# Cotinga

The cotingas are a large family, Cotingidae, of suboscine passerine birds found in Central America and tropical South America. Cotingas are birds of forests or forest edges, that are primary frugivorous. They all have broad bills with hooked tips, rounded wings, and strong legs. They range in size from 12–13 cm (4.7–5.1 in) of the fiery-throated fruiteater (Pipreola chlorolepidota) up to 48–51 cm (19–20 in) of the Amazonian umbrellabird (Cephalopterus ornatus).

## Description

Cotingas vary widely in social structure. There is a roughly 50/50 divide in the family between species with biparental care, and those in which the males play no part in raising the young. The purple-throated fruitcrow lives in mixed-sex groups in which one female lays an egg and the others help provide insects to the chick.
In cotinga species where only the females care for the eggs and young, the males have striking courtship displays, often grouped together in leks. Such sexual selection results in the males of these species, including the Guianan cock-of-the-rock, being brightly coloured, or decorated with plumes or wattles, like the umbrellabirds, with their umbrella-like crest and long throat wattles. Other lekking cotingids like the bellbirds and screaming piha, have distinctive and far-carrying calls. In such canopy-dwelling genera as Carpodectes, Cotinga, and Xipholena, males gather high in a single tree or in adjacent trees, but male cocks-of-the-rock, as befits their more terrestrial lives, give their elaborate displays in leks on the ground.
The females of both lekking and biparental species are duller than the males.

## Breeding

Nests range from tiny to very large. Many species lay a single egg in a nest so flimsy that the egg can be seen from underneath. This may make the nests hard for predators to find. Fruiteaters build more solid cup nests, and the cocks-of-the-rock attach their mud nests to cliffs. The nests may be open cups or little platforms with loosely woven plant material, usually placed in a tree. The clutches comprise one to four eggs. Incubation typically takes 15–28 days. Fledging usually occurs at 28–33 days.

## Habitat

Deserts, open woodlands, coastal mangroves, and humid tropical forests comprise their habitats. Cotingas face very serious threats from the loss of their habitats.

## Taxonomy and systematics

The family Cotingidae was introduced by French naturalist Charles Lucien Bonaparte in 1849. According to the International Ornithological Committee, as of July 2021, the family contains 66 species divided into 24 genera.
A 2014 molecular phylogenetic study of the cotingas by Jacob Berv and Richard Prum found that the genera formed five monophyletic clades and they proposed that the family could be divided into five subfamilies. The following cladogram is based on a molecular phylogenetic study of the suboscines by Michael Harvey and collaborators published in 2020.
The genus Tijuca was found to be embedded in Lipaugus, a position that was confirmed by a more detailed 2020 study.
| Genus | Living species |
| Ampelioides Verreaux, 1867 | Scaled fruiteater, Ampelioides tschudii |
| Pipreola Swainson, 1838 | Fiery-throated fruiteater, Pipreola chlorolepidota Scarlet-breasted fruiteater, Pipreola frontalis Handsome fruiteater, Pipreola formosa Red-banded fruiteater, Pipreola whitelyi Black-chested fruiteater, Pipreola lubomirskii Orange-breasted fruiteater, Pipreola jucunda Masked fruiteater, Pipreola pulchra Golden-breasted fruiteater, Pipreola aureopectus Barred fruiteater, Pipreola arcuata Band-tailed fruiteater, Pipreola intermedia Green-and-black fruiteater, Pipreola riefferii |
| Snowornis Prum, 2001 | Grey-tailed piha, Snowornis subalaris Olivaceous piha, Snowornis cryptolophus |
| Carpornis G.R. Gray, 1846 | Hooded berryeater, Carpornis cucullata Black-headed berryeater, Carpornis melanocephala |
| Rupicola Brisson, 1760 | Andean cock-of-the-rock, Rupicola peruvianus Guianan cock-of-the-rock, Rupicola rupicola |
| Phoenicircus Swainson, 1832 | Guianan red cotinga, Phoenicircus carnifex Black-necked red cotinga, Phoenicircus nigricollis |
| Zaratornis Koepcke, 1954 | White-cheeked cotinga, Zaratornis stresemanni |
| Phytotoma Molina, 1782 | Rufous-tailed plantcutter or Chilean plantcutter, Phytotoma rara Peruvian plantcutter, Phytotoma raimondii White-tipped plantcutter or reddish plantcutter, Phytotoma rutila |
| Phibalura Vieillot, 1816 | Swallow-tailed cotinga, Phibalura flavirostris |
| Doliornis Taczanowski, 1874 | Bay-vented cotinga, Doliornis sclateri Chestnut-bellied cotinga, Doliornis remseni |
| Ampelion Tschudi, 1845 | Red-crested cotinga, Ampelion rubrocristatus Chestnut-crested cotinga, Ampelion rufaxilla |
| Haematoderus Bonaparte, 1854 | Crimson fruitcrow, Haematoderus militaris |
| Querula Vieillot, 1816 | Purple-throated fruitcrow, Querula purpurata |
| Pyroderus G.R. Gray, 1840 | Red-ruffed fruitcrow, Pyroderus scutatus |
| Cephalopterus E. Geoffroy Saint-Hilaire, 1809 | Bare-necked umbrellabird, Cephalopterus glabricollis Long-wattled umbrellabird, Cephalopterus penduliger Amazonian umbrellabird, Cephalopterus ornatus |
| Perissocephalus Oberholser, 1899 | Capuchinbird, Perissocephalus tricolor |
| Lipaugus F. Boie, 1828 | Rufous piha, Lipaugus unirufus Cinnamon-vented piha, Lipaugus lanioides Rose-collared piha, Lipaugus streptophorus Screaming piha, Lipaugus vociferans Dusky piha, Lipaugus fuscocinereus Scimitar-winged piha, Lipaugus uropygialis Chestnut-capped piha, Lipaugus weberi Black-and-gold cotinga, Lipaugus ater Grey-winged cotinga, Lipaugus conditus |
| Procnias Illiger, 1811 | White bellbird, Procnias albus Three-wattled bellbird, Procnias tricarunculatus Bare-throated bellbird, Procnias nudicollis Bearded bellbird, Procnias averano |
| Cotinga Brisson, 1760 | Plum-throated cotinga, Cotinga maynana Spangled cotinga, Cotinga cayana Lovely cotinga, Cotinga amabilis Blue cotinga, Cotinga nattererii Turquoise cotinga, Cotinga ridgwayi Banded cotinga, Cotinga maculata Purple-breasted cotinga, Cotinga cotinga |
| Porphyrolaema Bonaparte, 1854 | Purple-throated cotinga, Porphyrolaema porphyrolaema |
| Conioptilon Lowery & O'Neill, 1966 | Black-faced cotinga, Conioptilon mcilhennyi |
| Gymnoderus E. Geoffroy Saint-Hilaire, 1809 | Bare-necked fruitcrow, Gymnoderus foetidus |
| Xipholena Gloger, 1841 | Pompadour cotinga, Xipholena punicea White-tailed cotinga, Xipholena lamellipennis White-winged cotinga, Xipholena atropurpurea |
| Carpodectes Salvin, 1865 | Black-tipped cotinga, Carpodectes hopkei Yellow-billed cotinga, Carpodectes antoniae Snowy cotinga, Carpodectes nitidus |
A number of species previously placed in this family are now placed in the family Tityridae (genera Laniisoma, Laniocera and Iodopleura)