# Taxonomy of Lucanidae

The Lucanidae are a family of beetles that include the stag beetles. The classification presented here follows Smith (2006), with the exception of the tribal classification within the Lucaninae.

## Subfamily Aesalinae

Subfamily Aesalinae
    Tribe Aesalini
       Genus Aesalus (Fabricius, 1801)
         * Aesalus asiaticus Lewis, 1883
         * Aesalus himalayicus Kurosawa, 1985
         * Aesalus imanishii Inahara & Ratti, 1991
         * Aesalus meridionalis Bartolozzi, 1989
         * Aesalus neotropicalis Bates, 1886
         * Aesalus saburoi
         * Aesalus sawaii Fujita & Ichikawa, 1985
         * Aesalus scarabaeoides Panzer, 1793
         * Aesalus sichuanensis
         * Aesalus smithi Bates, 1889
         * Aesalus trogoides Albers, 1883
         * Aesalus ulanowskii Ganglbauer, 1886
       Genus Cretaesalus (fossil)
       Genus Echinoaesalus Zelenka, 1993
         * Echinoaesalus barriesi (Zelenka, 1993)
         * Echinoaesalus hidakai (Araya, 1993)
         * Echinoaesalus jaechi (Zelenka,1993)
         * Echinoaesalus matsuii (Araya, 1993)
         * Echinoaesalus sabahensis (Zelenka, 1994)
         * Echinoaesalus schuhi (Zelenka, 1994)
         * Echinoaesalus timidus (Krikken, 1975)
         * Echinoaesalus yongi (Araya, 1993)
       Genus Lucanobium Howden & Lawrence, 1974
         * Lucanobium squamosum Howden & Lawrence, 1974
    Tribe Ceratognathini
       Genus Ceratognathus Westwood, 1838
         * Ceratognathus abdominalis Parry, 1870
         * Ceratognathus bitumulatus Carter, 1925
         * Ceratognathus flabellatus Boileau, 1905
         * Ceratognathus frenchi Blackburn, 1897
         * Ceratognathus froggatti Blackburn, 1894
         * Ceratognathus gilesi Blackburn, 1895
         * Ceratognathus macrognathus Boileau, 1905
         * Ceratognathus mentiferus Westwood, 1863
         * Ceratognathus minutus Lea, 1929
         * Ceratognathus niger Westwood, 1838
         * Ceratognathus ocularis Carter, 1925
         * Ceratognathus rufipennis Westwood, 1872
         * Ceratognathus tasmanus Benesh, 1943
         * Ceratognathus westwoodi Thomson, 1862
       Genus Hilophyllus Paulsen & Mondaca, 2006
         * Hilophyllus argentinensis (Martinez, 1981)
         * Hilophyllus martinezi Paulsen & Mondaca, 2006
         * Hilophyllus penai (Martinez, 1976)
       Genus Mitophyllus Parry, 1843
         * Mitophyllus alboguttatus (Bates, 1867)
         * Mitophyllus dispar (Sharp, 1882)
         * Mitophyllus foveolatus (Broun, 1880)
         * Mitophyllus gibbosus (Broun, 1884)
         * Mitophyllus insignis Broun, 1921
         * Mitophyllus irroratus Parry, 1842
         * Mitophyllus macrocerus (Broun, 1886)
         * Mitophyllus parrianus Westwood, 1863
         * Mitophyllus reflexus Broun, 1908
    Tribe Nicagini
       Genus Nicagus LeConte, 1861
         * Nicagus obscurus (LeConte, 1848)
         * Nicagus occultus Paulsen & Smith, 2005
         * Nicagus japonicus Nagel, 1928
       Genus Holloceratognathus Nikolajev, 1998
         * Holloceratognathus cylindricus (Broun, 1895)
         * Holloceratognathus helotoides (Thomson, 1862)
         * Holloceratognathus passaliformis (Holloway, 1962)

## Subfamily Lampriminae

Subfamily Lampriminae
       Genus Dendroblax White, 1846
         * Dendroblax earlianus White, 1846
       Genus Homolamprima
         * Homolamprima crenulata (MacLeay, 1885)
       Genus Lamprima
         * Lamprima adolphinae (Gestro, 1875)
         * Lamprima aenea Fabricius, 1792
         * Lamprima aurata Latreille, 1817
         * Lamprima imberbis Barber, 1926
         * Lamprima insularis Macleay, 1885
       Genus Phalacrognathus
         * Phalacrognathus muelleri MacLeay, 1885
       Genus Streptocerus Dejean, 1833
         * Streptocerus speciosus Fairmaire, 1850

## Subfamily Lucaninae

Subfamily Lucaninae Latreille, 1804
    Tribe Aegini
       Genus Aegus MacLeay, 1819
    Tribe Allotopini
       Genus Allotopus
         * Allotopus moellenkampi (Fruhstorfer, 1894)
         * Allotopus rosenbergi (Vollenhoven, 1872)
       Genus Mesotopus
    Tribe Chiasognathini
       Genus Casignetus
       Genus Chiasognathus Stephens, 1831
         * Chiasognathus beneshi Lacroix, 1978
         * Chiasognathus grantii Stephens, 1831
         * Chiasognathus jousselini Reiche, 1850
         * Chiasognathus latreillei Solier, 1851
         * Chiasognathus mniszechii Kiesche 1919
       Genus Protognathinus
       Genus Sphaenognathus
    Tribe Cladognathini
       Genus Aphanognathus
         * Aphanognathus laterotarsoides (Houlbert, 1915)
       Genus Capreolucanus
         * Capreolucanus sicardi Didier, 1928
       Genus Cladophyllus
       Genus Eligmodontus
       Genus Gonometopus
       Genus Macrodorcas Motschulsky, 1862
         * Macrodorcas adachii
         * Macrodorcas amamiana
         * Macrodorcas barbata
         * Macrodorcas bisignata
         * Macrodorcas bomansi
         * Macrodorcas davidi
         * Macrodorcas dierli
         * Macrodorcas elsiledis
         * Macrodorcas fulvonotata
         * Macrodorcas humilis
         * Macrodorcas intricata
         * Macrodorcas kobayashii
         * Macrodorcas koyamai
         * Macrodorcas melliana
         * Macrodorcas mochizukii
         * Macrodorcas moellenkampi
         * Macrodorcas montivaga
         * Macrodorcas pieli
         * Macrodorcas pseudaxis
         * Macrodorcas recta Motschulsky, 1858
         * Macrodorcas ruficrus
         * Macrodorcas songiana
         * Macrodorcas striatipennis Motschulsky, 1862
         * Macrodorcas ursulae
         * Macrodorcas vernicata
         * Macrodorcas yaeyamaensis
       Genus Palaeognathus
       Genus Prismognathus Motschulsky, 1860
         * Prismognathus alessandrae
         * Prismognathus angularis Waterhouse, 1874
         * Prismognathus branczicki
         * Prismognathus cheni
         * Prismognathus dauricus Motschulsky, 1860
         * Prismognathus davidis
         * Prismognathus delislei
         * Prismognathus formosanus
         * Prismognathus klapperichi
         * Prismognathus lucidus
         * Prismognathus morimotoi
         * Prismognathus nigerrimus
         * Prismognathus nigricolor
         * Prismognathus piluensis
         * Prismognathus platycephalus
         * Prismognathus ruficephalus
         * Prismognathus sinensis
         * Prismognathus subnitens
         * Prismognathus tokui
       Genus Prosopocoilus Westwood, 1845
       Genus Pseudorhaetus
       Genus Rhaetulus
       Genus Rhaetus
         * Rhaetus westwoodi (Parry, 1862)
       Genus Tetrarthrius
       Genus Weinreichius
         * Weinreichius perroti Lacroix, 1978
    Tribe Colophonini
       Genus Colophon
    Tribe Cyclommatini
       Genus Cyclommatus Parry, 1863
    Tribe Dendeziini
       Genus Dendezia
       Genus Oonotus Parry, 1864
       Genus Xiphodontus
    Tribe Dorcini
       Genus Bartolozziolucanus
       Genus Cantharolethrus
         * Cantharolethrus luxeri
       Genus Cyclommatus
         * Cyclommatus scutellaris
       Genus Dorcasoides
       Genus Dorcus MacLeay, 1819
         * Dorcus akahorii
         * Dorcus alcides
         * Dorcus alexisi
         * Dorcus amamianus
         * Dorcus antaeus
         * Dorcus anteus
         * Dorcus arfakianus
         * Dorcus binodulosus
         * Dorcus brevis
         * Dorcus bucephalus
         * Dorcus carinulatus
         * Dorcus consentaneus
         * Dorcus curvidens
         * Dorcus cylindricus
         * Dorcus davidis
         * Dorcus emikoae
         * Dorcus eurycephalus
         * Dorcus formosanus
         * Dorcus gracilicornis
         * Dorcus garndis
         * Dorcus hopei
         * Dorcus hyperion
         * Dorcus japonicus
         * Dorcus kyanrauensis
         * Dorcus metacostatus
         * Dorcus miwai
         * Dorcus musimon
         * Dorcus parallelus
         * Dorcus parallelipipedus - lesser stag beetle. Linnaeus, 1758
         * Dorcus parryi
         * Dorcus parvulus
         * Dorcus peyronis
         * Dorcus prochazkai
         * Dorcus ratiocinativus
         * Dorcus rectus
         * Dorcus reichei
         * Dorcus rudis
         * Dorcus sawaii
         * Dorcus schenklingi
         * Dorcus sewertzowi
         * Dorcus suturalis
         * Dorcus taiwanicus
         * Dorcus tanakai
         * Dorcus taurus
         * Dorcus tenuecostatus
         * Dorcus titanus
         * Dorcus tormosanus
         * Dorcus ursulus
         * Dorcus vavrai
         * Dorcus velutinus
         * Dorcus vicinus
         * Dorcus yamadai
       Genus Homoderus
       Genus Leptinopterus
       Genus Prosopocoilus
         * Prosopocoilus astacoides
         * Prosopocoilus biplagiatus
         * Prosopocoilus bison
         * Prosopocoilus giraffa
         * Prosopocoilus inclinatus
         * Prosopocoilus savagei
       Genus Rhaetulus
         * Rhaetulus crenatus
       Genus Serrognathus
       Genus Pycnosiphorus
       Genus Sclerostomus
    Tribe Figulini
       Genus Cardanus
       Genus Dinonigidius
       Genus Figulus
       Genus Ganelius
       Genus Nigidionus
       Genus Nigidius
       Genus Novonigidius
       Genus Penichrolucanus
    Tribe Lissapterini
       Genus Bomansius
       Genus Dorculus
       Genus Geodorcus
       Genus Hoplogonus
       Genus Lissapterus
       Genus Lissotes
       Genus Paralissotes
       Genus Pseudodorcus
    Tribe Lucanini
       Genus Hexarthrius Hope, 1842
         * Hexarthrius aduncusJordan & Rothschild, 1894
         * Hexarthrius andreasi Schenk, 2003
         * Hexarthrius bowringii Parry, 1862
         * Hexarthrius buquettii (Hope, 1843)
         * Hexarthrius davisoniWaterhouse, 1888
         * Hexarthrius forsteri (Hope, 1840)
         * Hexarthrius howdeni De Lisle, 1972
         * Hexarthrius kirchneri Schenk, 2003
         * Hexarthrius mandibularis Deyrolle, 1881
         * Hexarthrius melchioritis Séguy, 1954
         * Hexarthrius mniszechi(Thomson, 1857)
         * Hexarthrius nigritus Lacroix, 1990
         * Hexarthrius parryi Hope, 1842
         * Hexarthrius rhinoceros(Olivier, 1789)
         * Hexarthrius vitalisi Didier, 1925
       Genus Lucanus
         * Lucanus capreolus
         * Lucanus cervus
         * Lucanus elaphus
         * Lucanus formosanus
         * Lucanus maculifemoratus
         * Lucanus mazama
         * Lucanus placidus
         * Lucanus swinhoei
         * Lucanus tetraodon
       Genus Pseudolucanus
         * Pseudolucanus barbarossa (Fabricius, 1801)
    Tribe Odontolabini
       Genus Calcodes
       Genus Neolucanus
         * Neolucanus castanopterus
       Genus Odontolabis
         * Odontolabis castelnaudi
         * Odontolabis cuvera
         * Odontolabis femoralis
         * Odontolabis siva
    Tribe Platycerini
       Genus Platyceroides Benesh, 1946
       Genus Platyceropsis Benesh, 1946
         * Platyceropsis keeni (Casey, 1895)
       Genus Platycerus Geoffroy, 1762
    Tribe Ryssonotini
       Genus Cacostomus
         * Cacostomus floralis (Lea, 1914)
         * Cacostomus squamosus Newman, 1840
         * Cacostomus subvittata (Moore, 1994)
       Genus Ryssonotus

## Subfamily Syndesinae

Subfamily Syndesinae
    Tribe Ceruchini
       Genus Ceruchus MacLeay, 1819
         * Ceruchus piceus (Weber, 1801)
         * Ceruchus punctatus LeConte, 1869
         * Ceruchus striatus LeConte, 1859
         * Ceruchus yangi Huang, Imura & Chen, 2011
    Tribe Sinodendronini
       Genus Sinodendron Hellwig in Schneider, 1792
         * Sinodendron cylindricum (Linnaeus, 1758)
         * Sinodendron rugosum Mannerheim, 1843
    Tribe Syndesini
       Genus Psilodon Perty, 1830
         * Psilodon aequinoctiale (Buquet, 1840)
         * Psilodon gilberti Boucher, 1993
         * Psilodon schuberti Perty, 1830
         * Psilodon seguyi (Didier, 1929)
         * Psilodon xerophilicum Martinez & Reyes-Castillo, 1985
       Genus Syndesus MacLeay, 1819
         * †Syndesus ambericus Woodruff, 2009 (Miocene?; Dominican amber)
         * Syndesus cancellatus Montrouzier, 1860
         * Syndesus cornutus (Fabricius, 1801)
         * Syndesus macleayi Boileau, 1905