# Clavus (gastropod)

Clavus is a genus of small sea snails, marine gastropod mollusks in the family Drilliidae.

## Description

Apart from the general characteristics of the genera in the family Drilliidae, the species in the genus Clavus have a shell with peripheral tubercles, spines or wing-like processes. The aperture is rather large. The outer lip is produced below the sinus.
The shell of Clavus is characterised by the following features:— Flat indefinite fasciole, indicated only by the curve of growth lines. A smooth subulate protoconch. An insinuation of the outer lip, near the base, like that of Strombus. A major sculpture of prominent arched scales on the shoulder, and a minor sculpture of dense, microscopic, waved, spiral striae.
Most species in this genus have a dorsal varix. But this is absent in a few species such as Clavus beckii, Clavus humilis and Clavus pica.
G.W. Tryon correctly preserved the genus for smooth specimens with a short body whorl, long spire, nodulous shoulder, no spiral sculpture, a wide, deep anal sulcus adjacent to the suture and, in the completely adult, a marked subsutural callus on the body.

## Distribution

The species in this genus occur in the Red Sea, In the Indian Ocean off Mozambique; also off Indonesia, New Caledonia, Papua New Guinea, the Philippines, Solomon Islands, Japan and Australia (New South Wales, Northern Territory, Queensland, Western Australia).

## Species

According to the World Register of Marine Species (WoRMS) the following species with valid names are included within the genus Clavus :
* Clavus aenigmaticus Wells, 1991
* Clavus aglaia (Dall, 1918)
* Clavus alboroseus Horro, Gori, Rosado & Rolán, 2021
* Clavus albotuberculatus (Schepman, 1889)
* Clavus andreolbrichi Fedosov & Puillandre, 2020
* Clavus androyensis (Bozzetti, 2007)
* Clavus angulatus Stahlschmidt, Poppe & Tagaro, 2018
* Clavus basipunctatus (Kilburn, 1988)
* Clavus beckii (Reeve, 1842)
* Clavus berenice (Dall, 1918) (synonym: Clavus spinosa Smith, E.A., 1882 )
* Clavus biancae (Bozzetti, 2008)
* Clavus bilineatus (Reeve, 1845)
* Clavus boucheti Kilburn, Fedosov & Kantor, 2014
* Clavus brianmayi Fedosov & Puillandre, 2020
* Clavus burnupi (G. B. Sowerby III, 1897)
* Clavus cadenasi (Clench & Aguayo, 1939)
* Clavus canalicularis (Röding, 1798)
* Clavus candens (Smith E. A., 1879)
* Clavus cantharis (Reeve, 1845)
* Clavus clara (Reeve, 1845)
* Clavus clavata (Sowerby II, 1870)
* Clavus cloveri (Poppe, Tagaro & Goto, 2018)
* Clavus coffea (Smith E. A., 1882)
* Clavus cygneus (Melvill & Standen, 1897)
* Clavus davidgilmouri Fedosov & Puillandre, 2020
* Clavus delphineae Kilburn, Fedosov & Kantor, 2014
* Clavus devexistriatus Kilburn, Fedosov & Kantor, 2014
* Clavus dolichurus Stahlschmidt, Poppe & Tagaro, 2018
* Clavus dubrucqueae T. Cossignani, 2021
* Clavus ebur (Reeve, 1845)
* Clavus exasperatus (Reeve, 1843)
* Clavus exilis (Pease, 1868)
* Clavus falcicosta (Barnard, 1958)
* Clavus flammulatus Montfort, 1810
* Clavus formosus (Reeve, 1846)
* Clavus fulvus (Hinds, 1843)
* Clavus fusconitens (Sowerby III, 1901)
* Clavus glaucozona Kilburn & Dekker, 2008
* Clavus groschi Kilburn, 1988
* Clavus herberti (Kilburn, 1988)
* Clavus heryi (Bozzetti, 2007)
* Clavus hewittae Wells, 1991
* Clavus hottentotus (E. A. Smith, 1882)
* Clavus humilis (E. A. Smith, 1879)
* Clavus hylikos Kilburn, Fedosov & Kantor, 2014
* Clavus idae (Poppe, Tagaro & Goto, 2018)
* Clavus infrafusca (G.B. Sowerby III, 1893)
* Clavus infuscatus Kilburn & Dekker, 2008
* Clavus isibopho (Kilburn, 1988)
* Clavus isowai Poppe, Tagaro & Goto, 2018
* Clavus japonicus (Lischke, 1869)
* Clavus johnsoni (Bartsch, 1934)
* Clavus juani Horro, Gori, Rosado & Rolán, 2021
* Clavus kirkhammetti Fedosov & Puillandre, 2020
* Clavus laetus (Hinds, 1843)
* Clavus lamberti (Montrouzier, 1860)
* Clavus maestratii Kilburn, Fedosov & Kantor, 2014
* Clavus malva (Morassi, 1998)
* Clavus minutissimus Stahlschmidt, Poppe & Tagaro, 2018
* Clavus moquinianus (Montrouzier, 1874)
* Clavus nodifera (Pease, 1860)
* Clavus nodulosa (Pease, 1863)
* Clavus obliquatus (Reeve, 1845)
* Clavus obliquicostatus (Reeve, 1845)
* Clavus occiduus Wells, 1991
* Clavus papilio (Kilburn, 1988)
* Clavus paroeca (Melvill, 1923)
* Clavus particolor Stahlschmidt, Poppe & Tagaro, 2018
* Clavus peristera Melvill, 1927
* Clavus pica (Reeve, 1843)
* Clavus picoides Kilburn, Fedosov & Kantor, 2014
* Clavus powelli Kay, 1979
* Clavus protentus Hervier, 1896
* Clavus pulicarius Wells, 1991
* Clavus pusilla (Garrett, 1873)
* Clavus putillus (Reeve, 1845)
* Clavus quadratus (Kilburn, 1988)
* Clavus rissoiniformis Kay, 1979
* Clavus roseofuscus (Bozzetti, 2007)
* Clavus rugizonatus Hervier, 1896
* Clavus sacra (Reeve, L.A., 1845)
* Clavus similis Stahlschmidt, Poppe & Tagaro, 2018
* Clavus squamiferus Kilburn, Fedosov & Kantor, 2014
* Clavus subobliquatus (E. A. Smith, 1879)
* Clavus subtilifasciatus Horro, Gori, Rosado & Rolán, 2021
* Clavus suduirauti Bozzetti, 1997
* Clavus sulekile (Kilburn, 1988)
* Clavus triantoniorum Horro, Gori, Rosado & Rolán, 2021
* Clavus unizonalis (Lamarck, 1822)
* Clavus velcolorum Horro, Gori, Rosado & Rolán, 2021
* Clavus vibicinus (Helbling, 1779)
* Clavus virginieae Kilburn, Fedosov & Kantor, 2014
* Clavus wilmeri (Smith E. A., 1879)

### Species brought into synonymy

* Clavus abdera (Dall, 1919): synonym of Crassispira abdera (Dall, 1919)
* Clavus aeneus (Hedley, 1922): synonym of Antiguraleus aeneus (Hedley, 1922)
* Clavus alabaster (Reeve, 1843): synonym of Inquisitor alabaster (Reeve, 1843)
* Clavus auriculifera Lamarck, J.B.P.A. de, 1816: synonym of Clavus canalicularis (Röding, 1798)
* Clavus candidulus Hedley, 1922: synonym of Splendrillia candidulus (Hedley, 1922)
* Clavus costatus Hedley, 1922: synonym of Graciliclava costata (Hedley, 1922)
* Clavus crassa (E.A. Smith, 1888): synonym of Clavus aglaia (Dall, 1918)
* Clavus enna (Dall, 1918): synonym of Drillia enna (Dall, 1918)
* Clavus filicinctus (E. A. Smith, 1882): synonym of Horaiclavus filicinctus (E. A. Smith, 1882)
* Clavus flavidulus (Lamarck, 1822): synonym of Clathrodrillia flavidula (Lamarck, 1822)
* Clavus flexus Shuto, 1983: synonym of Plagiostropha flexus (Shuto, 1983)
* Clavus gibberulus Hervier, 1896 : synonym of Drillia gibberulus (Hervier, 1896)
* Clavus inclinata (Sowerby III, 1893): synonym of Iredalea inclinata (Sowerby III, 1893)
* Clavus lacertosus (Hedley, 1922): synonym of Bathytoma lacertosus Hedley, 1922
* Clavus leforestieri Hervier, 1896: synonym of Clavus obliquicostatus (Reeve, 1845)
* Clavus maravignae (Bivona Ant. in Bivona And., 1838): synonym of Crassopleura maravignae (Bivona Ant. in Bivona And., 1838)
* Clavus mighelsi Kay, 1979: synonym of Pyrgocythara mighelsi (Kay, 1979)
* Clavus mighelsi of authors: synonym of Clavus humilis (E. A. Smith, 1879)
* Clavus opalus (Reeve, 1845): synonym of Plagiostropha opalus (Reeve, 1845)
* Clavus pseudoprincipalis (Yokoyama, 1920): synonym of Inquisitor pseudoprincipalis (Yokoyama, 1920)
* Clavus pulchellus (Reeve, 1845): synonym of Fenimorea halidorema Schwengel, 1940
* Clavus quintuplex (Melvill, 1927): synonym of Plagiostropha quintuplex Melvill, 1927
* Clavus regius Habe & Murakami, 1970: synonym of Drillia regia (Habe & Murakami, 1970)
* Clavus undatus (Hedley, 1907): synonym of Austroclavus undatus (Hedley, 1907)
* Clavus viduus (Reeve, 1845): synonym of Clavus unizonalis (Lamarck, 1822)

### Species inquirenda

* Clavus acuminata : species inquirenda
* Clavus auriculifera Lamarck : species inquirenda
The Indo-Pacific Molluscan Database adds the following species with names in current use :
* Clavus siebenrocki (Sturany, 1903)

### Subgenus Clavus

* Clavus intermaculatus (E. A. Smith, 1879)