# Holothyrida

The Holothyrida are a small order of mites in the superorder Parasitiformes. No fossils are known. With body lengths of more than 2 mm (3⁄32 in) they are relatively large mites, with a heavily sclerotized body. It is divided into three families, Allothyridae, Holothyridae, and Neothyridae. In a 1998 experimental study, members of the family Allothyridae were found to ignore living animals but readily fed on the body fluids of dead arthropods, making them scavengers.
The order has a distribution largely confined to former Gondwanan landmasses. They are the sister group to Ixodida (ticks).

## Systematics
### Allothyridae

Allothyridae van der Hammen, 1972 — Australia, New Zealand
* Allothyrus van der Hammen, 1961
   * Allothyrus australasiae (Womersley, 1935)
   * Allothyrus constrictus (Domrow, 1955)
* Australothyrus van der Hammen, 1983
   * Australothyrus ocellatus van der Hammen, 1983

### Holothyridae

Holothyridae Thorell, 1882 Sri Lanka, Indian Ocean islands, New Guinea, New Caledonia
* Sternothyrus Lehtinen, 1995
** Sternothyrus braueri (Thon, 1905) — Seychelles
* Lindothyrus Lehtinen, 1995
** Lindothyrus elongatus Lehtinen, 1995 — Lord Howe Island
** Lindothyrus rubellus Lehtinen, 1995 — New Caledonia
* Indothyrus Lehtinen, 1995
** Indothyrus greeni Lehtinen, 1995 — Sri Lanka
* Haplothyrus Lehtinen, 1995
** Haplothyrus expolitissimus (Berlese, 1924) — New Caledonia
** Haplothyrus hyatti Lehtinen, 1995 — unknown locality
* Holothyrus Gervais, 1842 — Mauritius
** Holothyrus coccinella Gervais, 1842
** Holothyrus legendrei Hammen, 1983
* Hammenius Lehtinen, 1981
** Hammenius armatus (Canestrini, 1897) — Tamara Island (Aitape): New Guinea
** Hammenius berlesei (Lehtinen, 1995) — New Guinea
** Hammenius braueri (Thon, 1906)
** Hammenius fujuge Lehtinen, 1981 — New Guinea (Central District, Oro Province)
** Hammenius grandjeani (Hammen, 1961) — Mount Bosavi: New Guinea
** Hammenius holthuisi van der Hammen, 1983
** Hammenius ingii Lehtinen, 1981 — New Guinea
** Hammenius insularis Lehtinen, 1995 — Louisiade Archipelago: New Guinea
** Hammenius longipes (Thorell, 1882) — Fly River, New Guinea (?)
** Hammenius mendi (Lehtinen, 1995) — Strickland River: New Guinea
** Hammenius montanus Hammen, 1983 — Irian Jaya
** Hammenius niger (Thon, 1906)

### Neothyridae

Neothyridae Lehtinen, 1981 Northern South America and the Caribbean
* Diplothyrus Lehtinen, 1999 Brazil, French Guiana
** Diplothyrus schubarti Lehtinen, 1999
** Diplothyrus lecorrei Klompen 2010
** Diplothyrus lehtineni Vázquez & de Araújo & Feres 2016
* Neothyrus Lehtinen, 1981 Peru
** Neothyrus ana Lehtinen, 1981
* Caribothyrus Kontschán & Mahunka 2004 Dominican Republic
** Caribothyrus barbatus Kontschán & Mahunka 2004

## Footnotes

* ↑ Walter, David Evans; Proctor, Heather C. (1998). "Feeding behaviour and phylogeny: observations on early derivative Acari". Experimental and Applied Acarology. 22 (1): 39–50. doi:10.1023/A:1006033407957.
* ↑ Dobson, Susan J.; Barker, Stephen C. (1999). "Phylogeny of the Hard Ticks (Ixodidae) Inferred from 18S rRNA Indicates That the GenusAponommaIs Paraphyletic". Molecular Phylogenetics and Evolution. 11 (2): 288–295. doi:10.1006/mpev.1998.0565.