleusea Bedel, 1897
          Boeomimetes Péringuey, 1896
          Bradybaenus Dejean, 1829
          Coleolissus Bates, 1892
          Cratacanthus Dejean, 1829
          Cratognathus Dejean, 1829
          Daptus Fischer von Waldheim, 1823
          Dioryche W.S.MacLeay, 1825
          Discoderus LeConte, 1853
          Dregus Motschulsky, 1864
          Ectinothorax Alluaud, 1941
          Eriophonus Tschitscherine, 1901
          Euryderus LeConte, 1846
          Geodromus Dejean, 1829
          Harpalinus Jeannel, 1946
          Harpaliscus Bates, 1892
          Harpalobrachys Tschitscherine, 1899
          Harpalodiodes Bousquet, 2002
          Harpalomimus Facchini & Giachino, 2020
          Harpalomorphus Péringuey, 1896
          Harpalus Latreille, 1802
          Harpathaumas Basilewsky, 1947
          Harponixus Basilewsky, 1950
          Hartonymus Casey, 1914
          Heteracantha Brullé, 1835
          Hyphaereon W.S.MacLeay, 1825
          Indiophonus N.Ito, 1996
          Kareya Andrewes, 1919
          Lampetes Andrewes, 1940
          Laparhetes Jeannel, 1946
          Liodaptus Bates, 1889
          Meroctenus Gemminger & Harold, 1868
          Microderes Faldermann, 1836
          Neoaulacoryssus Noonan, 1985
          Neodiachipteryx Noonan, 1985
          Neohyparpalus Clarke, 1981
          Neophygas Noonan, 1976
          Nesacinopus Tschitscherine, 1900
          Nesarpalus Bedel, 1897
          Nipponoharpalus Habu, 1973
          Nothodaptus Maindron, 1906
          Oesyperus Andrewes, 1923
          Omostropus Péringuey, 1896
          Ooidius Chaudoir, 1847
          Ophoniscus Bates, 1892
          Ophonus Dejean, 1821
          Oxycentrus Chaudoir, 1854
          Panagrius Andrewes, 1933
          Pangus Dejean, 1821
          Parasiopelus Basilewsky, 1946
          Paraulacoryssus Shpeley; Hunting & Ball, 2017
          Parophonus Ganglbauer, 1891
          Paulianoscirtus Basilewsky, 1976
          Penthophonus Reitter, 1900
          Piosoma LeConte, 1847
          Platymetopsis Ball & Maddison, 1987
          Platymetopus Dejean, 1829
          Prakasha Andrewes, 1919
          Pseudodiachipteryx Burgeon, 1936
          Pseudohyparpalus Basilewsky, 1946
          Pseudoselenophorus Péringuey, 1896
          Selenophorus Dejean, 1829
          Selenotichnus Kataev, 1999
          Siopelophonus Kataev, 2015
          Siopelus Murray, 1859
          Stenomorphus Dejean, 1831
          Trichopselaphus Chaudoir, 1843
          Trichotichnus A.Morawitz, 1863
          Trichoxycentrus N.Ito, 2000
          Typsiharpalus Tschitscherine, 1901
    Tribe Pelmatellini Bates, 1882
       Hakaharpalus Larochelle & Larivière, 2005
       Kupeharpalus Larochelle & Larivière, 2005
       Lecanomerus Chaudoir, 1850
       Nemaglossa Solier, 1849
       Notospeophonus B.Moore, 1962
       Pelmatellus Bates, 1882
       Syllectus Bates, 1878
       Trachysarus Reed, 1874
    Tribe Stenolophini Kirby, 1837
       Acupalpus Latreille, 1829
       Agonoleptus Casey, 1914
       Amerinus Casey, 1884
       Angionychus Klug, 1853
       Anthracus Motschulsky, 1850
       Batoscelis Dejean, 1836
       Bradycellus Erichson, 1837
       Cratosoma Jeannel, 1948
       Cyptomicrus Vinson, 1939
       Dicheirotrichus Jacquelin du Val, 1855
       Egadyla Alluaud, 1916
       Euthenarus Bates, 1874
       Fuminoria Morita, 2006
       Goniocellus Casey, 1914
       Gugheorites Basilewsky, 1951
       Haplanister B.Moore, 1996
       Hemiaulax Bates, 1892
       Hippoloetis Laporte, 1835
       Idiomelas Tschitscherine, 1900
       Kaffovatus Clarke, 1972
       Kenyacus Alluaud, 1917
       Kiwiharpalus Larochelle & Larivière, 2005
       Lioholus Tschitscherine, 1897
       Loxoncus Schmidt-Goebel, 1846
       Pachytrachelus Chaudoir, 1852
       Parabradycellus N.Ito, 2003
       Paramecus Dejean, 1829
       Philodes LeConte, 1861
       Pholeodytes Britton, 1962
       Pogonodaptus G.Horn, 1881
       Polpochila Solier, 1849
       Psychristus Andrewes, 1930
       Rhabidius Basilewsky, 1948
       Stenolophus Dejean, 1821
       Uenanthracus Kasahara, 1994

### Subfamily Hiletinae Schiödte, 1847

    Eucamaragnathus Jeannel, 1938
    Hiletus Schiödte, 1847

### Subfamily Lebiinae Bonelli, 1810

    Tribe Cyclosomini Laporte, 1834
       Subtribe Corsyrina Ganglbauer
          Corsyra Dejean, 1825
          Discoptera Semenov, 1889
       Subtribe Cyclosomina Laporte, 1834
          Cyclosomus Latreille, 1829
          Sarothrocrepis Chaudoir, 1850
          Tetragonoderus Dejean, 1829
       Subtribe Graphipterina Latreille, 1802
          Graphipterus Latreille, 1802
          Piezia Brullé, 1834
          Trichopiezia Nègre, 1955
       Subtribe Masoreina Chaudoir, 1871
          Anaulacus W.S.MacLeay, 1825
          Atlantomasoreus Mateu, 1984
          Leuropus Andrewes, 1947
          Lophidius Dejean, 1831
          Masoreus Dejean, 1821
          Mnuphorus Chaudoir, 1873
          Odontomasoreus Darlington, 1968
          Somoplatodes Basilewsky, 1986
          Somoplatus Dejean, 1829
       Subtribe Nemotarsina Bates, 1883
          Nemotarsus LeConte, 1853
    Tribe Lachnophorini LeConte, 1853
       Subtribe Calophaenina Jeannel, 1948
          Calophaena Klug, 1821
          Calophaenoidea Liebke, 1930
       Subtribe Lachnophorina LeConte, 1853
          Amphithasus Bates, 1871
          Anchonoderus Reiche, 1843
          Aporesthus Bates, 1871
          Asklepia Liebke, 1938
          Balligratus Moret & Ortuño, 2017
          Calybe Laporte, 1834
          Ega Laporte, 1834
          Eucaerus LeConte, 1853
          Euphorticus G.Horn, 1881
          Grundmannius Basilewsky, 1965
          Guatemalteca Erwin, 2004
          Lachnaces Bates, 1872
          Lachnophorus Dejean, 1831
          Peruphorticus Erwin & Zamorano, 2014
          Pseudophorticus Erwin, 2004
          Selina Motschulsky, 1858
    Tribe Lebiini Bonelli, 1810
       Subtribe Agrina Kirby, 1837
          Abaditicus Ball & Hilchie, 1983
          Abrodiella Bousquet, 2002
          Agra Fabricius, 1801
          Allardina Basilewsky, 1963
          Amelus Chaudoir, 1873
          Anomotariella Baehr, 2012
          Anomotarus Chaudoir, 1875
          Apterodromites Mateu, 1976
          Aspasiola Chaudoir, 1877
          Calleida Latreille, 1824
          Calleidomorpha Motschulsky, 1855
          Callidadelpha Steinheil, 1875
          Chaudoirina Mateu, 1955
          Cryptobatis Eschscholtz, 1829
          Cyanotarus Reed, 1874
          Cylindronotum Putzeys, 1845
          Demetrida White, 1846
          Diabaticus Bates, 1878
          Do Baehr, 2009
          Dromidea Perroud & Montrouzier, 1864
          Epikastea Liebke, 1935
          Eujalmenus Bousquet, 2002
          Euplatia Chaudoir, 1873
          Eurycalleida Maindron, 1905
          Falsodromius Mateu, 1976
          Glycia Chaudoir, 1842
          Hyboptera Chaudoir, 1873
          Hybopteroides Erwin & Ball, 2012
          Infernophilus Larson, 1969
          Kteatus Liebke, 1936
          Lipostratia Chaudoir, 1873
          Merizomena Chaudoir, 1873
          Mimodromius Chaudoir, 1873
          Mimophilorhizus Mateu, 1993
          Oechalius Liebke, 1935
          Ogygium Liebke, 1935
          Onota Chaudoir, 1873
          Otoglossa Chaudoir, 1873
          Paraglycia Bedel, 1904
          Phacocerus Chaudoir, 1873
          Philophuga Motschulsky, 1859
          Phloeocarabus W.J.MacLeay, 1871
          Plochionus Dejean, 1821
          Pontonoa Liebke, 1935
          Pseudotoglossa Mateu, 1961
          Pylartesius Liebke, 1939
          Rugitarus Baehr, 2009
          Speotarus B.Moore, 1964
          Straneotia Mateu, 1961
          Tecnophilus Chaudoir, 1877
          Teiresia Liebke, 1935
          Thoasia Liebke, 1939
          Titaresius Liebke, 1935
          Trigonothops W.J.MacLeay, 1864
          Valeriaaschero Erwin, 2004
          Vianasia Mateu, 1955
       Subtribe Apenina Ball
          Apenes LeConte, 1851
          Cymindoidea Laporte, 1833
          Habutarus Ball & Hilchie, 1983
          Platytarus Fairmaire, 1850
          Trymosternus Chaudoir, 1873
       Subtribe Celaenephina Habu, 1982
          Celaenephes Schmidt-Goebel, 1846
       Subtribe Cymindidina Laporte, 1834
          Afrotarus Jeannel, 1949
          Assadecma Basilewsky, 1982
          Cymindis Latreille, 1805
          Hystrichopus Boheman, 1848
          Leptosarcus Péringuey, 1896
          Metaxymorphus Chaudoir, 1850
          Petrimagnia Kryzhanovskij & Mikhailov, 1971
          Pinacodera Schaum, 1857
          Plagiopyga Boheman, 1848
          Pseudomasoreus Desbrochers des Loges, 1904
          Taridius Chaudoir, 1875
       Subtribe Demetriadina Bates, 1886
          Cylindrocranius Chaudoir, 1878
          Demetrias Bonelli, 1810
       Subtribe Dromiusina Bonelli, 1810
          Afrodromius Basilewsky, 1958
          Axinopalpus LeConte, 1846
          Barrymooreana Baehr, 1997
          Brachynopterus Bedel, 1898
          Brigalowia Baehr, 2006
          Calodromius Reitter, 1905
          Carbonellia Mateu, 1968
          Disciferella Kataev & Muilwijk, 2020
          Dromiops Péringuey, 1898
          Dromius Bonelli, 1810
          Dromoceryx Schmidt-Goebel, 1846
          Geoffreyella Baehr, 2012
          Klepteromimus Péringuey, 1898
          Mesolestes Schatzmayr, 1943
          Metadromius Bedel, 1907
          Microdaccus Schaum in Baudi di Selve, 1864
          Microlestes Schmidt-Goebel, 1846
          Microlestodes Baehr, 1987
          Monnea Mateu, 1970
          Negrea Mateu, 1968
          Omophagus Andrewes, 1937
          Oxoides Solier, 1849
          Paradromius Fowler, 1887
          Philorhizus Hope, 1838
          Polyaulacus Chaudoir, 1878
          Psammodromius Peyerimhoff, 1927
          Pseudomonnea Mateu, 1983
          Somalodromius Mateu, 1967
          Xenodromius Bates, 1891
          Zolotarevskyella Mateu, 1953
       Subtribe Gallerucidiina Chaudoir, 1872
          Gallerucidia Chaudoir, 1872
          Lebidia A.Morawitz, 1862
       Subtribe Lebiina Bonelli, 1810
          Alkestis Liebke, 1939
          Aristolebia Bates, 1892
          Daer Semenov & Znojko, 1929
          Lachnolebia Maindron, 1905
          Lebia Latreille, 1802
          Lebidema Motschulsky, 1864
          Lebiomorphica Lorenz, 1998
          Lebistina Motschulsky, 1864
          Lebistinida Péringuey, 1898
          Lionedya Chaudoir, 1871
          Matabele Péringuey, 1896
          Megalebia Mateu, 1972
          Orthobasis Chaudoir, 1871
          Pachylebia Jeannel, 1949
          Pachylebiodes Mateu, 1972
          Paulianolebia Mateu, 1972
          Pseudopachylebia Mateu, 1972
          Rhopalostyla Chaudoir, 1850
          Scalidion Schmidt-Goebel, 1846
          Setolebia Jedlicka, 1941
          Sofota Jedlicka, 1951
       Subtribe Lichnasthenina J.Thomson
          Astastus Péringuey, 1896
          Australovelinda Baehr, 2012
          Imasakaia Arai & Morita, 2010
          Lichnasthenus J.Thomson, 1858
          Mimovelindopsis Mateu, 1963
          Velinda Andrewes, 1921
          Velindastus Schüle & Lorenz, 2008
          Velindomimus Jeannel, 1955
          Velindopsis Burgeon, 1937
       Subtribe Lionychina Jeannel, 1948
          Apristomimus Mateu, 1969
          Apristus Chaudoir, 1846
          Eremolestes Maindron, 1905
          Lionychus Wissmann, 1846
          Lorestania Anichtchenko, 2011
          Metablus Jedlicka, 1958
          Omobrus Andrewes, 1930
          Singiliomimus Péringuey, 1896
          Syntomus Hope, 1838
          Tilius Chaudoir, 1876
          Trichidema Basilewsky, 1956
       Subtribe Metallicina Basilewsky, 1984
          Euproctinus Leng & Mutchler, 1927
          Metallica Chaudoir, 1873
          Pachycallida Jeannel, 1949
          Parena Motschulsky, 1860
       Subtribe Peliocypadina Basilewsky, 1984
          Demetriola Jeannel, 1949
          Peliocypas Schmidt-Goebel, 1846
       Subtribe Pericalina Hope, 1838
          Agonocheila Chaudoir, 1848
          Allophanes Andrewes, 1939
          Allophanopsis Louwerens, 1952
          Amphimenes Bates, 1873
          Amphimenoides Kirschenhofer, 1999
          Antimerina Alluaud, 1897
          Arsinoe Laporte, 1834
          Auchmerus Andrewes, 1929
          Bellavalentis Hunting & Yang, 2019
          Brachichila Chaudoir, 1870
          Brachyctis Chaudoir, 1870
          Catascopellus Straneo, 1969
          Catascopus Kirby, 1825
          Cheilagona Baehr, 2006
          Coptodera Dejean, 1825
          Coptoglossus Chaudoir, 1870
          Coptoptera Chaudoir, 1837
          Coptopterella Basilewsky, 1961
          Coptopterina Basilewsky, 1956
          Crassagena Baehr, 2006
          Cylindropectus Lorenz, 1998
          Dolichoctis Schmidt-Goebel, 1846
          Dontolobus Basilewsky, 1970
          Drymatus Motschulsky, 1862
          Eucheila Dejean, 1829
          Eurycoleus Chaudoir, 1848
          Eurydera Laporte, 1831
          Formosiella Jedlicka, 1951
          Gidda Andrewes, 1920
          Glyphodactyla Chaudoir, 1837
          Holcoderus Chaudoir, 1870
          Horniulus Jedlicka, 1932
          Labocephalus Chaudoir, 1848
          Lelis Chaudoir, 1870
          Lioptera Chaudoir, 1870
          Lobodontidius Basilewsky, 1970
          Lobodontulus Basilewsky, 1970
          Lobodontus Chaudoir, 1842
          Madecassina Jeannel, 1949
          Mascarenhia Alluaud, 1933
          Menarus Andrewes, 1939
          Metascopus Basilewsky, 1970
          Minuphloeus Darlington, 1968
          Minuthodes Andrewes, 1941
          Miscelus Klug, 1834
          Mochtherus Schmidt-Goebel, 1846
          Mormolyce Hagenbach, 1825
          Mormolycina Jeannel, 1949
          Neocoptodera Jeannel, 1949
          Nycteis Laporte, 1835
          Oreodicastes Maindron, 1905
          Oxyodontus Chaudoir, 1870
          Paradolichoctis Baehr, 2006
          Pareurydera Jeannel, 1949
          Pectinitarsus Fairmaire, 1881
          Pericalus W.S.MacLeay, 1825
          Philophlaeus Chaudoir, 1844
          Phloeoxena Chaudoir, 1870
          Pristacrus Chaudoir, 1870
          Pseudoplatiella Baehr, 2012
          Serrimargo Chaudoir, 1870
          Sfitakantha Andrewes, 1919
          Sinurus Chaudoir, 1870
          Stenognathus Chaudoir, 1843
          Stenotelus Bouchard, 1903
          Stilboma Andrewes, 1933
          Stricklandiana Bousquet, 2002
          Tantillus Chaudoir, 1870
          Thyreochaetus Basilewsky, 1959
          Thyreopterus Dejean, 1831
          Thysanotus Chaudoir, 1848
          Trichocoptodera Louwerens, 1958
          Xanthos Kirschenhofer, 2003
          Xenitenopsis Basilewsky, 1956
          Xenitenus Péringuey, 1896
       Subtribe Physoderina Chaudoir, 1877
          Allocota Motschulsky, 1860
          Anchista Nietner, 1856
          Dasiosoma Britton, 1937
          Diamella Shi & Liang, 2013
          Endynomena Chaudoir, 1873
          Lachnoderma W.J.MacLeay, 1873
          Metallanchista Shi & Liang, 2013
          Orionella Jedlicka, 1964
          Paraphaea Bates, 1873
          Physodera Eschscholtz, 1829
       Subtribe Pseudotrechina Basilewsky, 1984
          Pseudotrechus Rosenhauer, 1856
       Subtribe Singilina Jeannel, 1949
          Singilis Rambur, 1837
       Subtribe Somotrichina Mateu, 1963
          Oecornis Britton, 1940
          Paulianites Jeannel, 1949
          Pephrica Alluaud, 1936
          Somotrichus Seidlitz, 1887
       Subtribe Sugimotoina Habu, 1975
          Sugimotoa Habu, 1975
       Subtribe Trichina Basilewsky, 1984
          Trichis Klug, 1832
       †Cymindoides Motschulsky, 1856
       †Lebina Germar, 1813
       †Miolebidia Zhang; Liu & Shangguan, 1989
       †Protoscalidion L.Schaufuss, 1888
    Tribe Odacanthini Laporte, 1834
       Subtribe Actenonycina Bates, 1871
          Actenonyx White, 1846
       Subtribe Homethina Liebherr, 2016
          Aeolodermus Andrewes, 1929
          Cyphocoleus Chaudoir, 1877
          Diplacanthogaster Liebke, 1932
          Homethes Newman, 1842
          Quammenis Erwin, 2000
          Stenocheila Laporte, 1832
       Subtribe Odacanthina Laporte, 1834
          Anasis Laporte, 1867
          Andrewesia Liebke, 1938
          Arame Andrewes, 1919
          Archicolliuris Liebke, 1931
          Asios Liebke, 1933
          Aulacolius Sloane, 1923
          Basistichus Sloane, 1917
          Clarencia Sloane, 1917
          Colliuris DeGeer, 1774
          Crassacantha Baehr, 1995
          Cryptocolliuris Basilewsky, 1955
          Deipyrodes Bousquet, 2002
          Dicraspeda Chaudoir, 1863
          Dobodura Darlington, 1968
          Erectocolliuris Liebke, 1931
          Eucolliuris Liebke, 1931
          Eudalia Laporte, 1867
          Gestroania Liebke, 1938
          Giachinoana Baehr, 2003
          Lachnothorax Motschulsky, 1862
          Lasiocera Dejean, 1831
          Mimocolliuris Liebke, 1933
          Myrmecodemus Sloane, 1923
          Neoeudalia Baehr, 2005
          Odacantha Paykull, 1798
          Ophionea Klug, 1821
          Polydamasium Liebke, 1938
          Porocara Sloane, 1917
          Protocolliuris Liebke, 1931
          Renneria Baehr, 1999
          Smeringocera Chaudoir, 1863
          Stenidia Brullé, 1834
          Tricharnhemia Baehr, 2009
       Subtribe Pentagonicina Bates, 1873
          Parascopodes Darlington, 1968
          Pentagonica Schmidt-Goebel, 1846
          Scopodes Erichson, 1842
    Tribe Perigonini G.Horn, 1881
       Diploharpus Chaudoir, 1850
       Mizotrechus Bates, 1872
       Perigona Laporte, 1835
       Ripogenites Basilewsky, 1954

### Subfamily Licininae Bonelli, 1810

    Tribe Chaetogenyini Emden, 1958
       Camptotoma Reiche, 1843
    Tribe Chlaeniini Brullé, 1834
       Subtribe Callistina Laporte, 1834
          Callistomimus Chaudoir, 1872
          Callistus Bonelli, 1810
       Subtribe Chlaeniina Brullé, 1834
          Actodus Alluaud, 1915
          Chlaenius Bonelli, 1810
          Eccoptomenus Chaudoir, 1850
          Ectenognathus Murray, 1858
          Globulipalpus Sciaky & Facchini, 2019
          Harpaglossus Motschulsky, 1858
          Hololeius LaFerté-Sénectère, 1851
          Holosoma Semenov, 1889
          Mirachlaenius Facchini, 2011
          Parachlaenius Kolbe, 1894
          Perissostomus Alluaud, 1930
          Procletodema Péringuey, 1898
          Procletus Péringuey, 1896
          Rhopalomelus Boheman, 1848
          Sphodroschema Alluaud, 1930
          Stenoodes Basilewsky, 1953
          Straneomelus Sciaky & Facchini, 2019
          Stuhlmannium Kolbe, 1894
          Viridagonum Lassalle, 2015
          †Rhopalochlaenius Zhang; Sun & Zhang, 1994
    Tribe Licinini Bonelli, 1810
       Subtribe Dicaelina Laporte, 1834
          Dicaelus Bonelli, 1813
          Diplocheila Brullé, 1835
       Subtribe Dicrochilina Ball, 1992
          Dicrochile Guérin-Méneville, 1846
       Subtribe Lestignathina Ball, 1992
          Atrotus Péringuey, 1896
          Dilonchus Andrewes, 1936
          Hormacrus Sloane, 1898
          Lacordairia Laporte, 1867
          Lestignathus Erichson, 1842
          Microferonia Blackburn, 1890
          Microzargus Sciaky & Facchini, 1997
          Platylytron W.J.MacLeay, 1873
          Siagonyx W.J.MacLeay, 1871
          Zargus Wollaston, 1854
       Subtribe Licinina Bonelli, 1810
          Badister Clairville, 1806
          Colpostoma Semenov, 1889
          Derostichus Motschulsky, 1860
          Eurygnathus Wollaston, 1854
          Eutogeneius Solier, 1849
          Licinus Latreille, 1802
          Omestes Andrewes, 1933
          Physolaesthus Chaudoir, 1850
    Tribe Oodini LaFerté-Sénectère, 1851
       Subtribe Dercylina Sloane, 1923
          Dercylus Laporte, 1832
       Subtribe Geobaenina Péringuey, 1896
          Geobaenus Dejean, 1829
       Subtribe Melanchitonina Jeannel, 1948
          Dicaelindus W.S.MacLeay, 1825
          Melanchiton Andrewes, 1940
          Melanchrous Andrewes, 1940
       Subtribe Oodina LaFerté-Sénectère, 1851
          Acanthoodes Basilewsky, 1953
          Acutosternus Lecordier & Girard, 1988
          Adelopomorpha Heller, 1916
          Anatrichis LeConte, 1853
          Bamaroodes Gueorguiev, 2014
          Brachyodes Jeannel, 1949
          Chaetocrepis Chaudoir, 1857
          Coptocarpus Chaudoir, 1857
          Dercylinus Chaudoir, 1883
          Evolenes LeConte, 1853
          Holcocoleus Chaudoir, 1883
          Hoplolenus LaFerté-Sénectère, 1851
          Lobatodes Basilewsky, 1956
          Lonchosternus LaFerté-Sénectère, 1851
          Macroprotus Chaudoir, 1878
          Megaloodes Lesne, 1896
          Microodes Jeannel, 1949
          Miltodes Andrewes, 1922
          Nanodiodes Bousquet, 1996
          Neoodes Basilewsky, 1953
          Nothoodes Gueorguiev & Liang, 2020
          Oodes Bonelli, 1810
          Oodinus Motschulsky, 1865
          Orthocerodus Basilewsky, 1946
          Polychaetus Chaudoir, 1882
          Prionognathus LaFerté-Sénectère, 1851
          Protopidius Basilewsky, 1949
          Pseudoodes Gueorguiev & Liang, 2020
          Pseudosphaerodes Jeannel, 1949
          Simous Chaudoir, 1882
          Sphaerodinus Jeannel, 1949
          Sphoerodes Chaudoir, 1883
          Stenocrepis Chaudoir, 1857
          Sundaoodes Gueorguiev & Liang, 2020
          Systolocranius Chaudoir, 1857
          Thaioodes Gueorguiev, 2014
          Thryptocerus Chaudoir, 1878
          Trichopalpoodes B.Gueorguiev & J.Schmidt, 2016

### Subfamily Loricerinae Bonelli, 1810

    Loricera Latreille, 1802

### Subfamily Melaeninae Csiki, 1933

    Cymbionotum Baudi di Selve, 1864
    Melaenus Dejean, 1831

### Subfamily Migadopinae Chaudoir, 1861

    Tribe Amarotypini Erwin, 1985
       Amarotypus Bates, 1872
       Migadopiella Baehr, 2009
    Tribe Migadopini Chaudoir, 1861
       Antarctonomus Chaudoir, 1861
       Aquilex Moret, 1989
       Calathosoma Jeannel, 1938
       Calyptogonia Sloane, 1920
       Decogmus Sloane, 1915
       Dendromigadops Baehr, 2013
       Lissopterus G.R.Waterhouse, 1843
       Loxomerus Chaudoir, 1842
       Migadopidius Jeannel, 1938
       Migadops G.R.Waterhouse, 1842
       Monolobus Solier, 1849
       Nebriosoma Laporte, 1867
       Pseudomigadops Jeannel, 1938
       Rhytidognathus Chaudoir, 1861
       Stichonotus Sloane, 1910
       Taenarthrus Broun, 1914

### Subfamily Nebriinae Laporte, 1834

    Tribe Cicindini Csiki, 1927
       Archaeocindis Kavanaugh & Erwin, 1991
       Cicindis Bruch, 1908
    Tribe Nebriini Laporte, 1834
       Leistus Frölich, 1799
       Nebria Latreille, 1802
       Nippononebria Ueno, 1955
       †Archaeonebria Kavanaugh & J.Schmidt, 2019
       †Ledouxnebria Deuve, 1998
    Tribe Notiokasiini Kavanaugh & Nègre, 1983
       Notiokasis Kavanaugh & Nègre, 1983
    Tribe Notiophilini Motschulsky, 1850
       Notiophilus Duméril, 1805
    Tribe Opisthiini Dupuis, 1912
       Opisthius Kirby, 1837
       Paropisthius Casey, 1920
    Tribe Pelophilini Kavanaugh, 1996
       Pelophila Dejean, 1821

### Subfamily Nototylinae Bänninger, 1927

    Nototylus Gemminger & Harold, 1868

### Subfamily Omophroninae Bonelli, 1810

    Omophron Latreille, 1802

### Subfamily Orthogoniinae Schaum, 1857

    Tribe Amorphomerini Sloane, 1923
       Amorphomerus Sloane, 1923
    Tribe Idiomorphini Bates, 1891
       Subtribe Idiomorphina Bates, 1891
          Idiomorphus Chaudoir, 1846
          Rathymus Dejean, 1831
          Strigia Brullé, 1835
       Subtribe Perochnoristhina Basilewsky, 1973
          Perochnoristhus Basilewsky, 1973
    Tribe Orthogoniini Schaum, 1857
       Subtribe Glyptina G.Horn, 1881
          Glyptus Brullé, 1837
          Neoglyptus Basilewsky, 1953
       Subtribe Orthogoniina Schaum, 1857
          Actenoncus Chaudoir, 1872
          Anoncopeucus Chaudoir, 1872
          Hexachaetus Chaudoir, 1872
          Neoorthogonius Tian & Deuve, 2006
          Nepalorthogonius Habu, 1979
          Orthogonius W.S.MacLeay, 1825

### Subfamily Panagaeinae Bonelli, 1810

    Tribe Brachygnathini Basilewsky, 1946
       Brachygnathus Perty, 1830
    Tribe Panagaeini Bonelli, 1810
       Subtribe Bascanina Basilewsky, 1953
          Bascanus Péringuey, 1896
       Subtribe Panagaeina Bonelli, 1810
          Adischissus Fedorenko, 2015
          Calathocosmus Emden, 1928
          Cintaroa Kasahara, 1989
          Cintaromorpha Häckel & Anichtchenko, 2016
          Coptia Brullé, 1835
          Craspedophorus Hope, 1838
          Dischissus Bates, 1873
          Epigraphodes Basilewsky, 1967
          Epigraphus Chaudoir, 1869
          Euschizomerus Chaudoir, 1850
          Geobius Dejean, 1831
          Micrixys LeConte, 1854
          Microschemus Andrewes, 1940
          Panagaeus Latreille, 1802
          Paregraphus Basilewsky, 1967
          Peronomerus Schaum, 1854
          Psecadius Alluaud, 1911
          Tefflus Leach in Samouelle, 1819
          Tinoderus Chaudoir, 1879
          Tinognathus Chaudoir, 1879
          Trichisia Motschulsky, 1865
    Tribe Peleciini Chaudoir, 1880
       Subtribe Agonicina Sloane, 1920
          Agonica Sloane, 1920
          Pseudagonica B.Moore, 1960
       Subtribe Peleciina Chaudoir, 1880
          Ardistomopsis Straneo & Ball, 1989
          Disphericus G.R.Waterhouse, 1842
          Dyschiridium Chaudoir, 1861
          Eripus Dejean, 1829
          Pelecium Kirby, 1819
          Stricteripus Straneo & Ball, 1989

### Subfamily Patrobinae Kirby, 1837

    Tribe Lissopogonini Zamotajlov, 2000
       Lissopogonus Andrewes, 1923
    Tribe Patrobini Kirby, 1837
       Subtribe Deltomerina Chaudoir, 1872
          Deltomerus Motschulsky, 1850
       Subtribe Deltomerodina Zamotajlov, 2002
          Deltomerodes Deuve, 1992
       Subtribe Patrobina Kirby, 1837
          Apatrobus Habu & Baba, 1960
          Apenetretus Kurnakov, 1960
          Archipatrobus Zamotajlov, 1992
          Caelopenetretus Zamotajlov & Ito, 2000
          Chaetapatrobus Lafer, 1996
          Chinapenetretus Kurnakov, 1963
          Dimorphopatrobus Casale & Sciaky, 1994
          Diplous Motschulsky, 1850
          Grandipenetretus Zamotajlov & Sciaky, 1999
          Himalopenetretus Zamotajlov, 2002
          Indopatrobus Zamotajlov & Wrase, 2006
          Ledouxius Zamotajlov, 1992
          Minipenetretus Zamotajlov, 2002
          Minypatrobus Ueno, 1955
          Naxipenetretus Zamotajlov, 1999
          Parapatrobus Zamotajlov, 1992
          Parapenetretus Kurnakov, 1960
          Patanitretus Zamotajlov, 2002
          Patrobus Dejean, 1821
          Penetretus Motschulsky, 1865
          Platidiolus Chaudoir, 1878
          Platypatrobus Darlington, 1938
          Prodiplous Zamotajlov & Sciaky, 2006
          Qiangopatrobus Zamotajlov, 2002
          Quasipenetretus Zamotajlov, 2002
          Tibetopenetretus Zamotajlov & Sciaky, 1999

### Subfamily Paussinae Latreille, 1806

    Tribe Metriini LeConte, 1853
       Subtribe †Kryzhanovskianina Deuve, 2019
          †Kryzhanovskiana Kataev & Kirejtshuk, 2019
       Metrius Eschscholtz, 1829
       Sinometrius Wrase & J.Schmidt, 2006
    Tribe Ozaenini Hope, 1838
       Subtribe Microzaenina Deuve, 2019
          Microzaena Fairmaire, 1901
       Subtribe Mystropomina G.Horn, 1881
          Mystropomus Chaudoir, 1848
       Subtribe Ozaenina Hope, 1838
          Crepidozaena Deuve, 2001
          Entomoantyx Ball & McCleve, 1990
          Filicerozaena Deuve, 2001
          Gibbozaena Deuve, 2001
          Goniotropis G.R.Gray, 1831
          Inflatozaena Deuve, 2001
          Mimozaena Deuve, 2001
          Ozaena Olivier, 1812
          Pachyteles Perty, 1830
          Physea Brullé, 1835
          Physeomorpha Ogueta, 1963
          Platycerozaena Bänninger, 1927
          Proozaena Deuve, 2001
          Serratozaena Deuve, 2001
          Tachypeles Deuve, 2001
          Tropopsis Solier, 1849
       Subtribe Pseudozaenina Sloane, 1905
          Anentmetus Andrewes, 1924
          Dhanya Andrewes, 1919
          Eustra Schmidt-Goebel, 1846
          Itamus Schmidt-Goebel, 1846
          Pseudozaena Laporte, 1834
          Sphaerostylus Chaudoir, 1848
    Tribe Paussini Latreille, 1806
       Subtribe Carabidomemnina Wasmann, 1928
          Carabidomemnus Kolbe, 1924
          Eohomopterus Wasmann, 1919
       Subtribe Cerapterina Billberg, 1820
          Arthropterus W.S.MacLeay, 1838
          Cerapterus Swederus, 1788
          Megalopaussus Lea, 1906
          Mesarthropterus Wasmann, 1926
       Subtribe Heteropaussina Janssens, 1950
          Heteropaussus J.Thomson, 1860
       Subtribe Homopterina Wasmann, 1920
          Homopterus Westwood, 1841
       Subtribe Paussina Latreille, 1806
          Allorhopalus Nagel, 2017
          Ceratoderus Westwood, 1841
          Euplatyrhopalus Desneux, 1905
          Lebioderus Westwood, 1838
          Leleupaussus Luna de Carvalho, 1962
          Melanospilus Westwood, 1847
          Paussomorphus Raffray, 1885
          Paussus Linnaeus, 1775
          Platyrhopalides Wasmann, 1918
          Platyrhopalopsis Desneux, 1905
          Platyrhopalus Westwood, 1833
          Pterorhopalus Maruyama, 2011
       Subtribe Pentaplatarthrina Jeannel, 1946
          Hexaplatarthrus Jeannel, 1955
          Pentaplatarthrus Westwood, 1833
       Subtribe †Arthropteritina Luna de Carvalho, 1961
          †Arthropterites Wasmann, 1926
          †Cerapterites Wasmann, 1926
          †Protocerapterus Wasmann, 1926
       Subtribe †Eopaussina Luna de Carvalho, 1951
          †Eopaussus Wasmann, 1926
    Tribe Protopaussini Gestro, 1892
       Protopaussus Gestro, 1892

### Subfamily Platyninae Bonelli, 1810

    Tribe Omphreini Ganglbauer, 1891
       Omphreus Dejean, 1828
    Tribe Platynini Bonelli, 1810
       Subtribe Enoicina Basilewsky, 1985
          Abacetodes Straneo, 1939
          Enoicus Péringuey, 1896
       Subtribe Platynina Bonelli, 1810
          Achaetocephala Habu, 1975
          Achaetoprothorax Habu, 1978
          Aepsera Chaudoir, 1874
          Agelaea Gené, 1839
          Agonidium Jeannel, 1948
          Agonobembix Jeannel, 1948
          Agonoriascus Basilewsky, 1985
          Agonorites Jeannel, 1951
          Agonum Bonelli, 1810
          Altagonum Darlington, 1952
          Anchomenus Bonelli, 1810
          Andinocolpodes Perrault, 1991
          Andrewesius Andrewes, 1939
          Aparupa Andrewes, 1930
          Archagonum Basilewsky, 1953
          Archicolpodes J.Schmidt, 2001
          Arhytinus Bates, 1889
          Atranodes Jedlicka, 1953
          Atranus LeConte, 1847
          Austroglyptolenus Roig-Juñent, 2003
          Beckeria Jedlicka, 1931
          Blackburnia Sharp, 1878
          Bothrocolpodes Basilewsky, 1985
          Bruskespar Morvan, 1998
          Bruskmoal Morvan, 1998
          Bryanites Valentine, 1987
          Callidagonum Lorenz, 1998
          Cardiomera Bassi, 1834
          Catacolpodes Basilewsky, 1985
          Celaenagonum Habu, 1978
          Chaetagonum Burgeon, 1933
          Chaetosaurus J.Schmidt, 2001
          Cinctagonum Baehr, 2012
          Cistelagonum Baehr, 2012
          Collagonum Baehr, 1995
          Colpodes W.S.MacLeay, 1825
          Colpoides Jedlicka, 1931
          Colpomimus Basilewsky, 1985
          Colposphodrus Jedlicka, 1953
          Ctenognathus Fairmaire, 1843
          Cymenopterus Jeannel, 1948
          Cyrtopilus Basilewsky, 1985
          Dalatagonum Fedorenko, 2011
          Deliaesianum Morvan, 1999
          Deltocolpodes Morvan, 1992
          Dendragonum Jeannel, 1948
          Diacanthostylus Habu, 1978
          Dicranoncoides Habu, 1978
          Dicranoncus Chaudoir, 1850
          Dinocolpodes J.Schmidt, 2001
          Dirotus W.S.MacLeay, 1825
          Dister Morvan, 2006
          Dolichocolpodes Basilewsky, 1985
          Dyscolus Dejean, 1831
          Elliptoleus Bates, 1882
          Epicolpodes Basilewsky, 1985
          Eucolpodes Jeannel, 1948
          Euleptus Klug, 1833
          Euplynes Schmidt-Goebel, 1846
          Feroniascus Jeannel, 1951
          Fortagonum Darlington, 1952
          Galiciotyphlotes Assmann, 1999
          Gastragonum Darlington, 1952
          Glaucagonum Habu, 1978
          Glyptolenoides Perrault, 1991
          Glyptolenus Bates, 1878
          Gyrochaetostylus Habu, 1978
          Habragonum Ueno, 1964
          Hannaphota Landin, 1955
          Haplocolpodes Jeannel, 1951
          Haplopeza Boheman, 1848
          Helluocolpodes Liebherr, 2005
          Hemiplatynus Casey, 1920
          Henvelik Morvan, 1999
          Herculagonum Baehr, 2002
          Hikosanoagonum Habu, 1954
          Idiagonum Darlington, 1952
          Idiastes Andrewes, 1931
          Idiocolpodes Basilewsky, 1985
          Incagonum Liebherr, 1994
          Iridagonum Darlington, 1952
          Ischnagonum Kasahara & Satô, 1997
          Jocqueius Basilewsky, 1988
          Jujiroa Ueno, 1952
          Kalchdigor Morvan, 1999
          Kar Morvan, 1998
          Karnes Morvan, 2010
          Kaszabellus Jedlicka, 1954
          Kiwiplatynus Larochelle & Larivière, 2021
          Klapperichella Jedlicka, 1956
          Kuceraianum Morvan, 2002
          Kupeplatynus Larochelle & Larivière, 2021
          Laevagonum Darlington, 1952
          Lassalleianum Morvan, 1999
          Lepcha Andrewes, 1930
          Leptagonum Kolbe, 1897
          Leptocolpodes Basilewsky, 1985
          Letouzeya Bruneau de Miré, 1982
          Liagonum Jeannel, 1948
          Liamegalonychus Basilewsky, 1963
          Liocolpodes Basilewsky, 1985
          Lissagonum Habu, 1978
          Lithagonum Darlington, 1952
          Lobocolpodes Basilewsky, 1985
          Lorostema Motschulsky, 1865
          Loxocrepis Eschscholtz, 1829
          Lucicolpodes J.Schmidt, 2000
          Maculagonum Darlington, 1952
          Maoriplatynus Larochelle & Larivière, 2021
          Meleagros Kirschenhofer, 1999
          Mesocolpodes Basilewsky, 1985
          Metacolpodes Jeannel, 1948
          Mexisphodrus Barr, 1965
          Montagonum Darlington, 1952
          Mooreagonum Baehr, 2016
          Morimotoidius Habu, 1954
          Nebriagonum Darlington, 1952
          Negreum Habu, 1958
          Neobatenus Jeannel, 1948
          Neocolpodes Jeannel, 1948
          Neodendragonum Basilewsky, 1953
          Neomegalonychus Jeannel, 1948
          Nesiocolpodes Jeannel, 1948
          Nipponagonum Habu, 1978
          Notagonum Darlington, 1952
          Notocolpodes Basilewsky, 1985
          Notoplatynus B.Moore, 1985
          Nymphagonum Habu, 1978
          Olisthopus Dejean, 1828
          Oncostylus Habu, 1978
          Onotokiba Alluaud, 1927
          Onycholabis Bates, 1873
          Onypterygia Dejean, 1831
          Orophicus Alluaud, 1925
          Orthotrichus Peyron, 1856
          Oxygonium Basilewsky, 1951
          Oxypselaphus Chaudoir, 1843
          Pachybatenus Basilewsky, 1973
          Pachyferonia Jeannel, 1951
          Paracolpodes Basilewsky, 1985
          Paraliagonum Basilewsky, 1957
          Paramegalonychus Basilewsky, 1953
          Paranchodemus Habu, 1978
          Paranchus Lindroth, 1974
          Paraplatynus Baehr, 2016
          Pawgammm Morvan, 1999
          Platyagonum Habu, 1978
          Platynus Bonelli, 1810
          Plaumannium Liebke, 1939
          Plicagonum Darlington, 1952
          Ponapagonum Darlington, 1970
          Potamagonum Darlington, 1952
          Praepristus Kirschenhofer, 1999
          Promecoptera Dejean, 1831
          Promegalonychus Basilewsky, 1953
          Prophenorites Basilewsky, 1985
          Prosphodrus Britton, 1959
          Protocolpodes Basilewsky, 1985
          Pseudanchomenus Tarnier, 1860
          Pseudobatenus Basilewsky, 1951
          Pseudomegalonychus Basilewsky, 1950
          Rhadine LeConte, 1846
          Rupa Jedlicka, 1935
          Scotagonum Habu, 1978
          Sericoda Kirby, 1837
          Shibataia Habu, 1978
          Sinocolpodes J.Schmidt, 2001
          Skoeda Morvan, 1995
          Skorlagad Morvan, 1999
          Skouedhirraad Morvan, 1999
          Sophroferonia Alluaud, 1933
          Speagonum B.Moore, 1977
          Speocolpodes Barr, 1974
          Speokokosia Alluaud, 1932
          Sperkanhir Morvan, 2010
          Stenocnemus Mannerheim, 1837
          Straneoa Basilewsky, 1953
          Syletor Tschitscherine, 1899
          Takasagoagonum Habu, 1977
          Tanystoma Motschulsky, 1845
          Tarsagonum Darlington, 1952
          Tetraleucus Casey, 1920
          Tostkar Morvan, 1998
          Trogloagonum Casale, 1982
          Tuiplatynus Larochelle & Larivière, 2021
          Vitagonum B.Moore, 1999
          Vulcanophilus Heller, 1898
          Xestagonum Habu, 1978
          †Praeanchodemus J.Schmidt; Göpel & Will, 2017
    Tribe Sphodrini Laporte, 1834
       Subtribe Atranopsina Baehr, 1982
          Amaroschema Jeannel, 1943
          Amphimasoreus Piochard de la Brûlerie, 1875
          Atranopsis Baehr, 1982
          Broter Andrewes, 1923
          Gomerina Bolivar y Pieltain, 1940
          Paraeutrichopus Mateu, 1954
          Platyderus Stephens, 1827
          Pseudomyas Uyttenboogaart, 1929
          Pseudoplatyderus Bolivar y Pieltain, 1940
       Subtribe Calathina Laporte, 1834
          Calathus Bonelli, 1810
          Thermoscelis Putzeys, 1873
       Subtribe Dolichina Brullé, 1834
          Acalathus Semenov, 1889
          Anchomenidius Heyden, 1880
          Casaleius Sciaky & Wrase, 1998
          Dolichus Bonelli, 1810
          Doliodactyla Sciaky & Wrase, 1998
          Morphodactyla Semenov, 1889
          Xestopus Andrewes, 1937
       Subtribe Pristosiina Lindroth, 1956
          Pristosia Motschulsky, 1865
       Subtribe Sphodrina Laporte, 1834
          Calathidius Putzeys, 1873
          Cephalosdrophus Lassalle & Marcilhac, 1999
          Eosphodrus Casale, 1988
          Eremosphodrus Semenov, 1909
          Himalosphodrus Casale, 1988
          Hystricosphodrus Casale & Giachino, 2004
          Ifridytes Deuve & Queinnec, 1994
          Laemostenus Bonelli, 1810
          Licinopsis Bedel, 1899
          Miquihuana Barr, 1982
          Pseudotaphoxenus L.Schaufuss, 1865
          Reflexisphodrus Casale, 1988
          Sphodropsis Seidlitz, 1887
          Sphodrus Clairville, 1806
          Stenolepta Semenov, 1889
          Taphoxenus Motschulsky, 1850
       Subtribe Synuchina Lindroth, 1956
          Nipponosynuchus Morita, 1998
          Parabroscus Lindroth, 1956
          Synuchus Gyllenhal, 1810
          Trephionus Bates, 1883

### Subfamily Promecognathinae LeConte, 1853

    Tribe Axinidiini Basilewsky, 1963
       Axinidium Sturm, 1843
       Holaxinidium Basilewsky, 1963
       Metaxinidium Basilewsky, 1963
       Paraxinidium Basilewsky, 1963
    Tribe Dalyatini Mateu, 2002
       Dalyat Mateu, 2002
    Tribe Promecognathini LeConte, 1853
       Promecognathus Chaudoir, 1846
    Tribe †Palaeoaxinidiini McKay, 1991
       †Palaeoaxinidium McKay, 1991

### Subfamily Pseudomorphinae Hope, 1838

    Adelotopus Hope, 1836
    Cainogenion Notman, 1925
    Cryptocephalomorpha Ritsema, 1875
    Guyanemorpha Erwin, 2013
    Manumorpha Erwin & Geraci, 2008
    Notopseudomorpha Baehr, 1997
    Paussotropus C.O.Waterhouse, 1877
    Pseudomorpha Kirby, 1823
    Samiriamorpha Erwin & Geraci, 2008
    Sphallomorpha Westwood, 1840
    Tuxtlamorpha Erwin & Geraci, 2008
    Yasunimorpha Erwin & Geraci, 2008

### Subfamily Psydrinae LeConte, 1853

    Tribe Gehringiini Darlington, 1933
       Subtribe Gehringiina Darlington, 1933
          Gehringia Darlington, 1933
       Subtribe Helenaeina Deuve, 2007
          Afrogehringia Baehr; Schüle & Lorenz, 2009
          Helenaea Schatzmayr & Koch, 1934
    Tribe Moriomorphini Sloane, 1890
       Subtribe Amblytelina Blackburn, 1892
          Amblytelus Erichson, 1842
          Dystrichothorax Blackburn, 1892
          Epelyx Blackburn, 1892
          Mecyclothorax Sharp, 1903
          Meonis Laporte, 1867
          Paratrichothorax Baehr, 2004
          Pseudamblytelus Baehr, 2004
          Raphetis B.Moore, 1963
          Selenochilus Chaudoir, 1878
          Trichamblytelus Baehr, 2004
       Subtribe Moriomorphina Sloane, 1890
          Celanida Laporte, 1867
          Melisodera Westwood, 1835
          Molopsida White, 1846
          Moriodema Laporte, 1867
          Moriomorpha Laporte, 1867
          Neonomius B.Moore, 1963
          Pterogmus Sloane, 1920
          Rhaebolestes Sloane, 1903
          Rossjoycea Liebherr, 2011
          Sitaphe B.Moore, 1963
          Spherita Liebherr, 2020
          Tarastethus Sharp, 1883
          Teraphis Laporte, 1867
          Theprisa B.Moore, 1963
          Trephisa B.Moore, 1963
          Trichopsida Larochelle & Larivière, 2013
       Subtribe Tropopterina Sloane, 1898
          Pharetis Liebherr, 2020
          Tropopterus Solier, 1849
    Tribe Psydrini LeConte, 1853
       Laccocenus Sloane, 1890
       Nomius Laporte, 1835
       Psydrus LeConte, 1846

### Subfamily Pterostichinae Bonelli, 1810

    Tribe Chaetodactylini Tschitscherine, 1903
       Chaetodactyla Tschitscherine, 1897
    Tribe Cnemalobini Germain, 1911
       Cnemalobus Guérin-Méneville, 1838
    Tribe Cratocerini Lacordaire, 1854
       Subtribe Catapieseina Bates, 1882
          Catapiesis Solier, 1835
          Homalomorpha Brullé, 1837
       Subtribe Cratocerina Lacordaire, 1854
          Brachidius Chaudoir, 1852
          Cratocerus Dejean, 1829
          Oxyglychus Straneo, 1938
       Subtribe Drimostomatina Chaudoir, 1873
          Abacaelostus Straneo, 1952
          Andrewesinulus Straneo, 1938
          Apsidocnemus Alluaud, 1936
          Barylaus Liebherr, 1985
          Caecocaelus Straneo, 1949
          Caelostomus W.S.MacLeay, 1825
          Camptogenys Tschitscherine, 1899
          Capabatus Csiki, 1930
          Crenulostrigus Straneo, 1942
          Cyrtolaus Bates, 1882
          Dactyleurys Tschitscherine, 1899
          Dactylinius Straneo, 1941
          Diachipteryx Alluaud, 1925
          Diceromerus Chaudoir, 1873
          Dromistomus Jeannel, 1948
          Drymonaxus Straneo, 1941
          Feostoma Straneo, 1941
          Hemitelestus Alluaud, 1895
          Hoplizomenus Chaudoir, 1873
          Leleuporites Straneo, 1960
          Madapelmus Dajoz, 1985
          Monodryxus Straneo, 1942
          Pachycaecus Straneo, 1971
          Pachyroxochus Straneo, 1942
          Platyxythrius Lorenz, 1998
          Stegazopteryx Will, 2004
          Stomonaxellus Tschitscherine, 1901
          Strigomerodes Straneo, 1939
          Strigomerus Chaudoir, 1873
          Trichillinus Straneo, 1938
    Tribe Microcheilini Jeannel, 1948
       Microcheila Brullé, 1835
    Tribe Morionini Brullé, 1837
       Buderes Murray, 1857
       Hyperectenus Alluaud, 1935
       Hyperion Laporte, 1834
       Megamorio Chaudoir, 1880
       Morion Latreille, 1810
       Morionidius Chaudoir, 1880
       Moriosomus Motschulsky, 1855
       Platynodes Westwood, 1847
       Stereostoma Murray, 1857
    Tribe Pterostichini Bonelli, 1810
       Subtribe Abacetina Chaudoir, 1873
          Abacaecus Allegro & Giachino, 2020
          Abacetus Dejean, 1828
          Aristopus LaFerté-Sénectère, 1853
          Celioinkosa Straneo, 1951
          Chlaeminus Motschulsky, 1865
          Colpodichius Straneo, 1952
          Cosmodiscus Sloane, 1907
          Cyrtomoscelis Chaudoir, 1874
          Distrigidius Jeannel, 1948
          Ecnomolaus Bates, 1892
          Haptoderidius Straneo, 1975
          Haptoderodes Straneo, 1986
          Holconotus Schmidt-Goebel, 1846
          Inkosa Péringuey, 1926
          Mateuellus Deuve, 1990
          Metabacetus Bates, 1892
          Metaxellus Straneo, 1960
          Metaxys Chaudoir, 1857
          Novillidius Straneo, 1941
          Oodinkosa Straneo, 1939
          Ophonichius Straneo, 1942
          Pediomorphus Chaudoir, 1878
          Pioprosopus Tschitscherine, 1899
          Pollicobius Vinson, 1939
          Prostalomus Basilewsky, 1950
          Pseudabacetus Burgeon, 1935
          Pterostillichus Straneo, 1949
          Rhagadillius Straneo, 1951
          Tiferonia Darlington, 1962
          Trachelocyphoides Straneo, 1942
          Trachelocyphus Tschitscherine, 1900
       Subtribe Euchroina Chaudoir, 1874
          Abacillius Straneo, 1949
          Abacillodes Straneo, 1988
          Abaris Dejean, 1831
          Apsaustodon Tschitscherine, 1901
          Argutoridius Chaudoir, 1876
          Blennidus Motschulsky, 1866
          Blenniventer Straneo, 1986
          Bothynoproctus Tschitscherine, 1900
          Cedrorum Borges & A.Serrano, 1993
          Cephalostichus Straneo, 1977
          Cynthidia Chaudoir, 1874
          Euchroa Brullé, 1835
          Eumara Tschitscherine, 1901
          Gastrogmus Sloane, 1915
          Haplobothynus Tschitscherine, 1901
          Litarthrum Sloane, 1915
          Lobobrachus Sharp, 1885
          Marsyas Putzeys, 1845
          Meropalpus Tschitscherine, 1900
          Microcephalus Dejean, 1828
          Neotalus Will, 2002
          Oribazus Chaudoir, 1874
          Orthomus Chaudoir, 1838
          Pachythecus Chaudoir, 1874
          Paniestichus Will, 2011
          Parorthomus B.Gueorguiev; Wrase & Farkac, 2014
          Phaenaulax Tschitscherine, 1898
          Prosopogmus Chaudoir, 1865
          Pseudabarys Chaudoir, 1874
          Setalimorphus Sloane, 1895
          Setalis Laporte, 1867
          Simodontus Chaudoir, 1843
          Trirammatus Chaudoir, 1835
       Subtribe Loxandrina Erwin & Sims, 1984
          Cerabilia Laporte, 1867
          Oxycrepis Reiche, 1843
          Zeodera Laporte, 1867
       Subtribe Metiina Straneo, 1951
          Abropus G.R.Waterhouse, 1842
          Antarctiola Straneo, 1977
          Feroniola Tschitscherine, 1900
          Kuschelinus Straneo, 1963
          Metius Curtis, 1839
       Subtribe Pterostichina Bonelli, 1810
          Abacidus LeConte, 1863
          Abacoleptus Fauvel, 1903
          Abacomorphus Chaudoir, 1878
          Abacophrastus Will, 2011
          Abacops Tschitscherine, 1902
          Abax Bonelli, 1810
          Allotriopus Bates, 1882
          Analoma Darlington, 1971
          Apophylon B.Gueorguiev & Sciaky, 2015
          Aristochroa Tschitscherine, 1898
          Aristochroodes Marcilhac, 1993
          Aulacopodus Britton, 1940
          Basilewskya Straneo, 1948
          Camptoscelis Dejean, 1828
          Castelnaudia Tschitscherine, 1891
          Catadromus W.S.MacLeay, 1825
          Chaetauchenium Tschitscherine, 1900
          Chalcochrous Chaudoir, 1838
          Cophosomorpha Tschitscherine, 1891
          Cratoferonia Tschitscherine, 1902
          Cratogaster Blanchard, 1843
          Cuneipectus Sloane, 1907
          Cyclotrachelus Chaudoir, 1838
          Cyrtoderus Hope, 1842
          Darodilia Laporte, 1867
          Delinius Westwood, 1864
          Eucamptognathus Chaudoir, 1837
          Eudromus Klug, 1835
          Euryabax Fauvel, 1903
          Euryaptus Bates, 1892
          Eurypercus Jeannel, 1948
          Eurystomis Chaudoir, 1878
          Gastrellarius Casey, 1918
          Gourlayia Britton, 1964
          Harpostomus Chaudoir, 1856
          Henrotiochoromus Busulini, 1958
          Henrotius Jeannel, 1953
          Holcaspis Chaudoir, 1865
          Hybothecus Chaudoir, 1874
          Leiolesticus Roux; Lassalle & Dubault, 2016
          Lesticus Dejean, 1828
          Licentius Jedlicka, 1939
          Liopasa Tschitscherine, 1901
          Lophoglossus LeConte, 1853
          Loxogenius Sloane, 1907
          Mecynognathus W.J.MacLeay, 1873
          Megadromus Motschulsky, 1866
          Molopidius Jeannel, 1942
          Molopinus Jeannel, 1948
          Molops Bonelli, 1810
          Myas Sturm, 1826
          Neoferonia Britton, 1940
          Nesites Andrewes, 1939
          Nirmala Andrewes, 1930
          Notabax B.Moore, 1976
          Notolestus Sloane, 1894
          Notonomus Chaudoir, 1865
          Nurus Motschulsky, 1866
          Oberthueria Vuillet, 1911
          Ogmophora Tschitscherine, 1896
          Onawea Johns, 2007
          Oscadytes Lagar Mascaro, 1975
          Pachymorphus Chaudoir, 1838
          Paranurus Tschitscherine, 1901
          Pareuryaptus Dubault; Lassalle & Roux, 2008
          Parhypates Motschulsky, 1866
          Pedius Motschulsky, 1850
          Percolaus Bates, 1882
          Percus Bonelli, 1810
          Peyrieraselus Deuve, 1981
          Piesmus LeConte, 1846
          Platycoelus Blanchard, 1843
          Platysmodes Fauvel, 1903
          Plocamostethus Britton, 1940
          Poecilinus Jeannel, 1948
          Poecilus Bonelli, 1810
          Pseudoceneus Tschitscherine, 1891
          Pterostichus Bonelli, 1810
          Rhabdotus Chaudoir, 1865
          Rhytiferonia Darlington, 1962
          Rhytisternus Chaudoir, 1865
          Sarticus Motschulsky, 1866
          Secatophus Laporte, 1867
          Setalidius Chaudoir, 1878
          Speluncarius Reitter, 1886
          Speomolops Patrizi, 1955
          Sphodrosomus Perroud, 1864
          Stenochoromus L.Miller, 1866
          Stereocerus Kirby, 1837
          Sthenocranion Tschitscherine, 1899
          Stomis Clairville, 1806
          Straneostichus Sciaky, 1994
          Styracoderus Chaudoir, 1874
          Tanythrix Schaum, 1858
          Taphoxomimus Straneo, 1975
          Tapinopterus Schaum, 1858
          Teratotarsa Tschitscherine, 1893
          Teropha Laporte, 1867
          Trichosternus Chaudoir, 1865
          Trigonaptus Fedorenko, 2020
          Trigonotoma Dejean, 1828
          Troglorites Jeannel, 1919
          Tropidocerus Chaudoir, 1878
          Typhlochoromus Moczarski, 1913
          Wahlbergiana Bousquet, 2002
          Xenion Tschitscherine, 1902
          Zariquieya Jeannel, 1924
          Zeopoecilus Sharp, 1886
    Tribe Zabrini Bonelli, 1810
       Subtribe Amarina Zimmermann, 1832
          Amara Bonelli, 1810
          Pseudamara Lindroth, 1968
       Subtribe Zabrina Bonelli, 1810
          Zabrus Clairville, 1806

### Subfamily Rhysodinae Laporte, 1840

    Tribe Clinidiini R.T. & J.R.Bell, 1978
       Clinidium Kirby, 1830
       Grouvellina R.T. & J.R.Bell, 1978
       Rhyzodiastes Fairmaire, 1895
    Tribe Dhysorini R.T. & J.R.Bell, 1978
       Dhysores Grouvelle, 1903
       Neodhysores R.T. & J.R.Bell, 1978
       Tangarona R.T. & J.R.Bell, 1982
    Tribe Leoglymmiini R.T. & J.R.Bell, 1978
       Leoglymmius R.T. & J.R.Bell, 1978
    Tribe Medisorini R.T. & J.R.Bell, 1987
       Medisores R.T. & J.R.Bell, 1987
    Tribe Omoglymmiini R.T. & J.R.Bell, 1978
       Arrowina R.T. & J.R.Bell, 1978
       Omoglymmius Ganglbauer, 1891
       Plesioglymmius R.T. & J.R.Bell, 1978
       Shyrodes Grouvelle, 1903
       Srimara R.T. & J.R.Bell, 1978
       Xhosores R.T. & J.R.Bell, 1978
       Yamatosa R.T. & J.R.Bell, 1979
    Tribe Rhysodini Laporte, 1840
       Kaveinga R.T. & J.R.Bell, 1978
       Kupeus R.T. & J.R.Bell, 1982
       Rhysodes Germar, 1822
    Tribe Sloanoglymmiini R.T. & J.R.Bell, 1991
       Sloanoglymmius R.T. & J.R.Bell, 1991

### Subfamily Scaritinae Bonelli, 1810

    Tribe Clivinini Rafinesque, 1815
       Subtribe Androzelmina R.T.Bell, 1998
          Androzelma Dostal, 1993
       Subtribe Ardistomina Putzeys, 1867
          Ardistomis Putzeys, 1846
          Aspidoglossa Putzeys, 1846
          Kearophus Dajoz, 2004
          Semiardistomis Kult, 1950
       Subtribe Clivinina Rafinesque, 1815
          Afrosyleter Basilewsky, 1959
          Ancus Putzeys, 1867
          Basilewskyana Kult, 1959
          Brachypelus Putzeys, 1867
          Cameroniola Baehr, 2000
          Climax Putzeys, 1861
          Clivina Latreille, 1802
          Clivinarchus Sloane, 1896
          Cryptomma Putzeys, 1846
          Lachenus Putzeys, 1846
          Leleuporella Basilewsky, 1956
          Nannoryctes Baehr, 2000
          Nyctosyles Putzeys, 1867
          Orictites Andrewes, 1931
          Platysphyrus Sloane, 1905
          Psilidius Jeannel, 1957
          Pyramoides Bousquet, 2002
          Rhysocara Sloane, 1916
          Rubidiclivina Baehr, 2015
          Rugiluclivina Balkenohl, 1996
          Semiclivina Kult, 1947
          Sinesetosa Balkenohl, 1996
          Syleter Andrewes, 1941
          Thliboclivina Kult, 1959
          Trilophidius Jeannel, 1957
          Trilophus Andrewes, 1927
          Trogloclivina Deuve, 2003
          Whiteheadiana Perrault, 1994
       Subtribe Forcipatorina Bänninger, 1938
          Camptidius Putzeys, 1867
          Camptodontus Dejean, 1826
          Forcipator Maindron, 1904
          Kultianella Perrault, 1994
          Mesus Chevrolat, 1858
          Obadius Burmeister, 1875
          Oxygnathopsis Louwerens, 1953
          Oxygnathus Dejean, 1826
          Scolyptus Putzeys, 1861
          Stratiotes Putzeys, 1846
       Subtribe Reicheiina Jeannel, 1957
          Alpiodytes Jeannel, 1957
          Asioreicheia Bulirsch & Magrini, 2014
          Catalanodytes Sciaky, 1989
          Dalmatoreicheia Magrini & Bulirsch, 2005
          Dimorphoreicheia Magrini; Fancello & Leo, 2002
          Galicioreicheia Felix & Bulirsch, 2015
          Guiodytes Tian, 2013
          Gymnetoreicheia Magrini; Fancello & Casale, 2019
          Iberodytes Jeannel, 1949
          Ichnusodytes Magrini; Fancello & Onnis, 2019
          Italodytes G.Müller, 1938
          Kenyoreicheia Bulirsch & Magrini, 2007
          Laoreicheia Balkenohl, 2005
          Madagascareicheia Magrini & Bulirsch, 2009
          Malagasyreicheia Magrini; Bulirsch & Fancello, 2021
          Orientoreicheia Bulirsch & Hurka, 1994
          Oxydrepanus Putzeys, 1867
          Parareicheia Jeannel, 1957
          Reicheadella Reitter, 1913
          Reicheia Saulcy, 1862
          Reicheidius Jeannel, 1957
          Sikelioreicheia Magrini & Fancello, 2019
          Spelaeodytes L.Miller, 1863
          Typhloreicheia Holdhaus, 1924
       Subtribe Schizogeniina Dostal, 2017
          Baehrogenius Dostal, 2017
          Coryza Putzeys, 1867
          Halocoryza Alluaud, 1919
          Lophocoryza Alluaud, 1941
          Paracoryza Basilewsky, 1952
          Psammocoryza Hogan, 2006
          Schizogenius Putzeys, 1846
       Subtribe Sparostesina Dostal, 2017
          Bohemaniella Bousquet, 2002
          Pseudoclivina Kult, 1947
          Sparostes Putzeys, 1867
    Tribe Corintascarini Basilewsky, 1973
       Corintascaris Basilewsky, 1952
    Tribe Dyschiriini Kolbe, 1880
       Akephorus LeConte, 1852
       Caledyschirius Bulirsch, 2010
       Clivinopsis Bedel, 1895
       Cribrodyschirius Bruneau de Miré, 1952
       Dyschirius Bonelli, 1810
       Neodyschirius Kult, 1954
       Reicheiodes Ganglbauer, 1891
       Setodyschirius Fedorenko, 1996
       Striganoviella Fedorenko, 2012
       Torretassoa Schatzmayr & Koch, 1933
       †Dyschiriomimus Iablokoff-Khnzorian, 1960
    Tribe Salcediini Alluaud, 1930
       Subtribe Salcediina Alluaud, 1930
          Salcedia Fairmaire, 1899
       Subtribe Solenogenyina R.T.Bell, 1998
          Holoprizus Putzeys, 1867
          Solenogenys Westwood, 1859
    Tribe Scaritini Bonelli, 1810
       Subtribe Carenina W.J.MacLeay, 1887
          Carenidium Chaudoir, 1868
          Carenum Bonelli, 1813
          Epilectus Blackburn, 1888
          Euryscaphus W.J.MacLeay, 1865
          Laccopterum W.J.MacLeay, 1878
          Monocentrum Chaudoir, 1868
          Mouhotia Laporte, 1862
          Neocarenum Laporte, 1867
          Neoscaphus Sloane, 1888
          Philoscaphus W.J.MacLeay, 1871
          Scaraphites Westwood, 1842
          Trichocarenum Blackburn, 1892
       Subtribe Pasimachina Putzeys, 1867
          Pasimachus Bonelli, 1813
       Subtribe Scapterina Putzeys, 1867
          Acanthoscelis Dejean, 1825
          Oxylobus Chaudoir, 1855
          Parathlibops Basilewsky, 1958
          Passalidius Chaudoir, 1863
          Scapterus Dejean, 1826
          Steganomma W.J.MacLeay, 1887
          Thlibops Putzeys, 1867
       Subtribe Scaritina Bonelli, 1810
          Anomophaenus Fauvel, 1882
          Antilliscaris Lorenz, 1998
          Baenningeria Reichardt, 1976
          Coptolobus Chaudoir, 1857
          Crepidopterus Chaudoir, 1855
          Cryptoscaphus Chaudoir, 1855
          Dinoscaris Alluaud, 1902
          Distichus Motschulsky, 1858
          Dyscaris Bänninger, 1940
          Dyscherinus Jeannel, 1955
          Dyscherus Chaudoir, 1855
          Geoscaptus Chaudoir, 1855
          Glyptogrus Bates, 1875
          Gnaphon Andrewes, 1920
          Haplogaster Chaudoir, 1879
          Haplotrachelus Chaudoir, 1855
          Macromorphus Chaudoir, 1857
          Madascaris Bänninger, 1938
          Mamboicus Bates, 1886
          Mecynoscaris Alluaud, 1930
          Menigius Chaudoir, 1879
          Neochryopus Bänninger, 1932
          Ochyropus Schiödte, 1847
          Pachyodontus Chaudoir, 1879
          Paradyscherus Basilewsky, 1971
          Pilades Heyne, 1895
          Prodyscherodes Jeannel, 1955
          Prodyscherus Jeannel, 1946
          Scarites Fabricius, 1775
          Storthodontus Chaudoir, 1855
          Tapinoscaris Jeannel, 1946
          Tibioscarites Bänninger, 1929
          Tonkinoscaris Bänninger, 1956
          Typhloscaris Kuntzen, 1914

### Subfamily Siagoninae Bonelli, 1813

    Tribe Enceladini G.Horn, 1881
       Enceladus Bonelli, 1813
    Tribe Siagonini Bonelli, 1813
       Luperca Laporte, 1840
       Siagona Latreille, 1804

### Subfamily Trechinae Bonelli, 1810

    Tribe Bembidarenini Maddison et al., 2019
       Andinodontis Erwin & Toledano, 2010
       Argentinatachoides Sallanave; Erwin & Roig-Juñent, 2008
       Bembidarenas Erwin, 1972
       Tasmanitachoides Erwin, 1972
    Tribe Bembidiini Stephens, 1827
       Subtribe Anillina Jeannel, 1937
          Afranillus Giachino, 2015
          Afrodipnus Giachino, 2015
          Angustanillus Baehr & Main, 2016
          Anillaspis Casey, 1918
          Anillinus Casey, 1918
          Anillodes Jeannel, 1963
          Anillopsidius Coiffait, 1969
          Anillopsis Jeannel, 1937
          Anillotarsus Mateu, 1980
          Anillus Jacquelin du Val, 1851
          Argiloborus Jeannel, 1937
          Austranillus Giachino, 2005
          Bafutyphlus Bruneau de Miré, 1986
          Bhutanillus M.E.Schmid, 1975
          Bulirschia Giachino, 2008
          Bylibaraphanus Giachino; Eberhard & Perina, 2021
          Caeconannus Jeannel, 1963
          Caecoparvus Jeannel, 1937
          Carayonites Bruneau de Miré, 1986
          Cryptocharidius M.Etonti & Mateu, 1992
          Cryptorites Jeannel, 1950
          Dicropterus Ehlers, 1883
          Elgonotyphlus Sciaky & Zaballos, 1993
          Erwinanillus Giachino; Eberhard & Perina, 2021
          Externanillus Baehr & Main, 2016
          Geocharidius Jeannel, 1963
          Geocharis Ehlers, 1883
          Gilesdytes Giachino; Eberhard & Perina, 2021
          Gracilanillus Baehr & Main, 2016
          Gregorydytes Giachino; Eberhard & Perina, 2021
          Hesperanillus Baehr & Main, 2016
          Honduranillus Zaballos, 1997
          Hygranillus B.Moore, 1980
          Hypodipnites Jeannel, 1963
          Hypotyphlus Jeannel, 1937
          Iason Giachino & Vailati, 2011
          Iberanillus Español, 1971
          Illaphanus W.J.MacLeay, 1865
          Kimberleytyphlus Giachino; Eberhard & Perina, 2021
          Leleupanillus Basilewsky, 1976
          Magnanillus Baehr, 2017
          Malagasydipnus Giachino, 2008
          Malagasytyphlus Giachino, 2008
          Megastylulus Giachino & Sciaky, 2003
          Mexanillus Vigna Taglianti, 1973
          Microdipnites Jeannel, 1957
          Microdipnodes Basilewsky, 1960
          Microdipnus Jeannel, 1937
          Microtyphlus Linder, 1863
          Mystroceridius Reichardt, 1972
          Neodipnus Jeannel, 1957
          Neoillaphanus Giachino; Eberhard & Perina, 2021
          Nesamblyops Jeannel, 1937
          Nothanillus Jeannel, 1962
          Orthotyphlus Mateu & Zaballos, 1998
          Paranillopsis Cicchino & Roig-Juñent, 2001
          Paranillus Jeannel, 1949
          Parillaphanus Baehr, 2018
          Parvocaecus Coiffait, 1956
          Pelocharis Jeannel, 1960
          Pelodiaetodes B.Moore, 1980
          Pelodiaetus Jeannel, 1937
          Pelonomites Jeannel, 1963
          Perucharidius Mateu & M.Etonti, 2002
          Pilbaradytes Giachino; Eberhard & Perina, 2021
          Pilbaranillus Baehr & Main, 2016
          Pilbaraphanus Giachino; Eberhard & Perina, 2021
          Prioniomus Jeannel, 1937
          Pseudanillus Bedel, 1896
          Pseudillaphanus Giachino, 2005
          Rhegmatobius Jeannel, 1937
          Scotodipnus Schaum, 1860
          Selenodipnus Jeannel, 1963
          Serranillus Barr, 1995
          Serratotyphlus Giachino, 2008
          Stylulus L.Schaufuss, 1882
          Tasmanillus Giachino, 2005
          Typhlocharis Dieck, 1869
          Typhlonesiotes Jeannel, 1937
          Winklerites Jeannel, 1937
          Zapotecanillus Sokolov, 2013
          Zeanillus Jeannel, 1937
          Zoianillus Sciaky, 1994
       Subtribe Bembidiina Stephens, 1827
          Amerizus Chaudoir, 1868
          Asaphidion Gozis, 1886
          Bembidion Latreille, 1802
          Caecidium Ueno, 1971
          Lionepha Casey, 1918
          Ocys Stephens, 1828
          Orzolina Machado, 1987
          Sinechostictus Motschulsky, 1864
       Subtribe Horologionina Jeannel, 1949
          Horologion Valentine, 1932
       Subtribe Lovriciina Giachino; B.Gueorguiev & Vailati, 2011
          Lovricia Pretner, 1979
          Neolovricia Lakota; Jalzic & J.Moravec, 2009
          Paralovricia Giachino; B.Gueorguiev & Vailati, 2011
       Subtribe Tachyina Motschulsky, 1862
          Aenigmatachys Baehr, 2020
          Afrotachys Basilewsky, 1958
          Anomotachys Jeannel, 1946
          Costitachys Erwin, 1974
          Elaphropus Motschulsky, 1839
          Kiwitachys Larochelle & Larivière, 2007
          Liotachys Bates, 1871
          Lymnastis Motschulsky, 1862
          Meotachys Erwin, 1974
          Micratopus Casey, 1914
          Moirainpa Erwin, 1984
          Nothoderis Boyd & Erwin, 2016
          Pericompsus LeConte, 1852
          Polyderidius Jeannel, 1962
          Porotachys Netolitzky, 1914
          Setitachys Baehr, 2016
          Stigmatachys Boyd & Erwin, 2016
          Straneoites Basilewsky, 1947
          Tachys Dejean, 1821
          Tachysbembix Erwin, 2004
          Tachyta Kirby, 1837
          Tachyxysta Boyd & Erwin, 2016
       Subtribe Xystosomina Erwin, 1994
          Erwiniana Paulsen & A.Smith, 2003
          Geballusa Erwin, 1994
          Gouleta Erwin, 1994
          Inpa Erwin, 1978
          Mioptachys Bates, 1882
          Philipis Erwin, 1994
          Xystosomus Schaum, 1863
    Tribe Pogonini Laporte, 1834
       Bedeliolus Semenov, 1900
       Cardiaderus Dejean, 1828
       Diodercarus Lutshnik, 1931
       Diplochaetus Chaudoir, 1872
       Ochtozetus Chaudoir, 1872
       Olegius Komarov, 1996
       Pogonistes Chaudoir, 1872
       Pogonopsis Bedel, 1898
       Pogonus Dejean, 1821
       Sirdenus Dejean, 1828
       Syrdenoidius Baehr & Hudson, 2001
       Thalassotrechus Van Dyke, 1918
    Tribe Sinozolini Deuve, 1997
       Chaltenia Roig-Juñent & Cicchino, 2001
       Phrypeus Casey, 1924
       Sinozolus Deuve, 1997
    Tribe Trechini Bonelli, 1810
       Subtribe Aepina Fowler, 1887
          Aepopsis Jeannel, 1922
          Aepus Leach in Samouelle, 1819
          Kenodactylus Broun, 1909
          Kiwitrechus Larochelle & Larivière, 2007
          Maoritrechus Brookes, 1932
          Neanops Britton, 1962
          Oarotrechus Townsend, 2010
          Temnostega Enderlein, 1905
          Thalassobius Solier, 1849
          Waiputrechus Townsend, 2010
       Subtribe Perileptina Sloane, 1903
          Apoplotrechus Alluaud, 1915
          Neoblemus Jeannel, 1923
          Perileptus Schaum, 1860
       Subtribe Trechina Bonelli, 1810
          Accoella Ueno, 1990
          Acheroniotes Lohaj & Lakota, 2010
          Adriaphaenops Noesske, 1928
          Aepiblemus Belousov & Kabak, 1993
          Agonotrechus Jeannel, 1923
          Agostinia Jeannel, 1928
          Alanorites Belousov, 1998
          Albanotrechus Casale & V.B.Gueorguiev, 1994
          Allegrettia Jeannel, 1928
          Allotrechiama Ueno, 1970
          Ameroduvalius Valentine, 1952
          Anchotrechus Jeannel, 1927
          Andinorites Mateu & Belles, 1980
          Andinotrechus Mateu, 1981
          Anillidius Jeannel, 1928
          Anophthalmus Sturm, 1844
          Aphaenopidius G.Müller, 1913
          Aphaenops Bonvouloir, 1862
          Aphaenopsis G.Müller, 1913
          Apocimmerites Belousov, 1998
          Apoduvalius Jeannel, 1953
          Aputrechisibus Trezzi, 2007
          Arctaphaenops Meixner, 1925
          Aspidaphaenops Ueno, 2006
          Austrotrechus B.Moore, 1972
          Awatrechus Ueno, 1955
          Balazucellus Deuve, 2001
          Bathytrechus Ueno, 2005
          Beronaphaenops B.V.Gueorguiev, 2012
          Birmaphaenops Deuve, 2017
          Blemus Dejean, 1821
          Boldoriella Jeannel, 1928
          Boreaphaenops Ueno, 2002
          Bothynotrechus B.Moore, 1972
          Casaleaphaenops Tian; Huang & Ma, 2021
          Cathaiaphaenops Deuve, 1996
          Caucasaphaenops Belousov, 1999
          Caucasorites Belousov & Zamotajlov, 1997
          Chaetoduvalius Jeannel, 1928
          Chaetotrechiama Ueno, 1982
          Chiapadytes Vigna Taglianti, 1977
          Chu Tian & He, 2020
          Cimbrodytes Piva, 2021
          Cimmerites Jeannel, 1928
          Cimmeritodes Deuve, 1996
          Ckacus Tian & Huang, 2021
          Columbitrechus Mateu, 1982
          Coreoblemus Ueno, 1969
          Croatotrechus Casale & Jalzic, 1999
          Dabatrechus Ueno, 2004
          Dactylotrechus Belousov & Kabak, 2003
          Daiconotrechus Ueno, 1971
          Dalmataphaenops Monguzzi, 1993
          Darlingtonea Valentine, 1952
          Derossiella Queinnec, 2008
          Deuveaphaenops Tian & Huang, 2017
          Dianotrechus Tian, 2016
          Doderotrechus Vigna Taglianti, 1968
          Dongoblemus Deuve & Tian, 2016
          Dongodytes Deuve, 1993
          Dracotrechus Ueno, 2010
          Duvalioblemus Deuve, 1995
          Duvaliomimus Jeannel, 1928
          Duvaliopsis Jeannel, 1928
          Duvalius Delarouzée, 1859
          Eocnides Jeannel, 1954
          Epaphiopsis Ueno, 1953
          Epaphiotrechus Deuve & Kavanaugh, 2016
          Epaphius Leach in Samouelle, 1819
          Erebotrechus Britton, 1964
          Escolatrechus Mateu, 2003
          Eutrechopsis B.Moore, 1972
          Eutrechus B.Moore, 1972
          Geotrechus Jeannel, 1919
          Giraffaphaenops Deuve, 2002
          Glabroduvalius Vrbica; Curcic; Antic & Curcic, 2013
          Goedetrechus B.Moore, 1972
          Gotoblemus Ueno, 1970
          Graciliblemus Deuve & Tian, 2016
          Guatemalatrechus Trezzi, 2003
          Guiaphaenops Deuve, 2002
          Guizhaphaenops Vigna Taglianti, 1997
          Guizhaphaenopsoides Tian; Huang & Li, 2021
          Gulaphaenops Ueno, 1987
          Himalaphaenops Ueno, 1980
          Himalotrechus Belousov; Kabak & J.Schmidt, 2019
          Homaloderodes Jeannel, 1962
          Huoyanodytes Tian & Huang, 2016
          Iberotrechus Jeannel, 1920
          Iga Ueno, 1953
          Incatrechus Mateu & Belles, 1982
          Inotrechus Dolzhanski & Ljovushkin, 1989
          Ishidatrechus Ueno, 1956
          Ishikawatrechus Habu, 1950
          Italaphaenops Ghidini, 1964
          Iyotrechus Ueno & Naito, 2009
          Jalzicaphaenops Lohaj & Lakota, 2010
          Jeannelius Kurnakov, 1959
          Jiangxiaphaenops Ueno & Clarke, 2007
          Jiulongotrechus Tian; Huang & Wang, 2015
          Junaphaenops Ueno, 1997
          Junnanotrechus Ueno & Yin, 1993
          Kettlotrechus Townsend, 2010
          Kosswigia Jeannel, 1947
          Kozlovites Jeannel, 1935
          Kupetrechus Larochelle & Larivière, 2007
          Kurasawatrechus Yoshida & Nomura, 1952
          Kusumia Ueno, 1952
          Lanxangaphaenops Deuve, 2012
          Laoblemus Ueno, 2006
          Laosaphaenops Deuve, 2000
          Lessinodytes Vigna Taglianti, 1982
          Libotrechus Ueno, 1998
          Luoxiaotrechus Tian & Yin, 2013
          Luyatrechus M.Etonti & Mateu, 2000
          Luzonotrechus Ueno, 1979
          Mamesdytes Trezzi, 2003
          Masuzoa Ueno, 1960
          Masuzonoblemus Ueno, 1989
          Mayaphaenops Vigna Taglianti, 1977
          Meganophthalmus Kurnakov, 1959
          Mexaphaenops Bolivar y Pieltain, 1942
          Mexitrechus Barr, 1982
          Miaotrechus Tian; Chen & Ma, 2020
          Microblemus Ueno, 2007
          Mimanillus B.Moore, 1972
          Mimotrechus B.Moore, 1972
          Minimaphaenops Deuve, 2000
          Minosaphaenops Queinnec, 2008
          Minutotrechus Deuve & Kavanaugh, 2016
          Nannotrechus Winkler, 1926
          Nanotrechiama Belousov & Kabak, 2018
          Neaphaenops Jeannel, 1920
          Nelsonites Valentine, 1952
          Neotrechus G.Müller, 1913
          Nesaeoduvalius Casale & Giachino, 2016
          Nesiotrechus Ueno, 1995
          Nipponaphaenops Ueno, 1971
          Nototrechus B.Moore, 1972
          Nunbergites Pawlowski & Stachowiak, 1991
          Omalodera Blanchard, 1842
          Oodinotrechus Ueno, 1998
          Oroblemites Ueno & Pawlowski, 1981
          Oroblemus Ueno & Yoshida, 1966
          Orotrechus G.Müller, 1913
          Oxytrechus Jeannel, 1927
          Panaphaenops Tian; Huang & Ma, 2021
          Paracimmerites Belousov, 1998
          Paraphaenops Jeannel, 1916
          Paratrechiotes Ueno, 1995
          Paratrechus Jeannel, 1920
          Petraphaenops Delic; Kapla & Colla, 2019
          Pheggomisetes Knirsch, 1923
          Pilosaphaenops Deuve & Tian, 2008
          Plesioaphaenops Deuve & Tian, 2011
          Pogonoschema Jeannel, 1927
          Pontodytes Casale & Giachino, 1989
          Porocimmerites Belousov, 1998
          Protrechiama Belousov & Kabak, 2003
          Pseudanophthalmus Jeannel, 1920
          Pseudaphaenops Winkler, 1912
          Pseudocnides Jeannel, 1927
          Pseudotrechisibus Mateu & Belles, 1982
          Puertrechus Belousov & Kabak, 2014
          Putzeysius Jeannel, 1962
          Qianaphaenops Ueno, 2000
          Qianotrechus Ueno, 2000
          Queinnectrechus Deuve, 1992
          Rakantrechus Ueno, 1951
          Ryugadous Habu, 1950
          Sardaphaenops Cerruti & Henrot, 1956
          Satotrechus Ueno, 2006
          Sbordoniella Vigna Taglianti, 1980
          Scotoplanetes Absolon, 1913
          Scototrechus Britton, 1962
          Shenaphaenops Ueno, 1999
          Shennongotrechus Ueno, 2004
          Shenoblemus Tian & Fang, 2020
          Shilinotrechus Ueno, 2003
          Shiqianaphaenops Tian, 2016
          Shuangheaphaenops Tian, 2017
          Shuaphaenops Ueno, 1999
          Sichuanotrechus Deuve, 2005
          Sidublemus Tian & Yin, 2013
          Sinaphaenops Ueno & Wang, 1991
          Sinocimmerites Deuve, 2007
          Sinotrechiama Ueno, 2000
          Sinotroglodytes Deuve, 1996
          Sloanella Jeannel, 1927
          Speotrechus Jeannel, 1922
          Stevensius Jeannel, 1923
          Stygiotrechus Ueno, 1958
          Subilsia Español, 1967
          Sumatrechus Deuve, 2005
          Superbotrechus Deuve & Tian, 2009
          Suzuka Ueno, 1956
          Taiwanotrechus Ueno, 1987
          Taniatrechus Belousov & Dolzhanski, 1994
          Tasmanorites Jeannel, 1927
          Tasmanotrechus B.Moore, 1972
          Taurocimmerites Belousov, 1998
          Thalassoduvalius Ueno, 1956
          Tianeotrechus Tian & Tang, 2016
          Tianzhuaphaenops Zhao & Tian, 2016
          Tienmutrechus Suenson, 1957
          Tonkinaphaenops Deuve, 2013
          Toshiaphaenops Ueno, 1999
          Trechepaphiama Deuve & Kavanaugh, 2016
          Trechepaphiopsis Deuve & Kavanaugh, 2016
          Trechiama Jeannel, 1927
          Trechiamiotes Deuve, 1998
          Trechicomimus Mateu & Nègre, 1972
          Trechiella Jeannel, 1927
          Trechimorphus Jeannel, 1927
          Trechinotus Jeannel, 1962
          Trechiotes Jeannel, 1954
          Trechisibus Motschulsky, 1862
          Trechistus B.Moore, 1972
          Trechoblemus Ganglbauer, 1891
          Trechus Clairville, 1806
          Trichaphaenops Jeannel, 1916
          Troglocimmerites Ljovuschkin, 1970
          Tropidotrechus Jeannel, 1927
          Typhlotrechus G.Müller, 1913
          Uenoaphaenops Tian & He, 2020
          Uenoites Belousov & Kabak, 2016
          Uenotrechus Deuve & Tian, 1999
          Ushijimaella Ueno, 1980
          Velebitaphaenops Casale & Jalzic, 2012
          Velesaphaenops S.Curcic et al., 2018
          Vietotrechus Ueno, 1995
          Wanhuaphaenops Tian & Wang, 2016
          Wanoblemus Tian & Fang, 2016
          Wulongoblemus Ueno, 2007
          Xenotrechus Barr & Krekeler, 1967
          Xiangxius Tian; Huang & Li, 2021
          Xuedytes Tian & Huang, 2017
          Yalongaphaenops Belousov & Kabak, 2021
          Yamautidius Ueno, 1957
          Yanzaphaenops Ueno, 2010
          Yunotrechus Tian & Huang, 2014
          Zhijinaphaenops Ueno & Ran, 2002
       Subtribe Trechodina Jeannel, 1926
          Amblystogenium Enderlein, 1905
          Canarobius Machado, 1987
          Cnides Motschulsky, 1862
          Cothresia Jeannel, 1964
          Cyphotrechodes Jeannel, 1926
          Eotrechodes Ueno; Lafer & Sundukov, 1995
          Himalotrechodes Ueno, 1981
          Iberotrechodes Faille et al., 2021
          Pachydesus Motschulsky, 1864
          Pachytrechodes Jeannel, 1960
          Paratrechodes Jeannel, 1926
          Scaurotrechodes Geginat, 2006
          Spelaeovulcania Machado, 1987
          Sporades Fauvel, 1882
          Thalassophilus Wollaston, 1854
          Trechobembix Jeannel, 1926
          Trechodes Blackburn, 1901
          Trechosia Jeannel, 1926
          Trechosiella Jeannel, 1960
       †Antarctotrechus Ashworth & Erwin, 2016
       †Trechinites Heer, 1862
       †Trechoides Motschulsky, 1856
    Tribe Zolini Sharp, 1886
       Subtribe Merizodontina Sloane, 1920
          Idacarabus Lea, 1910
          Maungazolus Larochelle & Larivière, 2017
          Merizodus Solier, 1849
          Percodermus Sloane, 1920
          Pseudoopterus Csiki, 1928
          Pterocyrtus Sloane, 1920
          Sloaneana Csiki, 1933
          Synteratus Broun, 1909
          Thayerella Baehr, 2016
       Subtribe Zolina Sharp, 1886
          Oopterus Guérin-Méneville, 1841
          Zolus Sharp, 1886

### Subfamily Xenaroswellianinae Erwin, 2007

    Xenaroswelliana Erwin, 2007

### Subfamily †Conjunctiinae Ponomarenko, 1977

    †Conjunctia Ponomarenko, 1977

### Subfamily †Protorabinae Ponomarenko, 1977

    †Aethocarabus Ren; Lu & Guo, 1995
    †Atrirabus Hong & Wang, 1990
    †Cordorabus Ponomarenko, 1977
    †Cretorabus Ponomarenko, 1977
    †Denudirabus Ren; Lu & Guo, 1995
    †Lirabus Hong, 1992
    †Lithorabus Ponomarenko, 1977
    †Magnirabus Hong & Wang, 1990
    †Mesorabus Ponomarenko, 1977
    †Nebrorabus Ponomarenko, 1989
    †Ovrabites Ponomarenko, 1977
    †Penecupes Ren, 1995
    †Protorabus Ponomarenko, 1977
    †Tica Hong, 1992

#### Not Assigned to Subfamily

    †Agatoides Motschulsky, 1856
    †Amphoxyne Bode, 1953
    †Carabites Heer, 1852
    †Cavicarabus Hong, 1991
    †Conexicoxa Lin, 1986
    †Cymatopterus Lomnicki, 1894
    †Fangshania Hong, 1981
    †Glenopterus Heer, 1847
    †Hebeicarabus Hong, 1983
    †Megacarabus Hong, 1983
    †Meileyingia Hong, 1987
    †Miocarabus Hong, 1983
    †Neothanes Scudder, 1890
    †Procarabus Oppenheim, 1888
    †Prosynactus Bode, 1953
    †Shanwangicarabus Hong, 1985
    †Sinis Heer, 1862
    †Sinocalosoma Hong & Wang, 1986
    †Sinocaralosoma Hong, 1984
    †Sunocarabus Hong, 1987
    †Tauredon Handlirsch, 1910
    †Wuchangicarabus Hong, 1991
    †Xishanocarabus Hong, 1984
    †Yunnanocarabus Lin, 1977