# Andricus

Andricus is a genus of oak gall wasps in the family Cynipidae.

## Life cycle

As in all Hymenoptera, sex-determination in species of the genus Andricus is governed by haplodiploidy: males develop from unfertilized eggs and are haploid, and females develop from fertilized eggs and are diploid.
Many species in the genus alternate between an asexual generation and a sexual generation.

## Species

The taxonomy of the genus Andricus is uncertain, with many of the approximately 375 species considered dubious. The following species are currently recognised in the genus Andricus:
* Andricus aciculatus Beutenmuller, 1909
* Andricus ahmeti Melika, Mutun & Dinç, 2014
* Andricus albipes Hartig, 1840
* Andricus albobalani Weld, 1926
* Andricus alniensis Folliot, 1964
* Andricus amblycerus (Giraud, 1859)
* Andricus amenti Giraud, 1859
* Andricus analis (Fonscolombe, 1832)
* Andricus anatolicus Melika, Mutun & Dinç, 2014
* Andricus archiboldi Melika & Abrahamson, 2021
* Andricus aries (Girauld, 1859)
* Andricus arslani Azmaz & Katılmış, 2022
* Andricus assarehi Melika & Sadeghi, 2008
* Andricus atkinsonae Melika, Stone, Sadeghi & Zargaran, 2008
* Andricus bakrachus Melika, Mutun & Dinç, 2014
* Andricus balanaspis (Weld, 1922)
* Andricus balanella (Weld, 1957)
* Andricus barriosi Medianero & Nieves-Aldrey, 2019
* Andricus biconicus Weld, 1959
* Andricus bimaculatus (Schenck, 1863)
* Andricus brachycentrus (Thomson, 1877)
* Andricus breviramuli Pujade-Villar, 2014
* Andricus brunneus Fullaway, 1911 — clustered gall wasp
* Andricus bulgaricus Vasileva-Symnalieva, 1978
* Andricus burgundus Giraud, 1859
* Andricus burnetti Dailey & Sprenger, 1983
* Andricus caliciformis (Giraud, 1859)
* Andricus californicus Ashmead, 1885
* Andricus callidoma (Hartig, 1841)
* Andricus capillatus (Weld, 1927)
* Andricus caputmedusae (Hartig, 1843)
* Andricus catalinensis Melika, Nicholls & Stone, 2021
* Andricus catilla (Darboux & Houard, 1907)
* Andricus cecconii Kieffer, 1901
* Andricus chapmanii Melika & Abrahamson, 2021
* Andricus chico Weld, 1957
* Andricus chinquapin (Fitch, 1859) — small oak spindle gall wasp
* Andricus chiricahuensis Melika, Nicholls & Stone, 2021
* Andricus chodjaii Melika, 2008
* Andricus chrysolepidicola Ashmead, 1896 — irregular spindle gall wasp
* Andricus cinnamomeus Ashmead, 1887
* Andricus clementinae (Giraud, 1859)
* Andricus coconinoensis Melika, Nicholls & Stone, 2021
* Andricus columbiensis Melika, Nicholls & Stone, 2021
* Andricus comata Weld, 1956
* Andricus confusus Lobato-Vila & Pujade-Villar, 2019
* Andricus confertus McCracken & Egbert, 1922 — convoluted gall wasp
* Andricus conglomeratus (Girauld, 1859)
* Andricus conificus (Hartig, 1843)
* Andricus cooki Melika, Nicholls & Stone, 2021
* Andricus coortus Weld, 1947 — club gall wasp
* Andricus coquilletti (Ashmead, 1897) - little oak apple gall wasp
* Andricus coriariformis Melika, Challis & Stone, 2008
* Andricus coriarius (Hartig, 1843)
* Andricus coronatus (Girauld, 1859)
* Andricus coronus Beutenmueller, 1907
* Andricus corruptrix (Schlechtendal, 1870)
* Andricus costaricensis Pujade-Villar & Melika, 2018
* Andricus costatus Weld, 1944
* Andricus crassicornis (Curtis, 1838)
* Andricus crispator Tschek, 1871
* Andricus cryptobius Wachtl, 1880
* Andricus csokai Melika & Tavakoli, 2008
* Andricus curtisii (Muller, 1870)
* Andricus curvator (Hartig, 1840)
* Andricus cydoniae Giraud, 1859
* Andricus deciduatus Weld, 1926
* Andricus dentimitratus (Rejtö, 1887)
* Andricus dimorphus (Beutenmuller, 1913) — clustered midrib gall wasp
* Andricus discalis (Weld, 1927)
* Andricus exclusus (Ratzeburg, 1844)
* Andricus fidelensis Kieffer, 1901
* Andricus fitzpatricki Melika & Abrahamson, 2021
* Andricus flavicornis Schenck, 1863
* Andricus flavus Pujade-Villar, Wang, Guo & Chen, 2014
* Andricus floridus Tavares, 1918
* Andricus foecundatrix (Hartig, 1840)
* Andricus flavohirtus Beutenmüller, 1913
* Andricus foliaformis Weld, 1927
* Andricus formosalis Weld, 1947
* Andricus formosanus Tang & Melika, 2009
* Andricus forni Pujade-Villar & Nicholls, 2020
* Andricus frondeum (Weld, 1957)
* Andricus fulviventris Schenck, 1863
* Andricus furnaceus Kinsey, 1920
* Andricus fuscicornis Hartig, 1841
* Andricus galeatus (Girauld, 1859)
* Andricus gallaecus Tavares, 1916
* Andricus gallaetinctoriae (Olivier, 1791)
* Andricus gallaeurnaeformis (Boyer de Fonscolombe, 1832)
* Andricus gemmeus (Giraud, 1859)
* Andricus gemmicola Kieffer, 1901
* Andricus geniculatus (Dufour, 1864)
* Andricus giardina Stefani, 1898
* Andricus glandulae (Hartig, 1840)
* Andricus glutinosus (Girauld, 1859)
* Andricus gracilicornis (Kieffer, 1900)
* Andricus grossulariae (Giraud, 1859)
* Andricus hartigi (Hartig, 1843)
* Andricus hastata (Kinsey, 1937)
* Andricus hedwigia (Kustenmacher, 1894)
* Andricus highlandensis Melika, Nicholls & Stone, 2021
* Andricus hispanicus (Hartig, 1856)
* Andricus howertoni Bassett, 1890
* Andricus hungaricus (Hartig, 1843)
* Andricus hyalinus Hartig, 1841
* Andricus hystrix Kieffer, 1897
* Andricus impropius Ballido & Pujade-Villar, 2003
* Andricus incertus Bassett, 1900
* Andricus indistinctus Bassett, 1890
* Andricus infectorius (Hartig, 1843)
* Andricus inflator Hartig, 1840
* Andricus insanus ](Westwood, 1837)
* Andricus istvani Melika, 2008
* Andricus kashiwaphilus Abe, 1998
* Andricus kollari (Hartig, 1843)
* Andricus korlevici Kieffer, 1902
* Andricus laevigatus Schenck, 1863
* Andricus lasius (Ashmead, 1896) - hairy gall wasp
* Andricus lateralis (Hartig, 1840)
* Andricus legitimus Wiebes-Rijks, 1980
* Andricus libani Melika, Challis & Stone, 2008
* Andricus lignicolus (Hartig, 1840)
* Andricus longipennis (Ashmead, 1887)
* Andricus lucidus (Hartig, 1843)
* Andricus luisieri Tavares, 1914
* Andricus luteicornis Kieffer, 1899
* Andricus malpighii (Adler, 1881)
* Andricus mammadovi Azmaz & Katilmis, 2021
* Andricus mamillaformis (Weld, 1926)
* Andricus marmoreus Kinsey, 1920
* Andricus maxwelli Bassett, 1890
* Andricus mayri (Wachtl, 1881)
* Andricus megatruncicolus Melika, 2008
* Andricus melikai Pujade-Villar & Kwast, 2002
* Andricus mellificus Melika, Nicholls & Stone, 2021
* Andricus mendocinensis Weld, 1957
* Andricus menkei Melika & Abrahamson, 2021
* Andricus miriami Shachar, 2015
* Andricus mitratus (Mayr, 1870)
* Andricus mogollonensis Melika, Nicholls & Stone, 2021
* Andricus montezumus Beutenmuller, 1913
* Andricus moniliatus Hartig, 1840
* Andricus moreae (Graeffe, 1905)
* Andricus morula Shachar, Inbar & Dorchin, 2017
* Andricus mukaigawae (Mukaigawa, 1913)
* Andricus multiplicatus (Girauld, 1859)
* Andricus murtfeldtae Ashmead, 1896
* Andricus nichollsi Melika & Stone, 2021
* Andricus niger (Fourcroy, 1785)
* Andricus nigricens (Gillette, 1888)
* Andricus nobrei Tavares, 1901
* Andricus nodifex Kieffer, 1900
* Andricus notholithocarpi Melika, Nicholls & Stone, 2018
* Andricus occultatus (Weld, 1926)
* Andricus octosporifex (Schrank, 1802)
* Andricus opertus (Weld, 1926) — fimbriate gall wasp
* Andricus ostrea (Hartig, 1840)
* Andricus pallidicornis (Curtis, 1838)
* Andricus pallipes (Schenck, 1863)
* Andricus panteli (Kieffer, 1896)
* Andricus paradoxus (Radoszkowski, 1866)
* Andricus partali Pujade-Villar, Cuesta-Porta & Hanson, 2022
* Andricus pedicellatus Kinsey, 1922 — hair stalk gall wasp
* Andricus peredurus (Kinsey, 1920)
* Andricus perfulvum (Weld, 1952)
* Andricus perlentus (Kinsey, 1937)
* Andricus petioli Hartig, 1843
* Andricus pictus (Hartig, 1856)
* Andricus pilula (Bassett, 1890)
* Andricus pilularis Weld, 1952
* Andricus pisiformis Beutenmueller, 1911
* Andricus polycerus (Girauld, 1859)
* Andricus prescotti Weld, 1952
* Andricus projectus Weld, 1952
* Andricus pseudoaries Melika, Stone & Sadegh, 2008
* Andricus pseudocurvator Tang & Melika, 2011
* Andricus pseudoinflator Tavares, 1901
* Andricus pujadevillari Melika, Stone, Sadeghi, Atkinson & Zargaran, 2008
* Andricus quadrilineatus (Hartig, 1840)
* Andricus quercuscalicis (Burgsdorf, 1783) — knopper gall
* Andricus quercuscalifornicus (Bassett, 1881) — California gall wasp
* Andricus quercuscorticis (Linnaeus, 1761)
* Andricus quercusfoliatus (Ashmead, 1881) — leafy oak gall wasp
* Andricus quercusformosus (Bassett, 1864)
* Andricus quercusfrondosus (Bassett, 1865)
* Andricus quercusinferus (Linnaeus, 1767)
* Andricus quercuslaurinus Melika and Pujade-Villar, 2009
* Andricus quercuspetioli (Linnaeus, 1758)
* Andricus quercuspetiolicola Bassett, 1863 — oak petiole gall wasp
* Andricus quercuspyramidalis (Landois, 1895)
* Andricus quercusradicis (Fabricius, 1798)
* Andricus quercusramuli (Linnaeus, 1761)
* Andricus quercussingularis Bassett, 1863 — small oak apple gall wasp
* Andricus quercustozae (Bosc, 1792)
* Andricus quercusstrobilanus (Osten-Sacken, 1862) — lobed oak gall wasp
* Andricus quercusutriculus (Bassett, 1881)
* Andricus reticulatus Bassett, 1890
* Andricus rhizoxenus (Ashmead, 1896)
* Andricus rhyzomae (Hartig, 1843)
* Andricus robustus Weld, 1926
* Andricus rotundula (Weld, 1952)
* Andricus rubripes (Thomson, 1877)
* Andricus ruficornis (Schenck, 1863)
* Andricus rufipes (Fabricius, 1804)
* Andricus rufiventris Schenck, 1863
* Andricus rufus (Thomson, 1877)
* Andricus rugatus Weld, 1926
* Andricus ruginosus Bassett, 1890
* Andricus sadeghii Melika, Stone, Atkinson & Aligolizade, 2008
* Andricus schencki Dalla Torre & Kieffer, 1910
* Andricus schickae Nicholls, Melika & Stone, 2021
* Andricus schoenroggei Melika & Stone, 2008
* Andricus schroeckingeri Wachtl, 1876
* Andricus scutella Weld, 1930
* Andricus seckendorffi (Wachtl, 1879)
* Andricus seminationis (Giraud, 1859)
* Andricus serotinus (Giraud, 1859)
* Andricus serricornis (Kinsey, 1922)
* Andricus sessilum Weld, 1926
* Andricus shuhuti Melika, Mutun & Dinç, 2013
* Andricus sieboldi (Hartig, 1843)
* Andricus singularis Mayr, 1870
* Andricus solitarius (Fonscolombe, 1832)
* Andricus songshui Tang & Melika, 2011
* Andricus sphaericus Pujade-Villar, 2016
* Andricus spicatus (Bassett, 1900)
* Andricus splendens Weld, 1919
* Andricus stefanii (Kieffer, 1897)
* Andricus stellaris Weld, 1926 — sunburst gall wasp
* Andricus stellulus (Burnett, 1974) — stellar gall wasp
* Andricus sternlichti Bellido, Pujade-Villar & Melika, 2003
* Andricus stonei Melika, Tavakoli & Sadeghi, 2014
* Andricus stramineus Weld, 1944
* Andricus stropus Ashmead, 1887
* Andricus strues (Kinsey, 1938)
* Andricus subterranea (Giraud, 1859)
* Andricus sufflator Mayr, 1882
* Andricus sulfureus Weld, 1926
* Andricus superfetationis (Giraud, 1859)
* Andricus surculi (Schrank, 1781)
* Andricus tavaresi Kieffer, 1904
* Andricus testaceipes (Hartig, 1840)
* Andricus testaceus (Gmelin, 1790)
* Andricus texanus Beutenmüller, 1909
* Andricus theophrasteus (Trotter, 1902)
* Andricus tomentosus (Trotter, 1901)
* Andricus torreyaensis Melika & Abrahamson, 2021
* Andricus toumeyi Weld, 1926
* Andricus trotteri Kieffer, 1898
* Andricus truncicola (Girauld, 1859)
* Andricus tubalis Weld, 1926
* Andricus tubularius Weld, 1926
* Andricus tumefaciens Pujade-Villar & Paretas-Martínez, 2014
* Andricus turcicus Melika, Mutun & Dinç, 2014
* Andricus turionum (Hartig, 1840)
* Andricus vacciniifoliae (Ashmead, 1896) — oak apple wasp
* Andricus villosulus (Gravenhorst, 1807)
* Andricus verensis Weld, 1955
* Andricus vindobonensis Müllner, 1901
* Andricus weldi (Beutenmüller, 1918)
* Andricus wheeleri Beutenmueller, 1907
* Andricus williami Melika, Nicholls & Stone, 2021
* Andricus wiltzae Fullaway, 1911
* Andricus zappellai Kieffer, 1901

## Previous species

These species were previously grouped in Andricus.
Currently in Druon:
* Druon flocculentum Lyon
* Druon fullawayi Beutenmüller, 1914
* Druon ignotum Bassett
* Druon linaria Kinsey
* Druon pattoni (Bassett, 1881)
* Druon protagion Kinsey
* Druon quercusflocci (Walsh, 1864)
* Druon quercuslanigerum Ashmead
Currently in Feron:
* Feron albicomus (Weld, 1952)
* Feron amphorus (Weld, 1926)
* Feron apiarium (Weld, 1944)
* Feron atrimentum Kinsey, 1922) — striped volcano gall wasp
* Feron bakkeri (Lyon, 1984)
* Feron caepula (Weld, 1926)
* Feron californicum (Beutenmueller, 1911)
* Feron clarkei (Bassett, 1890)
* Feron crystallinum (Bassett, 1900) — crystalline gall wasp
* Feron cylindratum (Kinsey, 1937)
* Feron discularis (Weld, 1926)
* Feron gigas (Kinsey, 1922)—saucer gall wasp
* Feron kingi (Bassett, 1900) — red cone gall wasp
* Feron parmula (Bassett, 1900) — disc gall wasp
* Feron pattersonae Fullaway, 1911 — plate gall wasp
* Feron tecturnarum Kinsey, 1920
* Feron tubifaciens Weld, 1926