# List of Cyphophthalmi species

This page lists all described species of the harvestman suborder Cyphophthalmi. Unless otherwise noted, information is taken from Giribet's 2000 taxonomic catalogue and from Giribet et al., 2012.

## Boreophthalmi
### Sironidae

* Arhesiro Karaman, 2022
   * Arhesiro clousi (Giribet & Shear, 2010)
   * Arhesiro sonoma (Shear, 1980)
* Cyphophthalmus Joseph, 1868
   * Cyphophthalmus beschkovi (Mitov, 1994)
   * Cyphophthalmus bithynicus (Gruber, 1969)
   * Cyphophthalmus conocephalus Karaman, 2009
   * Cyphophthalmus corfuanus (Kratochvíl, 1938)
   * Cyphophthalmus duricorius Joseph, 1868
   * Cyphophthalmus eratoae (Juberthie, 1968)
   * Cyphophthalmus ere Karaman, 2008
   * Cyphophthalmus gjorgjevici (Hadži, 1933)
   * Cyphophthalmus gordani Karaman, 2009
   * Cyphophthalmus hlavaci Karaman, 2009
   * Cyphophthalmus klisurae (Hadži, 1973)
   * Cyphophthalmus kratochvili Karaman, 2009
   * Cyphophthalmus markoi Karaman, 2008
   * Cyphophthalmus martensi Karaman, 2009
   * Cyphophthalmus minutus (Kratochvíl, 1938)
   * Cyphophthalmus montenegrinus (Hadzi, 1973)
   * Cyphophthalmus neretvanus Karaman, 2009
   * Cyphophthalmus noctiphilus (Kratochvíl, 1940)
   * Cyphophthalmus nonveilleri Karaman, 2008
   * Cyphophthalmus ognjenovici Karaman, 2009
   * Cyphophthalmus ohridanus (Hadži, 1973)
   * Cyphophthalmus paradoxus (Kratochvíl, 1958)
   * Cyphophthalmus paragamiani Karaman, 2009
   * Cyphophthalmus rumijae Karaman, 2009
   * Cyphophthalmus serbicus (Hadži, 1973)
   * Cyphophthalmus silhavyi (Kratochvíl, 1938)
   * Cyphophthalmus solentiensis Dreszer, Raða & Giribet, 2015
   * Cyphophthalmus teyrovskyi (Kratochvíl, 1938)
   * Cyphophthalmus thracicus Karaman, 2009
   * Cyphophthalmus trebinjanus Karaman, 2009
   * Cyphophthalmus yalovensis (Gruber, 1969)
   * Cyphophthalmus zetae Karaman, 2009
* Holosiro Ewing, 1923
   * Holosiro acaroides Ewing, 1923
   * Holosiro calaveras (Giribet & Shear, 2010)
   * Holosiro ewingi Karaman, 2022
   * Holosiro shasta (Giribet & Shear, 2010)
* Iberosiro de Bivort & Giribet, 2004
   * Iberosiro distylos de Bivort & Giribet, 2004
   * Iberosiro rosae Giribet, Merino-Sáinz & Benavides, 2017
* Neosiro Newell, 1943
   * Neosiro boyerae (Giribet & Shear, 2010)
   * Neosiro exilis (Hoffman, 1963)
   * Neosiro kamiakensis Newell, 1943
   * Neosiro ligiae (Giribet, 2017)
   * Neosiro martensi Karaman, 2022
   * Neosiro richarti (Benavides & Giribet, 2017)
   * [Also †Neosiro balticus (Dunlop & Mitov, 2011) (fossil, Baltic amber)]
* Odontosiro Juberthie, 1961
   * Odontosiro lusitanicus Juberthie, 1961
* Paramiopsalis Juberthie, 1962
   * Paramiopsalis anadonae Giribet, Merino-Sáinz & Benavides, 2017
   * Paramiopsalis eduardoi Murienne & Giribet, 2009
   * Paramiopsalis ramblae Benavides & Giribet, 2017
   * Paramiopsalis ramulosa Juberthie, 1962
* Siro Latreille, 1796
   * Siro carpaticus Rafalski, 1956
   * Siro crassus Novak & Giribet, 2006
   * Siro franzi Karaman & Raspotnig, 2022
   * Siro ozimeci Karaman, 2022
   * Siro rubens Latreille, 1804
   * Siro richarti Benavides & Giribet, 2017
   * Siro valleorum Chemini, 1990
   * [Also †Siro platypedibus Dunlop & Giribet, 2003 (fossil, Baltic amber)]
* Suzukielus Juberthie, 1970
   * Suzukielus sauteri (Roewer, 1916)
Insetae sedis:
* Marwe Shear, 1985
   * Marwe coarctata Shear, 1985

### Stylocellidae
#### Fangensinae

* Fangensis Rambla, 1994
   * Fangensis cavernarus Schwendinger & Giribet, 2005
   * Fangensis leclerci Rambla, 1994
   * Fangensis spelaeus Schwendinger & Giribet, 2005
* Giribetia Clouse, 2012
   * Giribetia insulanus (Schwendinger & Giribet, 2005)

#### Leptopsalidinae

* Leptopsalis Thorell, 1882
   * Leptopsalis beccarii Thorell, 1882
   * Leptopsalis dumoga (Shear, 1993)
   * Leptopsalis foveolata Clouse & Schwendinger, 2012
   * Leptopsalis hillyardi (Shear, 1993)
   * Leptopsalis javana (Thorell, 1882)
   * Leptopsalis laevichelis (Roewer, 1942)
   * Leptopsalis lydekkeri (Clouse & Giribet, 2007)
   * Leptopsalis modesta (Hansen & Sørensen, 1904)
   * Leptopsalis novaguinea (Clouse & Giribet, 2007)
   * Leptopsalis pangrango (Shear, 1993)
   * Leptopsalis ramblae (Giribet, 2002)
   * Leptopsalis sedgwicki (Shear, 1979)
   * Leptopsalis sulcata (Hansen & Sørensen, 1904)
   * Leptopsalis tambusisi (Shear, 1993)
   * Leptopsalis thorellii (Hansen & Sørensen, 1904)
   * Leptopsalis weberii (Hansen & Sørensen, 1904)
* Miopsalis Thorell, 1890
   * Miopsalis collinsi (Shear, 1993)
   * Miopsalis dillyi Schmidt, Clouse & Sharma, 2020
   * Miopsalis globosa (Schwendinger & Giribet, 2004)
   * Miopsalis gryllospeca (Shear, 1993)
   * Miopsalis kinabalu (Shear, 1993)
   * Miopsalis leakeyi (Shear, 1993)
   * Miopsalis lionota (Pocock, 1897)
   * Miopsalis mulu (Shear, 1993)
   * Miopsalis pocockii (Hansen & Sørensen, 1904)
   * Miopsalis pulicaria Thorell, 1890
   * Miopsalis sabah (Shear, 1993)
   * Miopsalis silhavyi (Rambla, 1991)
   * Miopsalis tarumpitao (Shear, 1993)

#### Stylocellinae

* Meghalaya Giribet, Sharma, & Bastawade, 2007
   * Meghalaya annandalei Giribet, Sharma, & Bastawade, 2007
* Stylocellus Westwood, 1874
   * Stylocellus lornei Clouse, 2012
   * Stylocellus spinifrons Roewer, 1942 (nomen dubium) [unidentifiable juvenile]
   * Stylocellus sumatranus Westwood, 1874

## Scopulophthalmi
### Pettalidae

* Aoraki Boyer & Giribet, 2007
   * Aoraki calcarobtusa (Forster, 1952)
   * Aoraki crypta (Forster, 1948)
   * Aoraki denticulata (Forster, 1948)
      * Aoraki denticulata denticulata (Forster, 1948)
      * Aoraki denticulata major (Forster, 1948)
   * Aoraki granulosa (Forster, 1952)
   * Aoraki healyi (Forster, 1948)
   * Aoraki inerma (Forster, 1948)
   * Aoraki longitarsa (Forster, 1952)
   * Aoraki stephenesis (Forster, 1952)
   * Aoraki tumidata (Forster, 1948)
   * Aoraki westlandica (Forster, 1952)
* Archaeopurcellia Giribet, Shaw, Lord & Derkarabetian, 2022
   * Archaeopurcellia eureka Giribet, Shaw, Lord & Derkarabetian, 2022
* Austropurcellia Juberthie, 1988
   * Austropurcellia absens Boyer & Popkin-Hall, 2015
   * Austropurcellia acuta Popkin-Hall & Boyer, 2014
   * Austropurcellia alata Boyer & Reuter, 2012
   * Austropurcellia arcticosa (Cantrell, 1980)
   * Austropurcellia barbata Popkin-Hall & Boyer, 2014
   * Austropurcellia cadens Baker & Boyer, 2015
   * Austropurcellia capricornia (Davies, 1977)
   * Austropurcellia clousei Boyer, Baker & Popkin-Hall, 2015
   * Austropurcellia culminis Boyer & Reuter, 2012
   * Austropurcellia daviesae (Juberthie, 1989)
   * Austropurcellis despectata Boyer & Reuter, 2012
   * Austropurcellia finniganensis Popkin-Hall, Jay & Boyer, 2016
   * Austropurcellia forsteri (Juberthie, 2000)
   * Austropurcellia fragosa Popkin-Hall, Jay & Boyer, 2016
   * Austropurcellia giribeti Boyer & Quay, 2015
   * Austropurcellia megatanka Jay, Coblens & Boyer, 2016
   * Austropurcellia monteithi Jay, Popkin-Hall, Coblens & Boyer, 2016
   * Austropurcellia nuda Popkin-Hall, Jay & Boyer, 2016
   * Austropurcellia riedeli Jay, Oberski & Boyer, 2016
   * Austropurcellia scoparia Juberthie, 1988
   * Austropurcellia sharmai Boyer & Quay, 2015
   * Austropurcellia superbensis Popkin-Hall & Boyer, 2014
   * Austropurcellia tholei Baker & Boyer, 2015
   * Austropurcellia vicina Boyer & Reuter, 2012
   * Austropurcellia woodwardi (Forster, 1955)
* Chileogovea Roewer, 1961
   * Chileogovea jocasta Shear, 1993
   * Chileogovea oedipus Roewer, 1961
* Karripurcellia Giribet, 2003
   * Karripurcellia peckorum Giribet, 2003
   * Karripurcellia sierwaldae Giribet, 2003
* Manangotria Shear & Gruber, 1996
   * Manangotria taolanaro Shear & Gruber, 1996
* Neopurcellia Forster, 1948
   * Neopurcellia salmoni Forster, 1948
* Parapurcellia Rosas Costa, 1950
   * Parapurcellia amatola de Bivort & Giribet, 2010
   * Parapurcellia convexa de Bivort & Giribet, 2010
   * Parapurcellia fissa (Lawrence, 1939)
   * Parapurcellia minuta de Bivort & Giribet, 2010
   * Parapurcellia monticola (Lawrence, 1939)
   * Parapurcellia natalia de Bivort & Giribet, 2010
   * Parapurcellia rumpiana (Lawrence, 1933)
   * Parapurcellia peregrinator (Lawrence, 1963)
   * Parapurcellia silvicola (Lawrence, 1939)
   * Parapurcellia staregai de Bivort & Giribet, 2010
   * Parapurcellia transvaalica (Lawrence, 1963)
* Pettalus Thorell, 1876
   * Pettalus brevicauda Pocock, 1897
   * Pettalus cimiciformis (Cambridge, 1875)
   * Pettalus lampetides Sharma & Giribet 2006
   * Pettalus thwaitesi Sharma, Karunarathna & Giribet, 2009
* Purcellia Hansen & Sørensen, 1904
   * Purcellia argasiformis (Lawrence, 1931)
   * Purcellia griswaldi de Bivort & Giribet, 2010
   * Purcellia illustrans Hansen and Sørensen, 1904
   * Purcellia lawrencei de Bivort & Giribet, 2010
   * Purcellia leleupi Staręga, 2008
* Rakaia Hirst, 1926
   * Rakaia antipodiana Hirst, 1926
   * Rakaia collaria Roewer, 1942
   * Rakaia dorothea Phillips and Grimmett, 1932
   * Rakaia florensis (Forster, 1948)
   * Rakaia isolata Forster, 1952
   * Rakaia lindsayi Forster, 1952
   * Rakaia macra Boyer and Giribet, 2003
   * Rakaia magna Forster, 1948
      * Rakaia magna australis Forster, 1952
      * Rakaia magna magna Forster, 1948
   * Rakaia media Forster, 1948
      * Rakaia media media Forster, 1948
      * Rakaia media insula Forster, 1952
   * Rakaia minutissima (Forster, 1948)
   * Rakaia pauli Forster, 1952
   * Rakaia solitaria Forster, 1948
   * Rakaia sorenseni Forster, 1952
      * Rakaia sorenseni digitata Forster, 1952
      * Rakaia sorenseni sorenseni Forster, 1952
   * Rakaia stewartiensis Forster, 1948
   * Rakaia uniloca Forster, 1952

## Sternophthalmi
### Neogoveidae

* Brasiliogovea Martens, 1969
   * Brasiliogovea aphantostylus Benavides, Hormiga, & Giribet, 2019
   * Brasiliogovea chiribiqueta Benavides & Giribet, 2013
   * Brasiliogovea microphaga Martens, 1969
   * Brasiliogovea microstylus Benavides, Hormiga, & Giribet, 2019
   * Brasiliogovea yacambuensis Benavides, Hormiga, & Giribet, 2019
* Canga DaSilva, Pinto-da-Rocha & Giribet, 2010
   * Canga renatae DaSilva, Pinto-da-Rocha & Giribet, 2010
* Huitaca Shear, 1979
   * Huitaca boyacanensis Benavides & Giribet, 2013
   * Huitaca bitaco Benavides & Giribet, 2013
   * Huitaca caldas Benavides & Giribet, 2013
   * Huitaca depressa Benavides & Giribet, 2013
   * Huitaca sharkeyi Benavides & Giribet, 2013
   * Huitaca tama Benavides & Giribet, 2013
   * Huitaca ventralis Shear, 1979
* Leggogovia Benavides & Giribet, 2019
   * Leggogovia pabsgarnoni (Legg. 1990)
* Metagovea Rosas-Costa, 1950
   * Metagovea disparunguis Rosas Costa, 1950
   * Metagovea ligiae Giupponi & Kury, 2015
   * Metagovea matapi Benavides, Hormiga, & Giribet, 2019
   * Metagovea philipi Goodnight & Goodnight, 1980
   * Metagovea planada Benavides, Hormiga, & Giribet, 2019
* Metasiro Juberthie, 1960
   * Metasiro americanus (Davis, 1933)
   * Metasiro savannahensis Clouse & Wheeler, 2014
   * Metasiro sassafrasensis Clouse & Wheeler, 2014
* Microgovia Benavides, Hormiga, & Giribet, 2019
   * Microgovia chenepau Benavides, Hormiga, & Giribet, 2019
   * Microgovia oviformis Martens, 1969
* Neogovea Hinton, 1938
   * Neogovea branstetteri Benavides, Hormiga, & Giribet, 2019
   * Neogovea enigmatica Martens, 2019
   * Neogovea hormigai Benavides & Giribet, 2013
   * Neogovea immsi Hinton, 1938
   * Neogovea kamakusa Shear, 1977
   * Neogovea kartabo Shear, 1977
   * Neogovea matawai Benavides, Hormiga, & Giribet, 2019
   * Neogovea virginie Jocqué & Jocqué, 2011
* Parogovia Hansen, 1921
   * Parogovia gabonica (Juberthie, 1969)
   * Parogovia montealensis Benavides & Giribet, 2019
   * Parogovia parasironoides Hiřman, Kotyk, Kotyková Varadínová,& Šťáhlavský, 2018
   * Parogovia prietoi Benavides & Giribet, 2019
   * Parogovia putnami Benavides & Giribet, 2019
   * Parogovia sironoides Hansen, 1921
* Tucanogovea Karaman, 2014
   * Tucanogovea schusteri Karaman, 2014
* Waiwaigovia Benavides, Hormiga, & Giribet, 2019
   * Waiwaigovia shultzi Benavides, Hormiga, & Giribet, 2019

### Ogoveidae

* Ogovea Roewer, 1923
   * Ogovea cameroonensis Giribet & Prieto, 2003
   * Ogovea grossa (Hansen & Sørensen, 1904)
   * Ogovea nasuta (Hansen, 1921)

### Troglosironidae

* Troglosiro Juberthie, 1979
   * Troglosiro aelleni Juberthie, 1979
   * Troglosiro brevifossa Sharma & Giribet, 2009
   * Troglosiro dogny Giribet & Baker, 2021
   * Troglosiro juberthiei Shear, 1993
   * Troglosiro longifossa Sharma & Giribet, 2005
   * Troglosiro monteithi Sharma & Giribet, 2009
   * Troglosiro ninqua Shear, 1993
   * Troglosiro oscitatio Sharma & Giribet, 2009
   * Troglosiro pin Giribet, Baker & Sharma, 2021
   * Troglosiro platnicki Shear, 1993
   * Troglosiro pseudojuberthiei Giribet, Baker & Sharma, 2021
   * Troglosiro raveni Shear, 1993
   * Troglosiro sharmai Giribet & Baker, 2021
   * Troglosiro sheari Sharma & Giribet, 2009
   * Troglosiro tillierorum Shear, 1993
   * Troglosiro urbanus Sharma & Giribet, 2009
   * Troglosiro wilsoni Sharma & Giribet, 2009

## Incertae sedis

* Ankaratra Shear & Gruber, 1996
   * Ankaratra franzi Shear & Gruber, 1996
* Shearogovea Giribet, 2011
   * Shearogovea mexasca (Shear, 1977)
Parasironidae Karaman, Mitov & Snegovaya, 2024
* Cimmerosiro Karaman, Mitov & Snegovaya, 2024
   * Cimmerosiro juberthiei Karaman, Mitov & Snegovaya, 2024
   * Cimmerosiro krivolutskyi Karaman, Mitov & Snegovaya, 2024
   * Cimmerosiro rhodiensis Karaman, Mitov & Snegovaya, 2024
* Ebrosiro Karaman, Mitov & Snegovaya, 2024
   * Ebrosiro coiffaiti (Juberthie, 1956)
* Parasiro Hansen & Sørensen, 1904
   * Parasiro corsicus (Simon, 1872)
* Tirrenosiro Karaman, Mitov & Snegovaya, 2024
   * Tirrenosiro axeli Karaman, Mitov & Snegovaya, 2024
   * Tirrenosiro minor (Juberthie, 1958)