# 1994 in paleontology

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1994.

## Plants
### Conifers

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Diploporus | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | Fossil Yew Genus, type species D. torreyoides |
| Taxus masonii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | Fossil Yew species |
| Torreya clarnensis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | Fossil nutmeg yew species |

### Angiosperms

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Actinidia oregonensis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Kiwi species | |
| Alangium eydei | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A dogwood species | |
| Alangium rotundicarpum | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A dogwood species | |
| Ampelocissus auriforma | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A vitaceous species | |
| Ampelocissus scottii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A vitaceous species | |
| Ampelopsis rooseae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A vitaceous species | |
| Anamirta leiocarpa | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species | |
| Ankistrosperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species A. spitzerae | |
| Anonaspermum bonesii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A custard apple relative | |
| Anonaspermum rotundum | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A custard apple relative | |
| Anonymocarpa | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species A. ovoidea | |
| Aphananthe maii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A cannabaceous species | |
| Ascosphaera | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species A. eocenis | |
| Atriaecarpum clarnense | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species | |
| Axinosperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species A. agnostum | |
| Bonesia | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species B. spatulata | |
| Bumelia? globosa | Sp nov | jr synonym | (Manchester) Manchester | Middle Eocene | Clarno Formation | USA Oregon | A lardizabalaceous species; Moved to Sargentodoxa globosa in 1999 | |
| Bursericarpum oregonense | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A torchwood family species | |
| Calycocarpum crassicrustae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species | |
| Castanopsis crepetii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A chinkapin species | |
| Celtis burnhamae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A cannabaceous species | |
| Cleyera grotei | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A pentaphylacaceous species | |
| Comicilabium | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | An icacinaceous relative. type species C. atkinsii. | |
| Comminicarpa | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species C. friisae | |
| Cornus clarnensis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A dogwood species | |
| Coryloides | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A hazelnut relative, Type species C. hancockii | |
| Cuneisemen | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species C. truncatum | |
| Curvitinospora | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed relative. Type species C. formanii | |
| Davisicarpum limacioides | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species. | |
| Dentisemen | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species D. parvum | |
| Deviacer | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A milkwort genus. Type species D. wolfei | |
| Diploclisia auriformis | Comb nov | jr synonym | (Hollick) Manchester | Paleocene/Eocene | "King Salmon Lake flora" | USA Alaska | A moonseed species. Moved from Carpolithes auriformis (1936) Moved to Stephania auriformis in 2018 | |
| Durocarpus | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species D. cordatus | |
| Emmenopterys dilcheri | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A rubiaceous species. | |
| Eohypserpa scottii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species. | |
| Eucommia constans | Sp nov | valid | Magallón-Puebl & Cevallos-Ferriz | Miocene - Pleistocene | Pié de Vaca Formation | Mexico | Youngest Eucommia sp. in North America. | |
| Ferrignocarpus | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species F. bivalvis | |
| Fimbrialata | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species F. wingii | |
| Fortunearites | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A witch-hazel relative, type species F. endressii | |
| Fragarites | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species F. ramificans | |
| Globulicarpium | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species G. levigatum | |
| Hexacarpellites | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species H. hallii | |
| Hydrangea knowltonii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Hydrangea species. | |
| Joejonesia | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species J. globosa | |
| lodes chandlerae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | An icacinaceous species. | |
| Iodicarpa | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | An icacinaceous relative. type species I. parvum, includes I. lenticularis | |
| Kardiasperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A hazelnut relative, type species K. parvum | |
| Laurocalyx wheelerae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A laural relative | |
| Laurocarpum hancockii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A laural relative | |
| Laurocarpum nutbedensis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A laural relative | |
| Laurocarpum raisinoides | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A laural relative | |
| Lignicarpus | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species L. crassimuri | |
| Ligniglobus | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species L. sinuosifibrae | |
| Lindera clarnensis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A spicebush | |
| Lunaticarpa | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species L. curvistriata | |
| Magnolia muldoonae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Magnolia species | |
| Magnolia paroblonga | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Magnolia species | |
| Magnolia tiffneyi | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Magnolia species | |
| Mastixicarpum occidentale | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Tupelo relative | |
| Meliosma beusekomii | sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A sabiaceous species. | |
| Meliosma bonesii | sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A sabiaceous species. | |
| Meliosma elongicarpa | sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A sabiaceous species. | |
| Meliosma leptocarpa | sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A sabiaceous species. | |
| Microphallus | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species M. perplexus | |
| Nephrosemen | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species N. reticulatus | |
| Nyssa scottii | sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Tupelo relative | |
| Nyssa spatulata | Comb nov | valid | (Scott) Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Tupelo, new comb for Palaeonyssa spatulata Scott, 1954 | |
| Omsicarpium | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species O. striatum | |
| Palaeoallophylus globosa | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A soapberry species. | |
| Palaeoallophylus gordonii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A soapberry species. | |
| Paleopanax | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A ginseng genus, type species P. oregonensis | |
| Paleoplatycarya? hickeyi | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Platycarya relative species. | |
| Parthenocissus clarnensis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A vitaceous species | |
| Pasternackia | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. pusilla | |
| Pentoperculum | Gen et comb nov | valid | Manchester | Middle Eocene | London Clay | UK England | A Sumac relative. A new genus for Dracontomelon minimus (Reid & Chandler, 1933) | |
| Pileosperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. minutum, also includes P. ovatum | |
| Pistachioides | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. striata | |
| Platanus hirticarpa | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A platanaceous Infructescence species. | |
| Polygrana nutbedense | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. dictyum | |
| Polygrana | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. nutbedense | |
| Prunus olsonii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A cherry/plum species. | |
| Prunus weinsteinii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A cherry/plum species. | |
| Pteronepelys | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. wehrii | Pteronepelys wehrii |
| Pulvinisperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. minutum | |
| Pyrenacantha occidentalis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | An icacinaceous species. | |
| Pyrisemen | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species P. attenuatum | |
| Quintacava | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species Q. velosida | |
| Rhus rooseae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Sumac species | |
| Sabia prefoetida | Comb nov | valid | (Becker) Manchester | Late Eocene | Ruby flora | USA Montana | A sabiaceous species. Moved from Symplocarpus prefoetidus Becker 1969 | |
| Sambucuspermites | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. rugulosus | |
| Saxifragispermum tetragonalis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A willow relative | |
| Scabraecarpium | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. clarnense | |
| Scalaritheca | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. biseriata | |
| Scaphicarpium | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. radiatum | |
| Schisandra oregonensis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A magnolia vine species. | |
| Sphaerosperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. riesii | |
| Sphenosperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. baccatum | |
| Stockeycarpa | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. globosa | |
| Striatisperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species S. coronapunctatum | |
| Symplocos nooteboomii | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A Symplocaceae species. | |
| Tanyoplatanus | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A platanaceous Infructescence genus. Type species T. cranei. | |
| Tapiscia occidentalis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A tapisciaceous species | |
| Tenuisperma | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. ellipticum | |
| Tiffneycarpa | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. scleroidea | |
| Tinomiscoidea occidentalis | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species | |
| Tinospora elongata | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species | |
| Thanikaimonia | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed genus. Type species T. geniculata. | |
| Tinospora hardmanae | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A moonseed species | |
| Trema nucilecta | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A cannabaceous species | |
| Trigonostela | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. oregonensis | |
| Tripartisemen | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. bonesii | |
| Triplascapha | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. collinsonae | |
| Triplexivalva | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. rugata | |
| Trisepticarpium | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. minutum | |
| Truncatisemen | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species T. sapotoides | |
| Ulospermum | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species U. hardingae | |
| Wheelera | Gen et sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A seed of uncertain placement. Type species W. lignicrusta | |
| Vitis tiffneyi | Sp nov | valid | Manchester | Middle Eocene | Clarno Formation | USA Oregon | A vitaceous species | |

## Arthropods
### Insects

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Acanthognathus poinari | Sp nov | Valid | Baroni Urbani & De Andrade | Burdigalian | Dominican amber | Dominican Republic | first fossil Acanthognathus species | |
| Aeshna andancensis | Sp nov | Valid | Nel & Brisac | Turolian | Montagne d'Andance | France | An Aeshna dragonfly | Aeshna andancensis |
| Aeshna cerdanica | Sp nov | Valid | Nel & Martínez-Delclòs | Vallesian | Bellver de Cerdanya | Spain | An Aeshna dragonfly | Aeshna cerdanica |
| Aeschnophlebia miocenica | Sp nov | Valid | Nel & Escuillé | Turolian | Montagne d'Andance | France | An Aeschnophlebia dragonfly | Aeschnophlebia miocenica |
| Anochetus ambiguus | Sp nov | Valid | De Andrade | Burdigalian | Dominican amber | Dominican Republic | A ponerin ant | Anochetus ambiguus |
| Anochetus conisquamis | Sp nov | Valid | De Andrade | Burdigalian | Dominican amber | Dominican Republic | A ponerin ant | Anochetus conisquamis |
| Anochetus dubius | Sp nov | Valid | De Andrade | Burdigalian | Dominican amber | Dominican Republic | A ponerin ant | Anochetus dubius |
| Anochetus exstinctus | Sp nov | Valid | De Andrade | Burdigalian | Dominican amber | Dominican Republic | A ponerin ant | Anochetus exstinctus |
| Anochetus intermedius | Sp nov | Valid | De Andrade | Burdigalian | Dominican amber | Dominican Republic | A ponerin ant | Anochetus intermedius |
| Anochetus lucidus | Sp nov | Valid | De Andrade | Burdigalian | Dominican amber | Dominican Republic | A ponerin ant | Anochetus lucidus |
| Calopteryx andancensis | Sp nov | Valid | Nel & Brisac | Turolian | Montagne d'Andance | France | A Calopteryx damselfly | Calopteryx andancensis |
| Dlusskyidris | Gen et comb nov. | Valid | Bolton | Santonian | Taimyr amber | Russia | A Sphecomyrminae ant. a new genus for Palaeomyrmex zherichini Dlussky 1975. | Palaeomyrmex zherichini |
| Forficula paleoligura | Sp nov | Valid | Nel, Albouy, Caussanel, & Jamet | Oligocene | | France | An earwig | Forficula paleoligura |
| Odontomachus pseudobauri | Sp nov. | Valid | De Andrade | Burdigalian | Dominican Amber | Dominican Republic | A myrmicin ant | Odontomachus pseudobauri |
| Odontomachus spinifer | Sp nov. | Valid | De Andrade | Burdigalian | Dominican Amber | Dominican Republic | A myrmicin ant | Odontomachus spinifer |
| Protoaristenymphes | Gen et sp nov | Valid | Nel & Henrotay | Toarcian | | Luxembourg | A Mesochrysopid lacewing, type species P. bascharagensis | Protoaristenymphes bascharagensis |
| Sympecma incerta | Comb. nov | Valid | Piton | Piacenzian | Lake Chambon | France | A Lestid damselfly, new combination for Lestes incertus | Sympecma incerta |

### Trilobites

| Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
| Parabouleia eldredgei | Sp nov | Valid | Edgecombe, Vaccari & Waisfeld | Early Devonian | Talacasto Formation | Argentina | A calmoniid | |
| Talacastops | Gen et sp nov | Valid | Edgecombe, Vaccari & Waisfeld | Early Devonian | Talacasto Formation | Argentina | A calmoniid, type species is T. zarelae | |

## Molluscs
### Bivalves

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Fordilla germanica | sp nov | Valid | Elicki | Early Cambrian | Zwetau Formation | Germany | early Cambrian bivalve species | |

## Fishes
### Newly named bony fishes

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Megalocoelacanthus | Gen. et sp. nov | valid | Schwimmer, Stewart & Williams | late Santonian to mid-Campanian | Blufftown Formation Niobrara Formation | United States | A latimeriid coelacanth. The type species is M. dobiei. | |

## Archosauromorphs

* Aff. Rebbachisaurus gastroliths documented.
* The "Talkeetna Mountains Hadrosaur" specimen was discovered in a quarry being excavated for road material. The quarry is near the Glenn Highway, approximately 150 miles northeast of Anchorage. That fall, excavation began, and was resumed in the summer of 1996.

### Newly named pseudosuchians

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Galtonia | Gen et sp nov | Valid | Hunt & Lucas | Late Triassic (late Carnian) | New Oxford Formation | United States | A relative of Revueltosaurus. | |
| Pekinosaurus | Gen et sp nov | Junior synonym | Hunt & Lucas | Late Triassic (late Carnian) | Cumnock Formation | United States | A junior synonym of Revueltosaurus. | |

### Newly named basal dinosauromorphs

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Marasuchus | Gen et comb nov | Valid | Sereno & Arcucci | Middle Triassic (Ladinian) | Chañares Formation | Argentina | A basal dinosauriform; new genus for "Lagosuchus" liloensis Romer (1972). | Marasuchus liloenisis |

### Newly named dinosaurs

Data courtesy of George Olshevsky's dinosaur genera list.
| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Afrovenator | Gen et sp nov | Valid | Sereno, Wilson, Larsson, Dutheil, & Sues | | Tiourarén Formation | Niger | | Afrovenator abakensis |
| Alwalkeria | Gen et comb nov | Valid | Chatterjee & Creisler | | Maleri Formation | India | | Alwalkeria |
| Alxasaurus | Gen et sp nov | Valid | Russell & Dong | | | Mongolia | | Alxasaurus |
| “Amphicoelicaudia” | | nomen nudum | Cheng | | | China | described as Huabeisaurus allocotus in 2000 | Huabeisaurus allocotus |
| Caenagnathasia | Gen et sp nov | Valid | Currie, Godfrey, & Nesov | Turonian - Coniacian | Bissekty Formation | Uzbekistan | | Caenagnathasia martinsoni |
| Cryolophosaurus | Gen et sp nov | Valid | Hammer & Hickerson | Pliensbachian | Hanson Formation | Ross Dependency | | Cryolophosaurus |
| Koparion | Gen et sp nov | Valid | Chure | Kimmeridgian | Morrison Formation | USA ( Utah) | | |
| Monolophosaurus | Gen et sp nov | Valid | Zhao & Currie | Bathonian - Callovian | Shishugou Formation | China | | Monolophosaurus jiangi |
| Mymoorapelta | Gen et sp nov | Valid | Kirkland & Carpenter | Kimmeridgian - Tithonian | Morrison Formation | USA ( Colorado) | | Mymoorapelta maysi |
| Pelecanimimus | Gen et sp nov | Valid | Perez-Moreno, Sanz, Buscalioni, Moratalla, Ortega, & Rasskin-Gutman | Barremian | Calizas de La Huérguina Formation | Spain | | Pelecanimimus |
| Phuwiangosaurus | Gen et sp nov | Valid | Martin, Buffetaut, & Suteethorn | Valanginian - Hauterivian | Sao Khua Formation | Thailand | | Phuwiangosaurus |
| Sinornithoides | Gen et sp nov | Valid | Russell & Dong | Aptian - Albian | Ejinhoro Formation | China | | Sinornithoides youngi |
| Sinraptor | Gen et sp nov | Valid | Currie & Zhao | Late Jurassic | Shishugou Formation | China | | Sinraptor dongi |
| Timimus | Gen et sp nov | Valid | Rich & Vickers-Rich | Albian | Eumeralla Formation | Australia | | Timimus hermani |
| “Tonouchisaurus” | | nomen nudum | Barsbold | | early Cretaceous | Mongolia | | |
| Yingshanosaurus | Gen et sp nov | Valid | Zhou | Kimmeridgian | Shaximiao Formation | China | | |

### Birds

| Name | Status | Novelty | Authors | Age | Unit | Location | Notes |
| Berruornis | Valid | Gen et sp nov. | Mourer-Chauviré | Late Paleocene | | France | A Strigidae. Type species B. orbisantiqui |
| Bubo osvaldoi | Valid | Sp. nov. | Arredondo & Olson | ?Pleistocene | Cave deposits | Cuba | A Strigidae. |
| Falco duboisi | Valid | Sp. nov. | Cowles | Holocene | Cave deposits | Réunion | A Falconidae. |
| Mascarenachen | jr synonym | Gen et sp nov. | Cowles | Holocene | Réunion | Réunion | An Anatidae, type species M. kervazoi; The species was transferred to Alopochen in 1999. |
| Mascarenotus grucheti | Valid | Sp. nov. | Mourer-Chauviré, Bour, Moutou, & Ribes | Holocene | | Réunion | A Strigidae. |
| Messelastur | Valid | Gen et sp nov. | Peters | Middle Eocene | Messel Formation | Germany Hessen | A Messelasturidae, Type species M. gratulator. |
| Microcarbo serventyorum | Valid | Sp. nov. | van Tets | Holocene | peat swamp | Australia Western Australia | A Phalacrocoracidae. |
| Neuquenornis | Valid | Gen. nov. et Sp. nov. | Chiappe & Calvo | Santonian-Coniacian | Rio Colorado Formation | Argentina | An Avisauridae, Type species N. volans. |
| Nycticorax borbonensis | jr synonym | Sp. nov. | Cowles | Holocene | | Réunion | An Ardeidae, junior synonym of Nycticorax duboisi. |
| Otogornis | Valid | Gen et sp nov. | Hou | Early Cretaceous | | China Inner Mongolia | A Longirostravidae, type species O. genghisi. |
| Presbyornis isoni | Valid | Sp. nov. | Olson | Late Paleocene | Aquia Formation | USA Maryland | A Presbyornithidae |
| Puffinus spelaeus | Valid | Sp. nov. | Holdaway & Worthy | Late Quaternary | South Island, cave deposits | New Zealand | A Procellariidae. |
| Stephanoaetus mahery | Valid | Sp. nov. | Goodman | Holocene. (Subrecent) | Madagascar | Madagascar | An Accipitridae. |
| Wanshuina | Valid | Gen et sp nov. | Hou | Paleocene | Doumu Formation | China | A Rallidae, type species W. lii |

## Plesiosaurs

* Plesiosaur gastroliths documented.

### New taxa

| Name | Status | Authors | Authors | Notes |
| Morturneria | Valid | Chatterjee Creisler | | |

## Pterosaurs
### New taxa

| Name | Status | Authors | Age | Unit | Location | Notes | Images |
| Arthurdactylus | Valid | Frey Martill | Early Cretaceous (late Aptian/early Albian) | Crato Formation | Brazil | A pteranodontoid pterosaur. | |
| Montanazhdarcho | Valid | Padian, K., Horner de Ricqlès, A.J. | Late Cretaceous (Judithian) | Two Medicine Formation | United States | An azhdarchid from Montana. | |
| "Wyomingopteryx" | nomen nudum | Bakker | Late Jurassic (Kimmeridgian-Tithonian) | Morrison Formation | United States | | |
| Zhejiangopterus | Valid | Cai, Z. Wei, F. | Late Cretaceous (early Campanian) | Tangshang Formation | China | A Chinese azhdarchid. | Zhejiangopterus |

## Other diapsids
### New taxa

| Name | Status | Authors | Authors | Age | Unit | Location | Notes |
| Lucianosaurus | Valid Non-Dinosaurian Taxon | Hunt | S. G. Lucas | Late Triassic (Norian) | Bull Canyon Formation | United States | |
| Tecovasaurus | Valid Non-Dinosaurian Taxon | Hunt | S. G. Lucas | Late Triassic (late Carnian) | Tecovas Formation | United States | |

## Footnotes

* Paleontology portal
* ↑ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
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* ↑ Manchester, S. R. (1999). "Biogeographical relationships of North American tertiary floras". Annals of the Missouri Botanical Garden. 86 (2): 472–522. doi:10.2307/2666183. JSTOR 2666183.
* ↑ Myers, J.; Erwin, D. (2015). "Deviacer pidemarmanii sp. nov. (Polygalaceae) from the Late Eocene–Early Oligocene Badger's Nose Paleoflora, Modoc County, California". International Journal of Plant Sciences. 176 (3): 259–268. doi:10.1086/679759. S2CID 84642512.
* ↑ Han, M.; Manchester, S. R.; Fu, Q.-Y.; Jin, J.-H.; Quan, C. (2018). "Paleogene fossil fruits of Stephania (Menispermaceae) from North America and East Asia". Journal of Systematics and Evolution. 56 (2): 81–91. doi:10.1111/jse.12288. S2CID 90749898.
* ↑ Call, V.B.; Dilcher, D.L. (1997). "The fossil record of Eucommia (Eucommiaceae) in North America". American Journal of Botany. 84 (6): 798–814. doi:10.2307/2445816. JSTOR 2445816. PMID 21708632. S2CID 20464075.
* ↑ Baroni Urbani, C.; De Andrade, M. L. (1994). "First description of fossil Dacetini ants with a critical analysis of the current classification of the tribe (Amber Collection Stuttgart: Hymenoptera, Formicidae. VI: Dacetini)". Stuttgarter Beiträge zur Naturkunde. Serie B (Geologie und Paläontologie). 198: 1–65.
* 1 2 3 4 Nel, A.; Martinez-Delclòs, X.; Escuillé, F; Brisac, P. (1994). "Names; Les Aeshnidae fossiles: Etat actuel des connaissances (Odonata, Anisoptera)". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 194 (2/3): 143–186. doi:10.1127/njgpa/194/1994/143. S2CID 247527646.
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* ↑ Grimaldi, D.; Agosti, D.; Carpenter, J. M. (1997). "New and rediscovered primitive ants (Hymenoptera, Formicidae) in Cretaceous amber from New Jersey, and their phylogenetic relationships". American Museum Novitates (3208): 1–43.
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