# Harpalus (beetle)

Harpalus is a genus of ground beetle first described by Pierre André Latreille in 1802.

## Species

Harpalus contains the following 430 species:

### Subgenus Cephalophonus Ganglbauer, 1891

    Harpalus cephalotes Fairmaire & Laboulbène, 1854

### Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982

    Harpalus agnatus Reiche, 1850
    Harpalus cyrenaicus Koch, 1939
    Harpalus fulvus Dejean, 1829
    Harpalus grilli Kataev, 2002
    Harpalus idiotus Bates, 1889
    Harpalus janinae Jeanne, 1984
    Harpalus litigiosus Dejean, 1829
    Harpalus melancholicus Dejean, 1829
    Harpalus schaumii Wollaston, 1864
    Harpalus tenebrosus Dejean, 1829

### Subgenus Glanodes Casey, 1914

    Harpalus cohni Ball, 1972
    Harpalus corpulentus (Casey, 1914)
    Harpalus huachuca Ball, 1972
    Harpalus obliquus G.Horn, 1880
    Harpalus puncticeps (Casey, 1914)
    Harpalus stephani Ball, 1972

### Subgenus Harpalus Latreille, 1802

    Harpalus acupalpoides Reitter, 1900
    Harpalus aeneipennis (Faldermann, 1836)
    Harpalus aequicollis Motschulsky, 1844
    Harpalus affinis (Schrank, 1781)
    Harpalus agakhaniantzi (Mikhailov, 1972)
    Harpalus akbensis Jedlicka, 1958
    Harpalus akinini Tschitscherine, 1895
    Harpalus alajensis Tschitscherine, 1898
    Harpalus albanicus Reitter, 1900
    Harpalus alexandrae Kataev, 1990
    Harpalus alexeevi Kataev, 1990
    Harpalus alpivagus Tschitscherine, 1899
    Harpalus amarellus Bates, 1891
    Harpalus amariformis Motschulsky, 1844
    Harpalus amplicollis Ménétriés, 1848
    Harpalus amputatus Say, 1830
    Harpalus anatolicus Tschitscherine, 1898
    Harpalus angulatus Putzeys, 1878
    Harpalus angustipennis Boheman, 1848
    Harpalus angustitarsis Reitter, 1887
    Harpalus animosus Casey, 1924
    Harpalus anisodactyliformis Solsky, 1874
    Harpalus antonowi (Tschitscherine, 1898)
    Harpalus anxioides Kataev, 1991
    Harpalus anxius (Duftschmid, 1812)
    Harpalus araraticus Mlynar, 1979
    Harpalus arcuatus Tschitscherine, 1898
    Harpalus arnoldii Kataev, 1988
    Harpalus ascetes Kataev & Wrase, 1997
    Harpalus asemus Basilewsky, 1946
    Harpalus aterrimus Casey, 1914
    Harpalus atratus Latreille, 1804
    Harpalus atrichatus Hatch, 1949
    Harpalus atripes Casey, 1914
    Harpalus attenuatus Stephens, 1828
    Harpalus australasiae Dejean, 1829
    Harpalus autumnalis (Duftschmid, 1812)
    Harpalus badakschanus Jedlicka, 1956
    Harpalus baleensis Clarke, 1973
    Harpalus balli Noonan, 1991
    Harpalus basanicus J.Sahlberg, 1913
    Harpalus basilewskyi Facchini, 2003
    Harpalus basuto Basilewsky, 1958
    Harpalus bellieri Reiche, 1861
    Harpalus beneshi Kataev & Wrase, 1997
    Harpalus brachypterus Tschitscherine, 1898
    Harpalus brevicornis Germar, 1823
    Harpalus brevis Motschulsky, 1844
    Harpalus breviusculus Chaudoir, 1846
    Harpalus bucharicus Tschitscherine, 1898
    Harpalus bulirschi (Facchini, 2012)
    Harpalus bungii Chaudoir, 1844
    Harpalus caeruleatus Bates, 1878
    Harpalus caiphus Reiche & Saulcy, 1855
    Harpalus calathoides Motschulsky, 1844
    Harpalus capicola Dejean, 1829
    Harpalus cardoni Antoine, 1922
    Harpalus caspius (Steven, 1806)
    Harpalus cautus Dejean, 1829
    Harpalus chalcentus Bates, 1873
    Harpalus chasanensis Lafer, 1989
    Harpalus chobautianus Lutshnik, 1922
    Harpalus chrysopus Reitter, 1887
    Harpalus circumpunctatus Chaudoir, 1846
    Harpalus cisteloides Motschulsky, 1844
    Harpalus clarkei Kataev & J.Schmidt, 2020
    Harpalus compressus Motschulsky, 1844
    Harpalus contemptus Dejean, 1829
    Harpalus convexus Faldermann, 1836
    Harpalus corporosus (Motschulsky, 1862)
    Harpalus corrugatus Basilewsky, 1958
    Harpalus crates Bates, 1883
    Harpalus cupreus Dejean, 1829
    Harpalus cursorius Péringuey, 1896
    Harpalus cyanopterus (Tschitscherine, 1897)
    Harpalus cyclogonus Chaudoir, 1844
    Harpalus danieli Reitter, 1900
    Harpalus davidianus Tschitscherine in Jakobson, 1903
    Harpalus decipiens Dejean, 1829
    Harpalus defector Péringuey, 1896
    Harpalus desertus LeConte, 1859
    Harpalus diligens Tschitscherine, 1898
    Harpalus dimidiatus (P.Rossi, 1790)
    Harpalus dispar Dejean, 1829
    Harpalus dissitus Antoine, 1931
    Harpalus distinguendus (Duftschmid, 1812)
    Harpalus diversicollis Basilewsky, 1958
    Harpalus dubius Boheman, 1848
    Harpalus dudkoi Kataev, 2011
    Harpalus durangoensis Bates, 1891
    Harpalus ebeninus Heyden, 1870
    Harpalus egenus Dejean, 1829
    Harpalus egorovi Lafer, 1989
    Harpalus ellipsis LeConte, 1847
    Harpalus elliptipennis Facchini, 2003
    Harpalus erosus Mannerheim, 1825
    Harpalus exiguus Boheman, 1848
    Harpalus famelicus Tschitscherine, 1898
    Harpalus farkaci Kataev & Wrase, 1995
    Harpalus fimetarius Dejean, 1829
    Harpalus flavescens (Piller & Mitterpacher, 1783)
    Harpalus flavicornis Dejean, 1829
    Harpalus foveiger Tschitscherine, 1895
    Harpalus franzi Mateu, 1954
    Harpalus frater Chaudoir, 1876
    Harpalus fraternus LeConte, 1852
    Harpalus froelichii Sturm, 1818
    Harpalus fulvicornis Thunberg, 1806
    Harpalus fulvilabris Mannerheim, 1853
    Harpalus fulvipennis Chaudoir, 1843
    Harpalus fuscicornis Ménétriés, 1832
    Harpalus fuscipalpis Sturm, 1818
    Harpalus fuscipennis Wiedemann, 1825
    Harpalus fuscoaeneus Dejean, 1829
    Harpalus ganssuensis Semenov, 1889
    Harpalus giacomazzoi Kataev & Wrase, 1996
    Harpalus gilgil Basilewsky, 1946
    Harpalus gisellae Csiki, 1932
    Harpalus glasunovi Kataev, 1987
    Harpalus gravis LeConte, 1858
    Harpalus gregoryi Alluaud, 1917
    Harpalus hartmanni Kataev, 2002
    Harpalus herbivagus Say, 1823
    Harpalus heyrovskyi Jedlicka, 1928
    Harpalus hiekei Kataev & Wrase, 2010
    Harpalus hirtipes (Panzer, 1796)
    Harpalus honestus (Duftschmid, 1812)
    Harpalus hospes Sturm, 1818
    Harpalus hybridus Boheman, 1848
    Harpalus impressus Roth, 1851
    Harpalus inconcinnus Chaudoir, 1876
    Harpalus indianus Csiki, 1932
    Harpalus indicola Bates, 1878
    Harpalus indigens Casey, 1924
    Harpalus inexspectatus Kataev, 1989
    Harpalus ingenuus Tschitscherine, 1898
    Harpalus innocuus LeConte, 1863
    Harpalus italus Schaum, 1860
    Harpalus jeanneli Basilewsky, 1946
    Harpalus jordanus Jedlicka, 1964
    Harpalus kabakianus Kataev, 1988
    Harpalus kabakovi Kataev, 1987
    Harpalus kadleci Kataev & Wrase, 1995
    Harpalus kadyrbekovi Kataev, 1988
    Harpalus kagyzmanicus Kataev, 1984
    Harpalus kandaharensis Jedlicka, 1956
    Harpalus karakorum Jedlicka, 1958
    Harpalus karamani Apfelbeck, 1902
    Harpalus kazanensis Jedlicka, 1958
    Harpalus kaznakovi Kataev & Wrase, 1997
    Harpalus kibonoti Alluaud, 1927
    Harpalus kirgisicus (Motschulsky, 1844)
    Harpalus kiritshenkoi Kataev, 1990
    Harpalus klapperichi Jedlicka, 1956
    Harpalus kmecoi Facchini, 2003
    Harpalus kozlovi Kataev, 1993
    Harpalus kryzhanovskii Kataev, 1988
    Harpalus kunarensis Kataev, 1993
    Harpalus laetus Reiche, 1843
    Harpalus laevipes Zetterstedt, 1828
    Harpalus lama Kataev & Wrase, 1997
    Harpalus lateralis Dejean, 1829
    Harpalus laticeps LeConte, 1850
    Harpalus latus (Linnaeus, 1758)
    Harpalus lederi Tschitscherine, 1899
    Harpalus lethierryi Reiche, 1860
    Harpalus lewisii LeConte, 1865
    Harpalus longipalmatus Mordkovitsh, 1969
    Harpalus lopezi J.Serrano & Lencina, 2009
    Harpalus lugubris Boheman, 1848
    Harpalus lumbaris Mannerheim, 1825
    Harpalus luteicornis (Duftschmid, 1812)
    Harpalus lutshniki Schauberger, 1932
    Harpalus macronotus Tschitscherine, 1893
    Harpalus mairei Peyerimhoff, 1928
    Harpalus major (Motschulsky, 1850)
    Harpalus makhekensis Basilewsky, 1958
    Harpalus manas Kataev, 1990
    Harpalus marginellus Gyllenhal, 1827
    Harpalus martini Van Dyke, 1926
    Harpalus masoreoides Bates, 1878
    Harpalus massarti Burgeon, 1936
    Harpalus mauritanicus Gaubil, 1844
    Harpalus medvedevi Kataev, 2006
    Harpalus megacephalus LeConte, 1847
    Harpalus melaneus Bates, 1878
    Harpalus metallinus Ménétriés, 1836
    Harpalus metarsius Andrewes, 1930
    Harpalus meteorus Basilewsky, 1946
    Harpalus michaili Kataev, 1990
    Harpalus michailovi Kataev, 1987
    Harpalus microthorax (Motschulsky, 1849)
    Harpalus miles Péringuey, 1896
    Harpalus minutissimus Facchini, 2015
    Harpalus minutulus Kataev & Liang, 2004
    Harpalus mitridati Pliginskiy, 1915
    Harpalus mlynari Kataev, 1990
    Harpalus modestus Dejean, 1829
    Harpalus morvani Kataev, 2002
    Harpalus murzini Kataev, 2008
    Harpalus nanniscus Péringuey, 1896
    Harpalus natalensis Boheman, 1848
    Harpalus natalicus Péringuey, 1896
    Harpalus neglectus Audinet-Serville, 1821
    Harpalus nevadensis K. & J.Daniel, 1898
    Harpalus nigrans A.Morawitz, 1862
    Harpalus nigritarsis C.R.Sahlberg, 1827
    Harpalus numidicus Bedel, 1893
    Harpalus nyassicus Basilewsky, 1946
    Harpalus oblitus Dejean, 1829
    Harpalus obnixus Casey, 1924
    Harpalus ochropus Kirby, 1837
    Harpalus oodioides Dejean, 1829
    Harpalus opacipennis (Haldeman, 1843)
    Harpalus optabilis Dejean, 1829
    Harpalus ovtshinnikovi Kataev, 1990
    Harpalus pallidipennis A.Morawitz, 1862
    Harpalus pallipes Chaudoir, 1837
    Harpalus parallelocollis Facchini, 2003
    Harpalus parasinuatus Kataev & Liang, 2007
    Harpalus parvulus Dejean, 1829
    Harpalus petri Tschitscherine, 1902
    Harpalus picipennis (Duftschmid, 1812)
    Harpalus plancyi Tschitscherine, 1897
    Harpalus plenalis Casey, 1914
    Harpalus politus Dejean, 1829
    Harpalus polyglyptus Schaum, 1862
    Harpalus potanini Tschitscherine, 1906
    Harpalus praecurrens Schauberger, 1934
    Harpalus procognatus Lorenz, 1998
    Harpalus progrediens Schauberger, 1922
    Harpalus prosperus Basilewsky, 1972
    Harpalus providens Casey, 1914
    Harpalus pseudoserripes Reitter, 1900
    Harpalus pterostichus (Reitter, 1900)
    Harpalus puetzi Kataev & Wrase, 1997
    Harpalus pulchrinulus Reitter, 1900
    Harpalus pulvinatus Ménétriés, 1848
    Harpalus pumilus Sturm, 1818
    Harpalus punctatostriatus Dejean, 1829
    Harpalus punctipennis Mulsant, 1852
    Harpalus pusillus (Motschulsky, 1850)
    Harpalus pygmaeus Dejean, 1829
    Harpalus quadratus Chaudoir, 1846
    Harpalus raphaili Kataev, 1997
    Harpalus reflexus Putzeys in Schneider & Leder, 1878
    Harpalus reitteri Wrase & Kataev, 2011
    Harpalus remboides Solsky, 1874
    Harpalus reversus Casey, 1924
    Harpalus rhemboides Solsky, 1874
    Harpalus rivalsi Jeannel, 1948
    Harpalus rotundus Facchini, 2003
    Harpalus rougemonti Clarke, 1973
    Harpalus rubripes (Duftschmid, 1812)
    Harpalus rufipalpis Sturm, 1818
    Harpalus rufiscapus Gebler, 1833
    Harpalus rufitarsoides Schauberger, 1934
    Harpalus rufomarginatus (Boheman, 1848)
    Harpalus rumelicus Apfelbeck, 1904
    Harpalus salinulus Reitter, 1900
    Harpalus salinus Dejean, 1829
    Harpalus sanctaehelenae Basilewsky, 1972
    Harpalus saxicola Dejean, 1829
    Harpalus semenowi Tschitscherine, 1901
    Harpalus semipunctatus Dejean, 1829
    Harpalus serripes (Quensel, 1806)
    Harpalus servus (Duftschmid, 1812)
    Harpalus setiporus (Reitter, 1894)
    Harpalus siculus Dejean, 1829
    Harpalus silipes Dejean, 1831
    Harpalus sinuaticollis Facchini, 2015
    Harpalus sinuatus Tschitscherine, 1893
    Harpalus smaragdinus (Duftschmid, 1812)
    Harpalus smyrnensis Heyden, 1888
    Harpalus solitaris Dejean, 1829
    Harpalus somereni Basilewsky, 1946
    Harpalus somnulentus Dejean, 1829
    Harpalus spadiceus Dejean, 1829
    Harpalus spretus Péringuey, 1896
    Harpalus spurius Péringuey, 1896
    Harpalus stevenii Dejean, 1829
    Harpalus stevensi Kataev, 2011
    Harpalus stoetznerianus Schauberger, 1932
    Harpalus strenuus Tschitscherine, 1898
    Harpalus subaeneus Boheman, 1848
    Harpalus subcylindricus Dejean, 1829
    Harpalus subtruncatus Chaudoir, 1846
    Harpalus sulphuripes Germar, 1823
    Harpalus sundaicus Schauberger, 1933
    Harpalus sushenicus Kataev, 1990
    Harpalus suturangulus Reitter, 1887
    Harpalus szalliesi Kataev & Wrase, 1995
    Harpalus taciturnus Dejean, 1829
    Harpalus tangutorum Kataev, 1993
    Harpalus tardus (Panzer, 1796)
    Harpalus tarsalis Mannerheim, 1825
    Harpalus terrestris (Motschulsky, 1844)
    Harpalus tibeticus Andrewes, 1930
    Harpalus tichonis Jakobson, 1907
    Harpalus tiridates Reitter, 1900
    Harpalus tithonus Reitter, 1900
    Harpalus tjanschanicus Semenov, 1889
    Harpalus torosensis Jedlicka, 1961
    Harpalus torridoides Reitter, 1900
    Harpalus trichophorus Tschitscherine, 1897
    Harpalus triseriatus A.Fleischer, 1897
    Harpalus turcicus Jedlicka, 1958
    Harpalus turmalinus Erichson, 1847
    Harpalus udege Lafer, 1989
    Harpalus uniformis (Motschulsky, 1844)
    Harpalus ussuricus Mlynar, 1979
    Harpalus vanemdeni Schauberger, 1932
    Harpalus venator Boheman, 1848
    Harpalus ventralis LeConte, 1847
    Harpalus vereschaginae Kataev, 1988
    Harpalus vernicosus Kataev & Liang, 2007
    Harpalus viridanus (Motschulsky, 1844)
    Harpalus viridicupreus Reiche, 1843
    Harpalus vittatus Gebler, 1833
    Harpalus wadiensis Jedlicka, 1964
    Harpalus wohlberedti Emden, 1932
    Harpalus xanthopus Gemminger & Harold, 1868
    Harpalus zabroides Dejean, 1829
    Harpalus zhdankoi Kataev, 1990

### Subgenus Loboharpalus Schauberger, 1932

    Harpalus platynotus Bates, 1873
    Harpalus rubefactus Bates, 1873

### Subgenus Opadius Casey, 1914

    Harpalus apache Kataev, 2010
    Harpalus cordatus (LeConte, 1853)
    Harpalus cordifer Notman, 1919

### Subgenus Plectralidus Casey, 1914

    Harpalus erraticus Say, 1823
    Harpalus retractus LeConte, 1863

### Subgenus Pseudoophonus Motschulsky, 1844

    Harpalus actiosus Casey, 1914
    Harpalus aenigma (Tschitscherine, 1897)
    Harpalus alienus Bates, 1878
    Harpalus aogashimensis (Habu, 1957)
    Harpalus azumai Habu, 1968
    Harpalus babai Habu, 1973
    Harpalus calceatus (Duftschmid, 1812)
    Harpalus caliginosus (Fabricius, 1775)
    Harpalus capito A.Morawitz, 1862
    Harpalus compar LeConte, 1847
    Harpalus coreanus (Tschitscherine, 1895)
    Harpalus davidi (Tschitscherine, 1897)
    Harpalus disimuciulus Huang; Lei; Yan & Hu, 1996
    Harpalus eous Tschitscherine, 1901
    Harpalus erythropus Dejean, 1829
    Harpalus faunus Say, 1823
    Harpalus fokienensis Schauberger, 1930
    Harpalus griseus (Panzer, 1796)
    Harpalus hatchi Ball & Anderson, 1962
    Harpalus hauserianus Schauberger, 1929
    Harpalus indicus Bates, 1891
    Harpalus jureceki (Jedlicka, 1928)
    Harpalus katiae F.Battoni, 1985
    Harpalus liobasis Chaudoir, 1868
    Harpalus meghalayensis Kataev, 2001
    Harpalus meridianus Andrewes, 1923
    Harpalus paratus Casey, 1924
    Harpalus pastor Motschulsky, 1844
    Harpalus pensylvanicus (DeGeer, 1774)
    Harpalus poncei Will, 2002
    Harpalus protractus Casey, 1914
    Harpalus pseudohauserianus Kataev, 2001
    Harpalus pseudophonoides Schauberger, 1930
    Harpalus roninus Bates, 1873
    Harpalus rufipes (DeGeer, 1774)
    Harpalus sericatus Tschitscherine, 1906
    Harpalus simplicidens Schauberger, 1929
    Harpalus singularis Tschitscherine, 1906
    Harpalus sinicus Hope, 1845
    Harpalus suensoni Kataev, 1997
    Harpalus texanus Casey, 1914
    Harpalus tridens A.Morawitz, 1862
    Harpalus ussuriensis Chaudoir, 1863
    Harpalus vagans LeConte, 1865
    Harpalus yinchuanensis Huang, 1993

### Subgenus Semiophonus Schauberger, 1933

    Harpalus signaticornis (Duftschmid, 1812)

### Subgenus Zangoharpalus Huang, 1998

    Harpalus mariae Kataev, 1997
    Harpalus microdemas Schauberger, 1932
    Harpalus praticola Bates, 1891
    Harpalus pseudotinctulus Schauberger, 1932
    Harpalus tinctulus Bates, 1873

### Extinct, not assigned to subgenus

    †Harpalus abolitus C. & L.Heyden, 1866
    †Harpalus anactus Giebel, 1856
    †Harpalus bruckmanni Heer, 1862
    †Harpalus burmeisteri Giebel, 1856
    †Harpalus conditus Scudder, 1900
    †Harpalus constrictus Heer, 1862
    †Harpalus deletus Oustalet, 1864
    †Harpalus dilatatus Theobald, 1937
    †Harpalus ewaldi Giebel, 1856
    †Harpalus excavatus Forster, 1891
    †Harpalus heeri Giebel, 1856
    †Harpalus knorri Giebel, 1856
    †Harpalus lheritieri Piton, 1940
    †Harpalus liasinus Giebel, 1856
    †Harpalus maceratus Wickham, 1911
    †Harpalus nero Oustalet, 1864
    †Harpalus nitens Birket-Smith, 1977
    †Harpalus nuperus Scudder, 1900
    †Harpalus obtusus Forster, 1891
    †Harpalus oustaleti Meunier, 1915
    †Harpalus pleistocenicus Lomnicki, 1894
    †Harpalus redivivus Wickham, 1917
    †Harpalus schlotheimi Giebel, 1856
    †Harpalus sinis Heer, 1847
    †Harpalus stierlini Heer, 1862
    †Harpalus stygius Heer, 1862
    †Harpalus tabidus Heer, 1847
    †Harpalus tardigradus Heer, 1862
    †Harpalus ulomaeformis Wickham, 1917
    †Harpalus veterum Cockerell, 1920
    †Harpalus whitfieldi Scudder, 1900