# 2022 in paleomalacology

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This list of fossil molluscs described in 2022 is a list of new taxa of fossil molluscs that were described during the year 2022, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2022.

## Ammonites
### New taxa

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Acanthohoplites albedocostatus | Sp. nov | In press | Matsukawa | Early Cretaceous | | Japan | |
| Acantholytoceras costaevariavilis | Sp. nov | In press | Matsukawa | Early Cretaceous | | Japan | |
| Aulatornoceras beyrichi | Sp. nov | Valid | Söte & Becker | Devonian (Frasnian) | Büdesheim Formation | Germany | A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023. |
| Buedesheimites | Gen. et sp. nov | Valid | Söte & Becker | Devonian (Frasnian) | Büdesheim Formation | Germany | A member of the family Tornoceratidae. Genus includes new species B. housei. Announced in 2022; the final article version was published in 2023. |
| Cadoceras (Catacadoceras) imlayi | Sp. nov | Valid | Kiselev | Middle Jurassic (Callovian) | Chinitna Formation | United States ( Alaska) | |
| Cadoceras kniazevi | Sp. nov | Valid | Kiselev | Middle Jurassic (Callovian) | | Russia | |
| Cadoceras (Paracadoceras) pseudofrearsi | Sp. nov | Valid | Kiselev | Middle Jurassic (Callovian) | | Russia | |
| Cadomites zelenchukensis | Sp. nov | Valid | Mitta | Middle Jurassic (Bathonian) | | Russia ( Karachay-Cherkessia) | |
| Collignoniceras rhodanicum | Sp. nov | Valid | Amédro, Baudouin & Delanoy in Amédro et al. | Late Cretaceous (Turonian) | | France | |
| Crassotornoceras tenue | Sp. nov | Valid | Söte & Becker | Devonian (Frasnian) | Büdesheim Formation | Germany | A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023. |
| Crioceratites tepehuaensis | Sp. nov | Valid | Esquivel-Macías et al. | Early Cretaceous | | Mexico | |
| Cryptomorphiceras | Gen. et comb. et 2 sp. nov | Valid | Marchesi, Balini & Jenks | Late Triassic (Carnian) | | United States ( Nevada) | A member of Trachyceratinae. The type species is C. compressum (Johnston, 1941); genus also includes new species C. crassum, and C. levicostatum. Announced in 2022; the final article version was published in 2023. |
| Dactylioceras kopiki | Sp. nov | Valid | Rogov in Rogov & Lutikov | Early Jurassic (Toarcian) | | Canada ( Northwest Territories)  Norway  United States ( Alaska)  Russia? | A member of the family Dactylioceratidae. |
| Diplesioceras fischeri | Sp. nov | Valid | Dietze, Schweigert & Chandler | Middle Jurassic (Bajocian) | Oolithe de Bayeux Formation | France | |
| Egoianites jauberti | Sp. nov | Valid | Delanoy et al. | Early Cretaceous (Barremian) | | France | A member of the family Heteroceratidae. |
| Epitornoceras transmediterraneum | Sp. nov | Valid | Korn & Bockwinkel | Devonian (Givetian) | | Morocco | A member of the family Tornoceratidae. |
| Forresteria (Harleites) seidli | Sp. nov | Valid | Summesberger et al. | Late Cretaceous (Santonian) | Gosau Group (Streiteck Formation) | Austria | |
| Heteroceras arnaudi | Sp. nov | Valid | Delanoy et al. | Early Cretaceous (Barremian) | | France | A member of the family Heteroceratidae. |
| Heteroceras munierchalmasi | Sp. nov | Valid | Delanoy et al. | Early Cretaceous (Barremian) | | France | A member of the family Heteroceratidae. |
| Heteroceras penagei | Sp. nov | Valid | Delanoy et al. | Early Cretaceous (Barremian) | | France | A member of the family Heteroceratidae. |
| Hildoceras appenninicus | Sp. nov | Valid | Ridente | Early Jurassic (Toarcian) | | Italy | |
| Kepplerites (Gowericeras) copernici | Sp. nov | Valid | Mönnig & Dietl | Middle Jurassic (Callovian) | | Germany | |
| Kepplerites (Gowericeras) halleyi | Sp. nov | Valid | Mönnig & Dietl | Middle Jurassic (Callovian) | | France  Germany  Russia ( Kostroma Oblast) | |
| Kepplerites (Toricellites) besseli | Sp. nov | Valid | Mönnig & Dietl | Middle Jurassic (Callovian) | | Germany | |
| Kutatissites sudai | Sp. nov | In press | Matsukawa | Early Cretaceous | | Japan | |
| Lentitornoceras | Gen. et sp. nov | Valid | Korn & Bockwinkel | Devonian (Givetian) | Red Ironstone Formation | Germany | A member of the family Tornoceratidae. The type species is L. materni. |
| Linguatornoceras acutilobatum | Sp. nov | Valid | Söte & Becker | Devonian (Frasnian) | Büdesheim Formation | Germany | A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023. |
| Linguatornoceras sandense | Sp. nov | Valid | Söte & Becker | Devonian (Frasnian) | Büdesheim Formation | Germany | A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023. |
| Lissoceras kubanense | Sp. nov | Valid | Mitta | Middle Jurassic (Bajocian) | | Russia ( Karachay-Cherkessia) | |
| Lissoceras paviai | Sp. nov | Valid | Mitta | Middle Jurassic (Bajocian) | | Russia ( Karachay-Cherkessia) | |
| Megatyloceras leteilensis | Sp. nov | Valid | Delanoy et al. | Early Cretaceous (Aptian) | | France | A member of the family Douvilleiceratidae. |
| Nejdia arabica | Sp. nov | Valid | Gelin | Early Jurassic (Toarcian) | Marrat Formation | Saudi Arabia | |
| Paracrochordiceras watanabei | Sp. nov | Valid | Ehiro | Middle Triassic (Anisian) | Fukkoshi Formation | Japan | A member of the family Acrochordiceratidae. |
| Paradanubites ozashiense | Sp. nov | Valid | Ehiro | Middle Triassic (Anisian) | Fukkoshi Formation | Japan | A member of Danubitoidea belonging to the family Danubitidae. |
| Parahedenstroemia? tatjanae | Sp. nov | Valid | Đaković, Krystyn & Sudar | Early Triassic | | Montenegro  Russia? | A member of the family Hedenstroemiidae. |
| Parapopanoceras involutum | Sp. nov | Valid | Ehiro | Middle Triassic (Anisian) | Fukkoshi Formation | Japan | A member of Ceratitida belonging to the group Megaphyllitoidea and the family Parapopanoceratidae. |
| Paucitornoceras | Gen. et comb. nov | Valid | Korn & Bockwinkel | Devonian | Red Ironstone Formation | Germany  United States ( New York) | A member of the family Tornoceratidae. The type species is "Goniatites" paucistriatus d`Archiac & de Verneuil (1842); genus also includes "Tornoceras" rhysum Clarke (1898). |
| Ponticeras materni | Sp. nov | Valid | Korn & Bockwinkel | Devonian (Givetian) | Red Ironstone Formation | Germany | A member of Agoniatitida belonging to the group Gephuroceratina and the family Ponticeratidae. |
| Procheloniceras yajimai | Sp. nov | In press | Matsukawa | Early Cretaceous | | Japan | |
| Pseudocrioceras mazierei | Sp. nov | Valid | Frau & Delanoy | Early Cretaceous (Barremian/Aptian transition) | | France | |
| Pseudosaynella pacifica | Sp. nov | In press | Matsukawa | Early Cretaceous | | Japan | |
| Psilokhvalynites | Gen. et sp. nov | Valid | Ehiro | Middle Triassic (Anisian) | Fukkoshi Formation | Japan | A member of Dinaritoidea belonging to the family Khvalynitidae. The type species is P. takaizumii. |
| Shasticrioceras nakazawai | Sp. nov | In press | Matsukawa | Early Cretaceous | | Japan | |
| Sigaloceras (Gulielmina) fabricius | Sp. nov | Valid | Mönnig & Dietl | Middle Jurassic (Callovian) | | Germany | |
| Tornoceras incisum | Sp. nov | Valid | Söte & Becker | Devonian (Frasnian) | Büdesheim Formation | Germany | A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023. |
| Tornoceras ventrovaricatum | Sp. nov | Valid | Söte & Becker | Devonian (Frasnian) | Büdesheim Formation | Germany | A member of the family Tornoceratidae. Announced in 2022; the final article version was published in 2023. |
| Witchellia companyi | Sp. nov | Valid | Sandoval | Middle Jurassic (Bajocian) | | Spain | A member of the family Sonniniidae. |
| Witchellia striata | Sp. nov | Valid | Sandoval | Middle Jurassic (Bajocian) | | Spain | A member of the family Sonniniidae. |

### Ammonite research

* Mironenko & Naugolnykh (2022) describe a collection of cephalopod jaws from the Permian (Artinskian) Divjinskian Formation (Sverdlovsk Oblast, Russia), interpreted as likely fossil material of goniatitid ammonites belonging to the genus Uraloceras, and infer from the anatomy of the studied jaws that representatives of the genus Uraloceras were likely active predators.
* A study on the morphological change through ontogeny of beyrichitine ammonites from the Anisian of Nevada is published by Bischof, Schlüter & Lehmann (2022).
* A study aiming to determine whether the environmental changes around the Triassic–Jurassic boundary influenced development of septal thickness in ammonites, as well aiming to determine the relation between septal thickness and paleolatitudinal occurrences, and whether ontogenetic septal thickness-trajectories reflected paleogeographic origin or phylogenetic relationships of ammonites, is published by Weber et al. (2022).
* A study on the hydrodynamic properties of shells of Late Triassic and Early Jurassic ammonites is published by Hebdon et al. (2022).
* A study on the functional morphology of ammonites from the Middle Triassic to the Early Jurassic, reevaluating possible causes of an overall shift of ammonite shell forms during the studied periods, is published by Hebdon et al. (2022), who interpret their findings as indicating that the trade-offs between coasting efficiency, volume accretion per unit surface area accreted and diameter accretion per unit of surface area (three factors proposed as explanations of the shift of shell forms) were identical in the Triassic and in the Jurassic.
* Jaws of ammonites belonging to the Boreal family Polyptychitidae, with lower jaws closely resembling jaws of craspeditids, cardioceratids and hoplitoids, are described for the first time from the Valanginian of Russia by Mironenko & Mitta (2022).
* A study on changes of septal spacing between succeeding chambers of the shell of Gaudryceras tenuiliratum during its ontogeny is published by Kawakami, Uchiyama & Wani (2022).
* A study on the impact of the conch shape on ammonite locomotion, based on data obtained from tests of 3D-printed, biomimetic robots representing the oxycone morphology (based on Sphenodiscus lobatus and S. lenticularis), the serpenticone morphology (based on Dactylioceras commune), the sphaerocone morphology (based on Goniatites crenistria) and the morphospace center, is published by Peterman & Ritterbush (2022).

## Other cephalopods
### New taxa

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Aphelaeceras azzelmattiense | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian–Viséan boundary interval) | | Algeria | A member of the family Trigonoceratidae. |
| Arctoteuthis angusta | Sp. nov | Valid | Efremenko | Early Cretaceous | | Russia | A belemnite belonging to the family Cylindroteuthididae. |
| Binoleniceras | Gen. et sp. nov | Valid | Afhüppe & Becker | Devonian (Givetian) | Hagen-Balve Formation | Germany | A member of Discosorida belonging to the family Entimoceratidae. The type species is B. stichlingi. |
| Boreioteuthis mirifica | Sp. nov | Valid | Efremenko | Early Cretaceous | | Russia | A belemnite belonging to the family Cylindroteuthididae. |
| Chisiloceras maocaopuense | Sp. nov | Valid | Fang in Fang et al. | Ordovician (Darriwilian) | | China | A member of the family Endoceratidae. |
| Domatoceras tuckeri | Sp. nov | Valid | Niko, Mapes & Seuss | Carboniferous (late Pennsylvanian) | Graham Formation | United States ( Texas) | A member of the family Grypoceratidae. |
| Endolobus sturgeoni | Sp. nov | Valid | Niko, Mapes & Seuss | Carboniferous (late Pennsylvanian) | Graham Formation | United States ( Texas) | A member of the family Koninckioceratidae. |
| Enoploceras rieberi | Sp. nov | Valid | Pieroni | Middle Triassic (Anisian) | Besano Formation | Switzerland | |
| Eximioceras | Gen. et sp. nov | Valid | Shchedukhin | Early Permian | | Russia ( Bashkortostan) | A member of the family Rhiphaeoceratidae. The type species is E. venustum. |
| Holmiceras havelense | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Red Orthoceratite Limestone | Germany | |
| Kotoceras minosiphonatum | Sp. nov | Valid | Fang in Fang et al. | Ordovician (Darriwilian) | | China | A member of the family Endoceratidae. |
| Lispoceras orbis | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Lituites baculus | Sp. nov | Valid | Aubrechtová & Korn | Middle Ordovician | | Sweden | |
| Lituites bottkei | Sp. nov | Valid | Aubrechtová & Korn | Middle Ordovician | | Poland  Sweden | |
| Lituites clavis | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Grey Orthoceratite Limestone | Germany | |
| Lituites dewitzi | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Grey Orthoceratite Limestone | Germany | |
| Lituites kruegeri | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Grey Orthoceratite Limestone | Germany | |
| Lituites nebeni | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Grey Orthoceratite Limestone | Germany | |
| Maccoyoceras concavum | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian–Viséan boundary interval) | | Algeria | A member of the family Trigonoceratidae. |
| Maccoyoceras habadraense | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Maccoyoceras pentagonum | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Maccoyoceras saharensis | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Metacoceras quadratum | Sp. nov | Valid | Niko, Mapes & Seuss | Carboniferous (late Pennsylvanian) | Graham Formation | United States ( Texas) | A member of the family Tainoceratidae. |
| Michelinoceras minutum | Sp. nov | Valid | Fang in Fang et al. | Ordovician (Darriwilian and Sandbian) | | China | |
| Michelinoceras multiseptum | Sp. nov | Valid | Fang in Fang et al. | Ordovician (Darriwilian) | | China | |
| Pararhiphaeoceras parvum | Sp. nov | Valid | Shchedukhin | Early Permian | | Russia ( Bashkortostan) | A member of the family Rhiphaeoceratidae. |
| Planetoceras destrictum | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Planetoceras transforme | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Pleuronautilus chisatoi | Sp. nov | Valid | Ehiro | Permian (Wordian) | Takakurayama Formation | Japan | A member of the family Tainoceratidae. |
| Proterovaginoceras rapidum | Sp. nov | Valid | Fang in Fang et al. | Ordovician (Darriwilian) | | China | A member of the family Endoceratidae. |
| Rhynchorthoceras kranepuhlense | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Red Orthoceratite Limestone | Germany  Poland | |
| Rhynchorthoceras rugium | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Red Orthoceratite Limestone | Germany | |
| Rineceras multituberculatum | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Rineceras rectangulatum | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Rineceras tenerum | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Solenochilus jackense | Sp. nov | Valid | Niko, Mapes & Seuss | Carboniferous (late Pennsylvanian) | Graham Formation | United States ( Texas) | A member of the family Solenochilidae. |
| Stroboceras ancilis | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Viséan) | | Algeria | A member of the family Trigonoceratidae. |
| Stroboceras mane | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Stroborineceras | Gen. et 2 sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. The type species is S. insalahensis; genus also includes new species S. felis and possibly also "Nautilus (Trematodiscus)" meekianus Winchell (1862) and "Nautilus (Trematodiscus)" strigatus Winchell (1862). |
| Syllipsimopodi | Gen. et sp. nov | Disputed | Whalen & Landman | Carboniferous (Serpukhovian) | Bear Gulch Limestone | United States ( Montana) | A member of Coleoidea of uncertain affinities. Originally assigned to the clade Vampyropoda; Klug et al. (2023) considered it to be a probable junior synonym of Gordoniconus beargulchensis. The type species is S. bideni, named for Joe Biden. |
| Thrincoceras devolvere | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Tohokubelus | Gen. et sp. nov | Valid | Niko & Ehiro | Early Triassic (Olenekian) | Osawa Formation | Japan | A sinobelemnitid belemnite. Genus includes new species T. takaizumii. |
| Trilacinoceras filix | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Grey Orthoceratite Limestone | Germany | A member of the family Lituitidae. |
| Trilacinoceras knoefleri | Sp. nov | Valid | Aubrechtová & Korn | Ordovician (Darriwilian) | Upper Grey Orthoceratite Limestone | Germany | A member of the family Lituitidae. |
| Trilobitoceras | Gen. et sp. et comb. nov | Valid | Korn & Bockwinkel | Carboniferous | | Algeria  United States ( Michigan) | A member of the family Trigonoceratidae. The type species is T. peculiaris; genus also includes "Nautilus (Trematodiscus)" planidorsalis Winchell (1862). |
| Vestinautilus angulatus | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Vestinautilus bicristatus | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Vestinautilus concinnus | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Vestinautilus inflexus | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Vestinautilus padus | Sp. nov | Valid | Korn, Miao & Bockwinkel | Early Carboniferous | | Algeria | A member of the family Trigonoceratidae. |
| Vestinautilus papilio | Sp. nov | Valid | Korn & Bockwinkel | Carboniferous (Tournaisian) | | Algeria | A member of the family Trigonoceratidae. |
| Xainzanoceras densum | Sp. nov | Valid | Fang in Fang et al. | Ordovician (Katian) | Pagoda Formation | China | Possibly a member of the family Diestoceratidae. |

### Other cephalopod research

* A study on the phylogenetic relationships of Cambrian and Ordovician cephalopods is published by Pohle et al. (2022).
* Putative tentaculitoid Iwakiella ichiroi is reinterpreted as a sphaerorthoceratid orthocerid cephalopod by Niko (2022).
* A study on the identity of enigmatic black amorphous fossils often associated with belemnoid remains, based on the analysis of fossil material from the Upper Triassic (Carnian) Polzberg Lagerstätte (Austria), is published by Lukeneder & Lukeneder (2022), who interpret the enigmatic fossils as likely representing the preservation of a mineralized and secondarily carbonized cephalic-ocular-arm-cartilage complex of the belemnoid Phragmoteuthis bisinuata.
* A study on structures interpreted by Eduard Suess as beaks of Phragmoteuthis bisinuata is published by Doguzhaeva et al. (2022), who reinterpret the studied structures as cartilaginous remains of a prey, presumably juvenile fish; the authors also report a gladius of a previously unknown Carnian teuthid from the Cave del Predil (Italy), possibly similar to the Permian taxon Glochinomorpha stifeli, as well as an upper beak of a coleoid from the Anisian Buchenstein Formation (Italy), demonstrating typical coleoid upper beak morphology and composition, and interpreted by the authors as likely indicative of similar upper beak structure in P. bisinuata. Their interpretation of purported beaks of P. bisinuata as remains of a prey is subsequently contested by Lukeneder & Lukeneder (2022).
* Belemnite species Liobelus acrei, otherwise known from the latest Jurassic and Early Cretaceous of the Boreal Realm, is reported from the Valanginian of the Vocontian Basin (France) by Mutterlose, Picollier & Dzyuba (2022), who interpret this finding as evidence of an isolated immigration of belemnites from the north in the early Valanginian, and establish two taxonomically different faunas of Tethyan belemnites.
* A study on the age of belemnites from the lower Santonian "Sponge horizon" of the Mozhzhelloovrazhnaya Formation in the Lower Volga region between Saratov and Volgograd (Russia), some of which are interpreted as redeposited from middle and upper Coniacian deposits on the basis of strontium, carbon and oxygen isotope data (with the first representatives of the genus Belemnitella interpreted as late Coniacian in age), is published by Zakharov et al. (2022).
* Revision of the type specimens of Eothinoceras americanum is published by Evans & Cichowolski (2022), who restrict the genus Eothinoceras to the type species, and propose a new scheme for classification of the genera belonging to the order Cyrtocerinida.
* A study on the distribution of nautiloids throughout the Cenozoic is published by Kiel, Goedert & Tsai (2022), who interpret their findings as indicating that from the Oligocene onward nautiloids became extinct in areas where pinnipeds appeared, while cetaceans (with possible exception of simocetids and agorophiids) were not found to significantly affect the demise of nautiloids.
* Fossil material of "Kummelonautilus" taiwanum is described from the Miocene Houdongkeng Formation (Taiwan) by Goedert, Kiel & Tsai (2022), who provisionally transfer this species to the genus Nautilus, and review the fossil record of the genus Nautilus in the Indo-Pacific region.
* A study on the external and internal morphology, affinities and ecology of Vampyronassa rhodanica, based on data on soft tissue of specimens from the La Voulte-sur-Rhône (France), is published by Rowe et al. (2022).
* First cephalopod statoliths from the Lower Cretaceous of Poland and United Kingdom are described by Pindakiewicz et al. (2022), who compare Mesozoic statoliths to those of extant cephalopods, and find the closest resemblance between Mesozoic forms and statoliths of extant idiosepiids.
* A study on the fossil material of Belosaepia from the Egemkapel Clay Member of the Tielt Formation and the Roubaix Clay Member of the Kortrijk Formation (Belgium), applying micro-CT imaging to the studied fossils, is published by Goolaerts et al. (2022), who identify growth lines in the studied fossils, and interpret their findings as indicating that the studied material belongs to representatives of a single species (B. tricarinata).

## Bivalves
### New taxa

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Acila (Truncacila) interferencia | Sp. nov | Valid | Kiel et al. | Miocene | Amlang Formation | Philippines | A species of Acila. |
| Anniedhondtella | Gen. et sp. nov | Valid | Jagt, Cooper & Jagt-Yazykova | Late Cretaceous (Maastrichtian) | Maastricht Formation | | A member of Trigoniida belonging to the family Rutitrigoniidae. Genus includes new species A. rieui. |
| Aphanaia sobopolensis | Sp. nov | Valid | Biakov & Kutygin | Early Permian | | Russia ( Sakha) | |
| Arcopagia esteponensis | Sp. nov | Valid | Brunetti & Della Bella | Pliocene | | Spain | A member of the family Tellinidae. |
| Arcopagia ligustica | Sp. nov | Valid | Brunetti & Della Bella | Early Pliocene | | Italy | A member of the family Tellinidae. |
| Arcopella crestii | Sp. nov | Valid | Brunetti & Della Bella | Late Pliocene | | Italy | A member of the family Tellinidae. |
| Arcopella pseudomista | Sp. nov | Valid | Brunetti & Della Bella | Early Pliocene | | Italy | A member of the family Tellinidae. |
| Bathymodiolus labayugensis | Sp. nov | Valid | Kiel et al. | Miocene | Amlang Formation | Philippines | A species of Bathymodiolus. |
| Choniocardia olivieri | Nom. nov | Valid | Pacaud | Eocene | | France | A member of the family Carditidae; a replacement name for Cardita (Miodon) bezanconi Cossmann (1887). |
| Conchocele pangasinanensis | Sp. nov | Valid | Kiel et al. | Miocene | Amlang Formation | Philippines | A member of the family Thyasiridae. |
| Crassatella (Bathytormus) ignava | Sp. nov | Valid | Berezovsky | Eocene | Malinovskaya Formation | Ukraine | A member of the family Crassatellidae. |
| Crassatella lassa | Sp. nov | Valid | Berezovsky | Late Eocene | | Ukraine | |
| Crassostrea sophiae | Nom. nov | Valid | Pacaud | Eocene | | France | A species of Crassostrea; a replacement name for Ostrea elegans Deshayes (1832). |
| Cyclotellina hugoi | Nom. nov | Valid | Pacaud | Eocene | | France | A member of the family Tellinidae; a replacement name for Tellina (Cyclotellina) bezanconi Cossmann (1906). |
| Gryphaea (Gryphaea) lavernockensis | Sp. nov | Valid | Hodges | Early Jurassic | | United Kingdom | |
| Kasimlara ? antiqua | Sp. nov | Valid | Fang & Carter | Permian | | Russia | A member of the family Kalenteridae belonging to the subfamily Healeyinae. |
| Liaoningoconcha | Nom. nov | | Fang & Carter | Late Triassic | Yangcaogou Formation | China | A member of Cardiida belonging to the family Ferganoconchidae; a replacement name for Liaoningia Yu & Dong (1993). Ceccolini & Cianferoni (2021) proposed a replacement name Sinoliaoningia for the same genus in an earlier publication. |
| Megaxinus gorrospei | Sp. nov | Valid | Kiel et al. | Miocene | Amlang Formation | Philippines | A member of the family Lucinidae. |
| Meleagrinella golberti | Sp. nov | Valid | Lutikov & Arp | Early Jurassic (Toarcian) | | Canada  Germany  Russia  United Kingdom | A member of the family Oxytomidae. Published online in 2023, but the issue date is listed as December 2022. |
| Meleagrinella prima | Sp. nov | Valid | Lutikov in Lutikov & Arp | Early Jurassic (Toarcian) | | Russia | A member of the family Oxytomidae. Published online in 2023, but the issue date is listed as December 2022. |
| Monginaia | Gen. et comb. nov | In press | Delvene et al. | Early Cretaceous (Barremian) | Camarillas Formation | Spain | A member of Unionida belonging to the new family Monginaiidae. Genus includes "Elliptio" galvensis Mongin (1966). |
| Myonia komiensiformis | Sp. nov | Valid | Biakov | Permian | | Russia | A member of Pholadomyida belonging to the family Sanguinolitidae. |
| Myonia kutygini | Sp. nov | Valid | Biakov | Permian | | Russia | A member of Pholadomyida belonging to the family Sanguinolitidae. |
| Oudardia pliorecens | Sp. nov | Valid | Brunetti & Della Bella | Plio-Pleistocene | | Italy | A member of the family Tellinidae. |
| Palaeomutela (Palaeanodonta) berrutii | Sp. nov | Valid | Silantiev et al. | Permian (Kungurian) | Collio Formation | Italy | A member of Actinodontida belonging to the family Palaeomutelidae. |
| Palaeomutela (Palaeanodonta) guncinaensis | Sp. nov | Valid | Silantiev et al. | Permian (Kungurian) | Guncina Formation | Italy | A member of Actinodontida belonging to the family Palaeomutelidae. |
| Peronidia iberica | Sp. nov | Valid | Brunetti & Della Bella | Early Pliocene | | Spain | A member of the family Tellinidae. |
| Peronidia intermedia | Sp. nov | Valid | Brunetti & Della Bella | Early Pliocene | | Italy | A member of the family Tellinidae. |
| Pliocardia ballesterosi | Sp. nov | Valid | Kiel et al. | Miocene | Amlang Formation | Philippines | A species of Pliocardia. |
| Praebarrettia magnifica | Sp. nov | | Mitchell | Late Cretaceous (Campanian) | Dry Hill Formation | Jamaica | A rudist bivalve. |
| Psammotreta santacatalinae | Sp. nov | Valid | Brunetti & Della Bella | Early Pliocene | | Spain | A member of the family Tellinidae. |
| Sisonia | Gen. et sp. nov | Valid | Kiel et al. | Miocene | Amlang Formation | Philippines | A bivalve of uncertain affinities. The type species is S. frijellanae. |
| Squiresica | Gen. et comb. nov | Valid | Hybertsen, Goedert & Kiel | Oligocene | Lincoln Creek Formation | United States ( Alaska Washington) | A member of the family Vesicomyidae. The type species is "Archivesica" knapptonensis Amano & Kiel (2007); genus also includes "Archivesica" marincovichi Kiel & Amano (2010). |
| Verneuilnodon | Gen. et comb. nov | Valid | Fang & Carter | Permian | | Russia | A member of the family Kalenteridae belonging to the subfamily Healeyinae; a new genus for "Mytilus (Modiola)" pallasi de Verneuil (1845). |
| Whitfieldiella catherinei | Sp. nov | | Mitchell | Late Cretaceous (Campanian) | Dry Hill Formation | Jamaica | A rudist bivalve. |
| Whitfieldiella gracilis | Sp. nov | | Mitchell | Late Cretaceous (Campanian) | Dry Hill Formation | Jamaica | A rudist bivalve. |
| Wolfgangella | Gen. et 2 sp. nov | Valid | Jagt, Cooper & Jagt-Yazykova | Late Cretaceous (Maastrichtian) | Maastricht Formation | | A member of Trigoniida belonging to the family Rutitrigoniidae. Genus includes new species W. neilpearti and W. ignota. |

### Bivalve research

* Late Triassic bivalves are identified for the first time in olistostromes from the Sêwa Formation in the Riganpeicuo area (Tibet, China) by Xiao et al. (2022).
* A study on the impact of the Triassic–Jurassic extinction event on the body-size distribution of bivalves, based on data from three study sites in the United Kingdom, is published by Opazo & Twitchett (2022).
* Wolkenstein (2022) presents evidence of abundance and diversity in the color patterns of Pleuronectites laevigatus from the Middle Triassic Muschelkalk of Central Europe.
* Asato, Nakayama & Imai (2022) study color patterns of freshwater bivalves from the Lower Cretaceous Kitadani Formation (Japan), providing evidence of the presence of similar shell color patterns in extant and Early Cretaceous bivalves.

## Gastropods
### New taxa

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Akburunella laminaris | Sp. nov | Valid | Guzhov | Miocene | | Russia | Published online in 2023, but the issue date is listed as December 2022. |
| Akburunella nefanda archaica | Ssp. nov | Valid | Guzhov | Miocene | | Russia | Published online in 2023, but the issue date is listed as December 2022. |
| Akburunella sinuosa | Sp. nov | Valid | Guzhov | Miocene | | Russia | Published online in 2023, but the issue date is listed as December 2022. |
| Akburunella spinosa | Sp. nov | Valid | Guzhov | Miocene | | Russia | Published online in 2023, but the issue date is listed as December 2022. |
| Ampezzopleura barremica | Sp. nov | Valid | Isaji, Haga & Kashiwagi | Early Cretaceous (Barremian) | Kimigahama Formation | Japan | |
| Angularia corallina | Sp. nov | Valid | Nützel et al. | Late Triassic (Rhaetian) | Oberrhät Formation | Austria | A member of Caenogastropoda belonging to the family Purpuroideidae. |
| Angulomphalus senowbarii | Sp. nov | Valid | Nützel et al. | Late Triassic (Rhaetian) | Oberrhät Formation | Austria | A member of Vetigastropoda belonging to the family Ptychomphalidae. |
| Anisocycla subcylindrica | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A member of the family Murchisonellidae. |
| Antiphora aurora | Sp. nov | Valid | Isaji, Haga & Kashiwagi | Early Cretaceous (Barremian) | Kimigahama Formation | Japan | |
| Aplexa canculae | Sp. nov | Valid | Pacaud | Paleocene (Thanetian) | | France | A species of Aplexa. |
| Aplexa cotiensis | Nom. nov | Valid | Pacaud | Eocene | | France | A species of Aplexa; a replacement name for Physa (Aplecta) brevispirata Cossmann & Pissarro (1913). |
| Aplexa iota | Sp. nov | Valid | Pacaud | Paleogene | | France | A species of Aplexa. |
| Aplexa oryza | Sp. nov | Valid | Pacaud | Paleogene | | France | A species of Aplexa. |
| Aplexa risleius | Sp. nov | Valid | Pacaud | Paleocene (Thanetian) | | France | A species of Aplexa. |
| Aplexa zhui | Nom. nov | Valid | Pacaud | Early Cretaceous (Albian) | Dalazi Formation | China | A species of Aplexa; a replacement name for Physa fusiformis Zhu (1980). |
| Arachnotropis | Gen. et comb. nov | Valid | Pérez, Ferrari & Ezcurra | Paleocene–Eocene | | Antarctica  Australia | A member of the family Eucyclidae. The type species is "Calliotropis" microglyptophorus Darragh (1997); genus also includes "Calliotropis" antarchais Stilwell (2005). |
| Archaeocyclotus brevivillosus | Sp. nov | In press | Bichain et al. | Cretaceous | Burmese amber | Myanmar | A member of the family Cyclophoridae. |
| Archaeostylus | Gen. et sp. nov | Valid | Brook & Hayward | Pliocene (Waipipian) | Kaawa Formation | New Zealand | A member of the family Bothriembryontidae belonging to the subfamily Placostylinae. The type species is A. manukauensis. |
| Astralium ena | Sp. nov | Valid | Tomida, Inoue & Kase | Miocene | Ena Limestone | Japan | A species of Astralium. |
| Austrocypraea jimgracei | Sp. nov | Valid | Southgate & Roberts | Pliocene | | Australia | A member of the family Cypraeidae. |
| Barycypraea iungo | Sp. nov | Valid | Seccombe & Aiken | Pleistocene | | South Africa | A species of Barycypraea. |
| Bathrotomaria teodorii | Sp. nov | Valid | Chaix in Chaix & Plicot | Late Cretaceous (Santonian) | | France | A member of the family Pleurotomariidae. |
| Baylea tenera | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Boggy Formation | United States ( Oklahoma) | A member of the family Eotomariidae. |
| Brachystomia succineiformis | Sp. nov | Valid | Guzhov | Miocene | | Russia | A species of Brachystomia. Published online in 2023, but the issue date is listed as December 2022. |
| Ceritella convexa | Sp. nov | Valid | Gründel et al. | Late Jurassic (Kimmeridgian) | | Germany | A member of the family Ceritellidae. |
| Choshipleura | Gen. et sp. nov | Valid | Isaji, Haga & Kashiwagi | Early Cretaceous (Barremian) | Kimigahama Formation | Japan | Genus includes new species C. striata. |
| Clathromangelia acuticostata | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Clathromangelia. |
| Clathromangelia mulderi | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Clathromangelia. |
| Clathromangelia oliverioi | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Clathromangelia. |
| "Clavatula" ariejansseni | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Serravallian) | Týrtar Formation | Turkey | A member of the family Clavatulidae. |
| "Clavatula" atatuerki | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Serravallian) | Týrtar Formation | Turkey | A member of the family Clavatulidae. |
| "Clavatula" iberica | Sp. nov | Valid | Landau & Harzhauser | Pliocene (Piacenzian) | | Spain | A member of the family Clavatulidae. |
| "Clavatula" irisae | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Baden Formation | Austria  Slovenia | A member of the family Clavatulidae. |
| "Clavatula" jarzynkae | Nom. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Grund Formation | Austria  Poland | A member of the family Clavatulidae; a replacement name for Clavatula auingeri Finlay (1927). |
| "Clavatula" nana | Sp. nov | Valid | Landau & Harzhauser | Pliocene (Piacenzian) | | Spain | A member of the family Clavatulidae. |
| "Clavatula" pliogradata | Sp. nov | Valid | Landau & Harzhauser | Pliocene (Piacenzian) | | Spain | A member of the family Clavatulidae. |
| Clavatula sorini | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Dej Formation | Romania  Italy? | A species of Clavatula. |
| Clithon cooncreekense | Sp. nov | Valid | Garvie & Symonds | Late Cretaceous (Maastrichtian) | Coon Creek Formation | United States ( Tennessee) | A species of Clithon. |
| Clithon duomillias | Sp. nov | Valid | Garvie & Symonds | Eocene | Cook Mountain Formation | United States ( Texas) | A species of Clithon. |
| Clithon lisae | Sp. nov | Valid | Garvie & Symonds | Eocene | Cook Mountain Formation | United States ( Texas) | A species of Clithon. |
| Clithon traceyi | Sp. nov | Valid | Garvie & Symonds | Eocene | Cook Mountain Formation | United States ( Texas) | A species of Clithon. |
| Columbellaria rara | Sp. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian) | St-Ursanne Formation | Switzerland | A member of the family Colombellinidae. |
| Conotomaria lecatae | Sp. nov | Valid | Chaix & Grenier | Late Cretaceous (Campanian) | | France | A member of the family Pleurotomariidae. |
| Conus damianakisi | Sp. nov | Valid | Psarras, Merle & Koskeridou | Miocene (Tortonian) | | Greece | A species of Conus. |
| Conus davolii | Sp. nov | Valid | Psarras, Merle & Koskeridou | Miocene (Tortonian) | | Greece  Italy  Hungary? | A species of Conus. |
| Conus dellabellai | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A species of Conus. |
| Conus ictini | Sp. nov | Valid | Psarras, Merle & Koskeridou | Miocene (Serravallian and Tortonian) | | Greece  Turkey | A species of Conus. |
| Conus lauriatragei | Sp. nov | Valid | Psarras, Merle & Koskeridou | Miocene (Langhian to Tortonian) | | Greece  Hungary | A species of Conus. |
| Conus moissettei | Sp. nov | Valid | Psarras, Merle & Koskeridou | Miocene (Tortonian) | | Greece | A species of Conus. |
| Conus villalvernensis | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A species of Conus. |
| Coptocheilus kachinensis | Sp. nov | | Yu, Zhuo & Páll-Gergely | Cretaceous | Burmese amber | Myanmar | A species of Coptocheilus. |
| Cornirostra anistratenkorum umbilicaris | Ssp. nov | Valid | Guzhov | Miocene | | Russia | Published online in 2023, but the issue date is listed as December 2022. |
| Crepidula bellardii | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A species of Crepidula. |
| Cretadiostoma | Gen. et 2 sp. nov | | Yu, Zhuo & Páll-Gergely | Cretaceous | Burmese amber | Myanmar | A member of the family Pupinidae. Genus includes new species C. caperatum and C. umbilicarinatum. |
| Cryptoptyxis? fortiter | Sp. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian) | St-Ursanne Formation | Switzerland | A member of the family Maoraxidae. |
| Cyrillia clathrataeformis | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Cyrillia. |
| Cyrillia saldubensis | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Cyrillia. |
| Dentitriangularum | Gen. et sp. nov | Valid | Chaix in Chaix & Plicot | Late Cretaceous (Santonian) | | France | A member of the family Cypraeidae. The type species is D. pacaudi. |
| Diaugasma mediterranea | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Diaugasma. |
| Dictyotomaria turrisbabel | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Caddo Creek Formation | United States ( Texas) | A member of the family Phymatopleuridae. |
| Diempterus? multicostatus | Sp. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian) | St-Ursanne Formation | Switzerland | A member of the family Aporrhaidae. |
| Edrozeba | Gen. et 4 sp. et comb. nov | Valid | Guzhov | Miocene | | Russia | A member of the family Hydrobiidae. The type species is E. caeca; genus also includes new species E. angulata, E. minuta and E. striata, as well as "Hydrobia" enikalensis Kolesnikov (1935). Published online in 2023, but the issue date is listed as December 2022. |
| Eirlysella | Gen. et sp. et comb. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Boggy Formation | United States ( Oklahoma) | A member of the family Luciellidae. The type species is E. buckhornensis; genus also includes "Trochus" hissingerianus de Koninck (1843), "Pleurotomaria" squamula Phillips (1836) and "Luciellina" ocultabanda Kues & Batten (2001). |
| Endoplocus acutus | Sp. nov | Valid | Gründel et al. | Late Jurassic (Kimmeridgian) | | Germany | A member of the family Nerineidae. |
| Endoplocus inflatus | Sp. nov | Valid | Gründel et al. | Late Jurassic (Kimmeridgian) | | Germany | A member of the family Nerineidae. |
| Ericusa ngayawang | Sp. nov | Valid | Yates | Miocene (Langhian) | Cadell Formation | Australia | A species of Ericusa. |
| Exelissa ursannella | Sp. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian) | St-Ursanne Formation | Switzerland | A member of Cerithioidea belonging to the family Cryptaulacidae. |
| Faureicerithium | Gen. et sp. nov | Valid | Chaix in Chaix & Plicot | Late Cretaceous (Santonian) | | France | A member of the family Campanilidae. The type species is F. bidentata. |
| Favartia plioelata | Sp. nov | Valid | Brunetti & Della Bella | Pliocene | | Italy | A species of Favartia. |
| Granulatocincta | Gen. et comb. et 3 sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene and Pliocene | | Austria  Bulgaria  Czech Republic  France  Hungary  Italy  Romania  Slovakia  Spain  Turkey | A member of the family Clavatulidae. The type species is "Pleurotoma" granulato-cincta Münster in Goldfuss (1841); genus also includes new species G. callim, G. theoderichi and G. pelliscrocodili, as well as "Pleurotoma (Clavatula)" angelae Hoernes & Auinger (1891), "Clavatula" contorta Švagrovský (1958), "Pleurotoma (Clavatula)" nataliae Hoernes & Auinger (1891), "Pleurotoma" schreibersi Hörnes (1854), "Clavatula" calcarai Bellardi (1877), "Clavatula" curionii Bellardi (1877), "Clavatula" pellegrinii Simonelli (1896), "Pleurotoma" sotterii Michelotti (1847), "Clavatula" turbinata Bellardi (1877), "Clavatula" turgidula Bellardi (1877), "Clavatula" vigolensis Bellardi (1877), "Clavatula" rugata Bellardi (1877), "Murex" rusticus Brocchi (1814), "Pleurotoma" capgrandi Tournouër (1873), "Pleurotoma" obtruta Millet (1865), "Clavatula" delgadoi Vera-Peláez & Lozano-Francisco (2001) and possibly "Clavatula" boreointerrupta Kautsky (1925). |
| Hologyra callosa | Sp. nov | Valid | Nützel et al. | Late Triassic (Rhaetian) | Oberrhät Formation | Austria | A member of the family Neritopsidae. |
| Hydrobia neofrauenfeldi | Sp. nov | Valid | Guzhov | Miocene | | Russia | A species of Hydrobia. Published online in 2023, but the issue date is listed as December 2022. |
| Hydrobia orientalis | Sp. nov | Valid | Zhu | Late Permian | Wutonggou Formation | China | A species of Hydrobia. |
| Iberellus colladoi | Sp. nov | Valid | Juárez-Ruiz & Altaba | Pliocene | | Spain | A species of Iberellus. |
| Itieroptygmatis cylindrata | Sp. nov | Valid | Gründel et al. | Late Jurassic (Kimmeridgian) | | Germany | A member of Nerineoidea belonging to the family Itieriidae. |
| Jenneria sandrae | Sp. nov | Valid | Daughenbaugh | Pleistocene (Chibanian) | Probably Bermont Formation | United States ( Florida) | A species of Jenneria. |
| Leufroyia gradata | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Leufroyia. |
| Loriolotrema? nodosa | Sp. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian) | St-Ursanne Formation | Switzerland | A member of the family Brachytrematidae. |
| Lyria estotiensis | Sp. nov | Valid | Bail, Ledon & Pacaud | Oligocene (Chattian) | | France | A species of Lyria. |
| Lyria lozoueti | Sp. nov | Valid | Bail, Ledon & Pacaud | Miocene (Aquitanian) | | France | A species of Lyria. |
| Maoristylus pliocenicus | Sp. nov | Valid | Brook & Hayward | Pliocene (Waipipian) | Kaawa Formation | New Zealand | A member of the family Bothriembryontidae belonging to the subfamily Placostylinae. |
| Megaclavatula | Gen. et 2 sp. et comb. nov | Valid | Harzhauser, Landau & Janssen | Miocene | | Austria  Czech Republic  Hungary  Poland  Romania  Slovakia  Slovenia  Ukraine  Italy? | A member of the family Clavatulidae. The type species is M. grunerti; genus also includes new species M. pilleri, as well as "Pleurotoma (Clavatula)" amaliae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" antoniae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" evae Hoernes & Auinger (1891), "Pleurotoma" laevigatum Eichwald (1830), "Perrona" nemethi Kovács & Vicián (2021), "Pleurotoma" neudorfensis Schaffer (1898), "Pleurotoma tuberculosa" polonica Pusch (1837), "Pleurotoma (Clavatula) asperulata" var. subsculpta Schaffer (1912), "Pleurotoma" ernesti Toula (1901), "Pleurotoma" asperulata Lamarck (1822), Clavatula saubrigiana laurensii Peyrot (1931) and possibly "Pleurotoma (Clavatula)" szontaghi Strausz (1926), "Clavatula" taurofusulata Sacco (1890), "Clavatula" subdepressa Sacco (1890) and "Clavatula" tortonica Peyrot (1931). |
| Metacerithium boshuae | Sp. nov | Valid | Isaji, Haga & Kashiwagi | Early Cretaceous (Barremian) | Kimigahama Formation | Japan | A member of the family Metacerithiidae. |
| Metalycaeus fossilis | Sp. nov | In press | Yu, Páll-Gergely & Salvador | Miocene | Zhangpu amber | China | A species of Metalycaeus. |
| Microandonia | Gen. et sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A member of the family Raphitomidae. The type species is M. minutissima. |
| Microdrillia plioserratula | Sp. nov | Valid | Landau & Harzhauser | Pliocene | | Italy  Spain | A species of Microdrillia. |
| Microschiza rhaetica | Sp. nov | Valid | Nützel et al. | Late Triassic (Rhaetian) | Oberrhät Formation | Austria | A member of the family Purpurinidae. |
| Mitromorpha (Mitrolumna) velerinensis | Sp. nov | Valid | Landau & Harzhauser | Pliocene | | Spain  Italy? | A species of Mitromorpha. |
| Muricopsis landaui | Sp. nov | Valid | Merle in Merle, Garrigues & Pointier | Miocene | | Turkey | A species of Muricopsis. |
| Napaeus guanche | Sp. nov | In press | Miller et al. | Middle Pleistocene | | Spain ( Canary Islands) | A species of Napaeus. |
| Napaeus lipauges | Sp. nov | In press | Miller et al. | Middle Pleistocene | | Spain ( Canary Islands) | A species of Napaeus. |
| Narrimania saldubensis | Sp. nov | Valid | Landau & Mulder | Pliocene | | Spain | |
| Neoperrona | Gen. et comb. et sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | | Croatia  Hungary  Italy | A member of the family Clavatulidae. The type species is "Perrona" harzhauseri Kovács & Vicián (2021); genus also includes new species N. zoltanorum, as well as "Clavatula" taurinensis Bellardi (1877). |
| Neridomus mccallae | Sp. nov | Valid | Garvie & Symonds | Early Cretaceous (Albian) | Edwards Formation | United States ( Texas) | A member of the family Neridomidae. |
| Neridomus minutissimus | Sp. nov | Valid | Garvie & Symonds | Early Cretaceous (Albian) | Walnut Formation | United States ( Texas) | A member of the family Neridomidae. |
| Nerinea donosa | Sp. nov | Valid | Gründel et al. | Late Jurassic (Kimmeridgian) | | Germany | A member of the family Nerineidae. |
| Neuburgensia | Gen. et comb. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian to Tithonian) | | Germany  Switzerland | Possibly a member of Cerithioidea. The type species is "Gymnocerithium" concavum Janicke (1966); genus also includes "Cerithium" amabile Zittel (1873), "Gymnocerithium"? convexoconcavum Gründel, Keupp & Lang (2019), "Cerithium" crenato-cinctum Zittel (1873), "Cerithium" involvens Zittel (1873) and "Cerithium" perrotundum Cossmann (1913). |
| Nystia conoidalis | Sp. nov | Valid | Kadolsky & Morton | Oligocene (Rupelian) | | United Kingdom | A species of Nystia. |
| Odetta chirlii | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A species of Odetta. |
| Odostomia caucasica | Sp. nov | Valid | Guzhov | Miocene | | Russia | A species of Odostomia. Published online in 2023, but the issue date is listed as December 2022. |
| Oleginina | Gen. et sp. et comb. nov | Valid | Harzhauser, Landau & Janssen in Landau, Harzhauser & Giannuzzi-Savelli | Miocene (Burdigalian to Tortonian) | | Austria  Bosnia and Herzegovina  Bulgaria  Croatia  Czech Republic  Italy  Moldova  Poland  Romania  Slovakia  Ukraine | A member of the family Clavatulidae. The type species is O. mandici; genus also includes "Pleurotoma (Clavatula)" agathae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" brigittae Hoernes & Auinger (1891), "Pleurotoma" doderleini Hörnes (1854), "Pleurotoma (Clavatula)" dorotheae Hoernes & Auinger (1891), "Pleurotoma" rumana Simionescu & Barbu (1940), Clavatula schreibersi szokolyensis Strausz (1960), "Pleurotoma" winterlingensis Quenstedt (1884), "Clavatula" aradasi Bellardi (1877), "Clavatula" baccifera Bellardi (1877), "Clavatula" consularis Bellardi (1877) and "Clavatula" stazzanensis Bellardi (1877). The original generic name was Olegia, but this name turned out to be preoccupied by Olegia Shaposhnikov (1979), necessitating the creation of a replacement name. |
| Oligoptyxis khorasanica | Sp. nov | Valid | Guzhov & Nazemi in Raisossadat et al. | Early Cretaceous (Aptian) | | Iran | A member of the family Nerineidae. |
| Ondina curta | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A species of Ondina. |
| Ondina elongata | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A species of Ondina. |
| Ondina pseudovitrea | Sp. nov | Valid | Pavia et al. | Pliocene | | Italy | A species of Ondina. |
| Ophieulima lobilloensis | Sp. nov | Valid | Landau & Mulder | Pliocene | | Spain | A species of Ophieulima. |
| Otostoma cattoi | Sp. nov | Valid | Garvie & Symonds | Early Cretaceous (Albian) | Edwards Formation | United States ( Texas) | A member of the family Otostomidae. |
| Ovalotrema | Gen. et comb. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian) | | France  Switzerland | A member of the family Brachytrematidae. The type species is "Turbo" cotteausius d'Orbigny (1853); genus might also include "Turbo" greppini De Loriol in De Loriol & Koby (1895). |
| Palaeoleuca edeni | Sp. nov | Valid | Pacaud | Paleocene (Thanetian) to Eocene (Ypresian) | | France | A member of the family Ellobiidae. |
| Palaeoleuca gradata | Sp. nov | Valid | Pacaud | Paleocene (Thanetian) to Eocene (Ypresian) | | France | A member of the family Ellobiidae. |
| Palaeoleuca hulvizensis | Nom. nov | Valid | Pacaud | Paleocene | | United Kingdom | A member of the family Ellobiidae; a replacement name for Auricula (Conovulus) pygmaea Morris (1854). |
| Paleothyca | Gen. et sp. nov | Valid | Forner, Domènech & Martinell | Early Cretaceous (Albian) | | Spain | A member of the family Eulimidae. The type species is P. quinionarii. |
| Panneritina | Gen. et comb. et sp. nov | In press | Salvador & Yu | Jurassic and Cretaceous | | China  Myanmar | A member of the new family Neritidae. Genus includes Chinese species formerly assigned to the genus Mesoneritina ("M." crassa Pan, 1982, "M." dakangensis Pan, 1978, "M." gansuensis Yu & Guo, 1982, "M." nanshanensis Pan, 1984, "M." opima Yu, 1974, "M." pustula Pan, 1980), as well as new species P. ambrae from the Kachin amber. |
| Papuliscala spinosa | Sp. nov | Valid | Landau & Mulder | Pliocene | | Spain | A species of Papuliscala. |
| Paragoniozona ornata | Nom. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Lazy Bend Formation | United States ( Texas) | A member of the family Phymatopleuridae; a replacement name for Pleurotomaria aspera Girty (1934). |
| Paragoniozona yanceyi | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Boggy Formation | United States ( Oklahoma) | A member of the family Phymatopleuridae. |
| Perrona grossi | Nom. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Florian Formation | Austria | A species of Perrona; a replacement name for Pleurotoma (Clavatula) auingeri Hilber (1879). |
| Perrona ilonae | Nom. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Sámsonháza Formation | Hungary  Ukraine | A species of Perrona; a replacement name for Clavatula vindobonensis nodosa Csepreghy-Meznerics (1954). |
| Perrona koeberli | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Grund Formation | Austria | A species of Perrona. |
| Perrona loetschi | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Grund Formation | Austria | A species of Perrona. |
| Perrona wanzenboecki | Nom. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Baden Formation | Austria | A species of Perrona; a replacement name for Pleurotoma concinna Handmann (1883). |
| Peruvispira kanchanaburiensis | Sp. nov | In press | Karapunar, Nützel & Ketwetsuriya | Permian | Ratburi Group | Thailand | A member of the family Goniasmatidae. |
| Peruvispira oklahomaensis | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Gene Autry Formation | United States ( Oklahoma) | A member of the family Goniasmatidae. |
| Phymatopleura conica | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Gene Autry Formation | United States ( Oklahoma) | A member of the family Phymatopleuridae. |
| Phymatopleura girtyi | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Wewoka Formation | United States ( Oklahoma) | A member of the family Phymatopleuridae. |
| Platyphysa nagensis | Nom. nov | Valid | Pacaud | Cretaceous | | India | A replacement name for Physa prinsepii elongata Hislop (1860). |
| Pleurotomaria cloutetsensis | Sp. nov | Valid | Chaix in Chaix & Plicot | Late Cretaceous (Santonian) | | France | |
| Pleurotomella obesula | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Pleurotomella. |
| Pleurotomella turrita | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Pleurotomella. |
| Pomatiasia | Gen. et comb. nov | Valid | Guzhov | Miocene | | Russia | A member of the family Hydrobiidae. Genus includes "Amnicola" cyclostomoides Sinzov (1880). Published online in 2023, but the issue date is listed as December 2022. |
| Propefavartia | Gen. et comb. nov | Valid | Merle in Merle, Garrigues & Pointier | Eocene | | France | A member of the family Muricidae belonging to the subfamily Muricopsinae. The type species is "Murex" distans Deshayes (1835). |
| Pseudamnicola roblesi | Sp. nov | Valid | Albesa, López & Crespo | Early Miocene | Ribesalbes-Alcora Basin | Spain | A species of Pseudamnicola. |
| Pseudamnicola taodonggouensis | Sp. nov | Valid | Zhu | Late Permian | Wutonggou Formation | China | A species of Pseudamnicola. |
| Pseudodaphnella iberica | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Pseudodaphnella. |
| Pseudodaphnella velerinensis | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Pseudodaphnella. |
| Pseudodostia lisbonensis | Sp. nov | Valid | Garvie & Symonds | Eocene | Lisbon Formation | United States ( Alabama) | A member of the family Neritidae. |
| Pseudomelania yamadai | Sp. nov | Valid | Isaji, Haga & Kashiwagi | Early Cretaceous (Barremian) | Kimigahama Formation | Japan | |
| Pseudonerinea ? pseudomelaniformis | Sp. nov | Valid | Gründel et al. | Late Jurassic (Kimmeridgian) | | Germany | A member of Nerineoidea belonging to the family Pseudonerineidae. |
| Purpuripullina | Gen. et comb. nov | Valid | Guzhov & Raisossadat in Raisossadat et al. | Cretaceous | | Iran  Lebanon | The type species is "Natica" scalaris Conrad (1852). |
| Purpuroidea moosleitneri | Sp. nov | Valid | Nützel et al. | Late Triassic (Rhaetian) | Oberrhät Formation | Austria | A member of Caenogastropoda belonging to the family Purpuroideidae. |
| Pusionella hofmanni | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Dej Formation | Romania | A species of Pusionella. |
| Ringicula costata chokrakensis | Ssp. nov | Valid | Guzhov | Miocene | | Russia | Published online in 2023, but the issue date is listed as December 2022. |
| Ringicula costata paratethica | Ssp. nov | Valid | Guzhov | Miocene | | Russia | Published online in 2023, but the issue date is listed as December 2022. |
| Ringicula subglobosa | Sp. nov | Valid | Guzhov | Miocene | | Russia | A species of Ringicula. Published online in 2023, but the issue date is listed as December 2022. |
| Saalensia lorioli | Sp. nov | Valid | Gründel, Hostettler & Menkveld-Gfeller | Late Jurassic (Oxfordian) | St-Ursanne Formation | Switzerland | A member of the family Brachytrematidae. |
| Shansiella (Oklahomaella) globilineata | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Gene Autry Formation | United States ( Oklahoma) | A member of the family Portlockiellidae. |
| Sigmesalia anatole | Sp. nov | Valid | Hoşgör & Pacaud | Paleocene (Danian) | Çaldağ Formation | Turkey | A member of the family Turritellidae. |
| Sigmesalia arguta | Sp. nov | Valid | Berezovsky in Berezovsky & Demyanov | Eocene | Mandrikovka Beds | Ukraine | A member of the family Turritellidae. |
| Sigmesalia captiosa | Sp. nov | Valid | Berezovsky & Demyanov | Eocene | Mandrikovka Beds | Ukraine | A member of the family Turritellidae. |
| Sigmesalia erifensis | Sp. nov | Valid | Hoşgör & Pacaud | Paleocene (Danian) | Çaldağ Formation | Turkey | A member of the family Turritellidae. |
| Skenea gofasi | Sp. nov | Valid | Landau & Mulder | Pliocene | | Spain | A species of Skenea. |
| Specula estebbunensis | Sp. nov | Valid | Landau & Mulder | Pliocene | | Spain | A species of Specula. |
| Spinatus ruizi | Sp. nov | Valid | Liverani & Wieneke | Miocene (Langhian) | | Indonesia | A member of the family Strombidae. |
| Spiroscala quasipulchra | Nom. nov | Valid | Karapunar et al. | Permian | | United States | A member of the family Eotomariidae; a replacement name for Euconospira pulchra Batten (1958). |
| Spiroscala shwedagoniformis | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Gene Autry Formation | United States ( Oklahoma) | A member of the family Eotomariidae. |
| Striopusionella | Gen. et comb. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | | Hungary | A member of the family Clavatulidae. The type species is "Clavatula" szekelyhidiae Kovács & Vicián (2021). |
| Stuoraxis kasei | Sp. nov | Valid | Isaji, Haga & Kashiwagi | Early Cretaceous (Barremian) | Kimigahama Formation | Japan | A member of the family Stuoraxidae. |
| Stuorella zapfei | Sp. nov | Valid | Nützel et al. | Late Triassic (Rhaetian) | Oberrhät Formation | Austria | A member of Vetigastropoda belonging to the family Stuorellidae. |
| Subpterynotus eocaenicus | Sp. nov | | Merle, Lesport & Ledon | Eocene (Bartonian) | | France | A species of Subpterynotus. |
| Subpterynotus mainotensis | Sp. nov | | Merle, Lesport & Ledon | Miocene (Aquitanian) | | France | A species of Subpterynotus. |
| Talassia plioalboranensis | Sp. nov | Valid | Landau & Mulder | Pliocene | | Spain | A species of Talassia. |
| Teretia inflatissima | Sp. nov | Valid | Landau, Harzhauser & Giannuzzi-Savelli | Pliocene (Piacenzian) | | Spain | A species of Teretia. |
| Thalassonerita hagai | Sp. nov | Valid | Kiel et al. | Miocene | Amlang Formation | Philippines | A member of the family Phenacolepadidae. |
| Tomellana aueri | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Baden Formation | Austria  Bulgaria  Hungary  Romania  Slovakia  Slovenia | A species of Tomellana. |
| Tomellana dulaii | Sp. nov | Valid | Harzhauser, Landau & Janssen | Miocene (Langhian) | Dej Formation | Hungary  Romania | A species of Tomellana. |
| Tomellana onubensis | Sp. nov | Valid | Landau & Harzhauser | Early Pliocene | | Spain | A species of Tomellana. |
| Tomellana postjouannetii | Sp. nov | Valid | Landau & Harzhauser | Pliocene (Piacenzian) | | Spain | A species of Tomellana. |
| Tortaxis zhangpuensis | Sp. nov | In press | Yu, Páll-Gergely & Salvador | Miocene | Zhangpu amber | China | |
| Tosapusia peregrina | Sp. nov | Valid | Bertaccini et al. | Pliocene | | Italy | A species of Tosapusia. |
| Trigonostoma ariejansseni | Sp. nov | Valid | Roosen | Miocene | | Germany | A species of Trigonostoma. |
| Tudorella lenneae | Sp. nov | Valid | Groh et al. | Pliocene | | Spain ( Canary Islands) | A species of Tudorella. |
| Tudorella rothei | Sp. nov | Valid | Groh et al. | Pliocene | | Spain ( Canary Islands) | A species of Tudorella. |
| Turbonilla yadongensis | Sp. nov | | Li, Li & Garvie | Eocene (Ypresian) | Zhupure Formation | China | A species of Turbonilla. |
| Valvata complanusa | Sp. nov | Valid | Zhu | Late Permian | Wutonggou Formation | China | A species of Valvata. |
| Varicoturbo | Gen. et sp. nov | Valid | Nützel, Schweigert & Karapunar | Late Triassic (Carnian) | San Cassiano Formation | Italy | A member of the family Seguenziidae. Genus includes new species V. microstriatus. |
| Worthenia (Yochelsonospira) kuesi | Sp. nov | Valid | Karapunar et al. | Carboniferous (Pennsylvanian) | Wetumka Formation | United States ( Oklahoma) | A species of Worthenia. |
| Xinjiangospira habita | Sp. nov | Valid | Zhu | Late Permian | Wutonggou Formation | China | A member of the family Hydrobiidae. |

### Gastropod research

* A study on transitions from water to land throughout the evolutionary history of gastropods is published by Vermeij & Watson-Zink (2022).
* A study on changes in gastropod diversity at the genus/subgenus level from the Norian to the Pliensbachian is published by Ferrari & Hautmann (2022), who report that gastropods lost 56% of genera/subgenera during the Triassic–Jurassic extinction event, which was much more than the average loss of marine life at the time, and attempt to determine potential causes of the end-Triassic extinctions of gastropods.
* A study aiming to determine the drivers of diversification for European freshwater gastropods over the past 100 million years is published by Neubauer et al. (2022).
* A study aiming to determine the factors which caused the diversification of European freshwater gastropods in the Late Cretaceous is published by Neubauer & Harzhauser (2022).
* Revision of the fossil material of Ancistrolepis from the Eocene (Ypresian) Llajas Formation (California, United States), representing the oldest occurrence of this genus reported to date, is published by Squires (2022).

## Other molluscs
### New taxa

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Calloplax roederi | Sp. nov | Valid | Vendrasco, Powell & LaFollette | Miocene | | United States ( California) | A chiton. |
| Chiton solaris | Sp. nov | Valid | Vendrasco, Powell & LaFollette | Miocene | | United States ( California) | A chiton. |
| Dorispira avannga | Sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A helcionellid. |
| Dorispira? lemdadensis | Sp. nov | | Mghazli et al. | Cambrian | | Morocco | |
| Dorispira septentrionalis | Sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A helcionellid. |
| Dorispira tavsenensis | Sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A helcionellid. |
| Dorispira tippik | Sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A helcionellid. |
| Erugoconus | Gen. et sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A member of Helcionelloida of uncertain affinities. The type species is E. acuminatus. |
| Hoarepora | Nom. nov | Valid | Mapes in Mazaev | Carboniferous (Pennsylvanian) | | United States | A rostroconch; a replacement name for Oxyprora Hoare, Mapes & Yancey (2002). |
| Ischnochiton loureiroi | Sp. nov | Valid | Dell'Angelo et al. | Pliocene | Mondego Basin | Portugal | A chiton, a species of Ischnochiton. |
| Lepidochitona rochae | Sp. nov | Valid | Dell'Angelo et al. | Pliocene | Mondego Basin | Portugal | A chiton, a species of Lepidochitona. |
| Scenella? siku | Sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A member of Helcionelloida. Originally tentatively assigned to the genus Scenella, but subsequently made the type species of the separate genus Hensoniconus. |
| Sermeqiconus | Gen. et comb. nov | Valid | Peel & Kouchinsky | Cambrian | Aftenstjernesø Formation | Greenland | A helcionellid. The type species is "Figurina" polaris Peel (2021). |
| Tavseniconus | Gen. et sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A member of Helcionelloida of uncertain affinities. The type species is T. erectus. |
| Vendrascospira | Gen. et 2 sp. nov | Valid | Peel & Kouchinsky | Cambrian (Wuliuan) | Henson Gletscher Formation | Greenland | A helcionellid. The type species is V. troelseni; genus also includes V. frykmani. |
| Yochelcionella gracilituba | Sp. nov | | Mghazli et al. | Cambrian | | Morocco | |
| Yochelcionella longituba | Sp. nov | | Mghazli et al. | Cambrian | | Morocco | |

### Other mollusc research

* Revision of the oldest known specimens of Jinonicella kolebabai from the Middle Ordovician strata of Ukraine and Belarus is published by Gubanov et al. (2022), who report the preservation of growth lines which were previously documented only in Silurian specimens.
* New information on the anatomy of Typhloesus wellsi is presented by Conway Morris & Caron (2022), who consider it plausible that Typhloesus was a pelagic mollusc with possible affinities with gastropods.