# Gonocephalum

Gonocephalum is a genus in the darkling beetle family Tenebrionidae. There are more than 350 described species in Gonocephalum, found on every continent except Antarctica.

## Description

In the species of this genus the body is quite elongated, the base of pronotum is usually slightly narrower than the base of elytra. The hind wings are developed.

## Species

These 364 species belong to the genus Gonocephalum:
* Gonocephalum aalbui Ferrer, 2000
* Gonocephalum abnormale Kaszab, 1952
* Gonocephalum acanthocnemis Gridelli, 1945
* Gonocephalum acoriaceum Chûjô, 1975
* Gonocephalum acuticolle Kaszab, 1952
* Gonocephalum adelaide (Blackburn, 1894)
* Gonocephalum adpressiforme Kaszab, 1951
* Gonocephalum adpressum (Germar, 1824)
* Gonocephalum aequale (Erichson, 1843)
* Gonocephalum aequatoriale (Blanchard, 1853)
* Gonocephalum aethiopicum (Ardoin, 1965)
* Gonocephalum affine (Billberg, 1815)
* Gonocephalum alaticolle (Fairmaire, 1893)
* Gonocephalum albertii Ferrer, 1995
* Gonocephalum alternatum Carter, 1915
* Gonocephalum alternicostis Gebien, 1910
* Gonocephalum alticola Chatanay, 1914
* Gonocephalum amplicollis Fairmaire, 1894
* Gonocephalum amplithorax (Fairmaire, 1894)
* Gonocephalum andamanense Kaszab, 1952
* Gonocephalum andrewesi Kaszab, 1952
* Gonocephalum angelicae Ferrer, 1995
* Gonocephalum angolense Erichson, 1843
* Gonocephalum angustatum Chevrolat, 1873
* Gonocephalum angusticolle (Gerstaecker, 1855)
* Gonocephalum annamita Chatanay, 1917
* Gonocephalum apterum Koch, 1955
* Gonocephalum ardoinicum Kaszab, 1972
* Gonocephalum arenarium (Fabricius, 1775)
* Gonocephalum arisi Reitter, 1905
* Gonocephalum asperatum Gebien, 1920
* Gonocephalum assimile (Küster, 1849)
* Gonocephalum aterrimum (Montrouzier, 1860)
* Gonocephalum australe Boisduval, 1835
* Gonocephalum balmeae Kaszab, 1952
* Gonocephalum bartolozzii Ferrer, 1995
* Gonocephalum batesi Ferrer, 2000
* Gonocephalum belli Kaszab, 1952
* Gonocephalum bengalense Kaszab, 1952
* Gonocephalum besnardi Kaszab, 1982
* Gonocephalum bigranulatum Kaszab, 1952
* Gonocephalum bilineatum (Walker, 1858)
* Gonocephalum bimaculatum Ferrer, 1995
* Gonocephalum binotatum Ardoin, 1967
* Gonocephalum birmanicum Kaszab, 1952
* Gonocephalum biroi Kaszab, 1952
* Gonocephalum biseriatum Kaszab, 1975
* Gonocephalum blairi (Gebien, 1922)
* Gonocephalum borosi Kaszab, 1952
* Gonocephalum brachelytra Kaszab, 1952
* Gonocephalum bradymeroides Chatanay, 1917
* Gonocephalum bremeri Ferrer, 1955
* Gonocephalum brenieri Chatanay, 1917
* Gonocephalum brevicolle Philippi, 1860
* Gonocephalum brevicorne Chatanay, 1917
* Gonocephalum brevisetosum Kaszab, 1952
* Gonocephalum brincki Ferrer, 2000
* Gonocephalum brittoni Kaszab, 1952
* Gonocephalum buitenzorgense Kaszab, 1952
* Gonocephalum caesum Steven, 1829
* Gonocephalum calcaripes (Karsch, 1881)
* Gonocephalum calvulum (Olliff, 1889)
* Gonocephalum camerunense Gridelli, 1945
* Gonocephalum carpentariae (Blackburn, 1894)
* Gonocephalum catenulatum (Fairmaire, 1896)
* Gonocephalum celebense Kaszab, 1952
* Gonocephalum charbonnelae Ferrer, 2004
* Gonocephalum chinense Gebien, 1910
* Gonocephalum civicum Kaszab, 1952
* Gonocephalum clavigerum Kaszab, 1980
* Gonocephalum clypeatum Gebien, 1920
* Gonocephalum coarcticolle Kaszab, 1952
* Gonocephalum coenosum Kaszab, 1952
* Gonocephalum compressum Ferrer, 1995
* Gonocephalum condamini Ardoin, 1963
* Gonocephalum confusum Ferrer, 2000
* Gonocephalum consobrinum Blair, 1923
* Gonocephalum contractum Gerstaecker, 1871
* Gonocephalum controversum Gridelli, 1948
* Gonocephalum cookae Kaszab, 1952
* Gonocephalum corallinum Kaszab, 1961
* Gonocephalum coriaceum Motschulsky, 1858
* Gonocephalum costatum (Brullé, 1832)
* Gonocephalum cowardense Blackburn, 1894
* Gonocephalum crassepunctatum Kaszab, 1952
* Gonocephalum cristovallense (Montrouzier, 1855)
* Gonocephalum csikii Kaszab, 1952
* Gonocephalum curiosum Kaszab, 1952
* Gonocephalum curlettii Ferrer, 2000
* Gonocephalum curvicolle Reitter, 1889
* Gonocephalum dasiforme Kaszab, 1952
* Gonocephalum deliense Kaszab, 1952
* Gonocephalum demeteri Ferrer, 1995
* Gonocephalum dentipes Kaszab, 1952
* Gonocephalum dentitibia Gebien, 1920
* Gonocephalum depressum (Fabricius, 1801)
* Gonocephalum dermestoides (Gerstaecker, 1871)
* Gonocephalum deserticola Ferrer, 2000
* Gonocephalum devylderi Ferrer, 1995
* Gonocephalum dilatatum (Wollaston, 1854)
* Gonocephalum dorsogranosum (Fairmaire, 1896)
* Gonocephalum dravidum Bremer et Ferrer, 1992
* Gonocephalum dubium (Arrow, 1900)
* Gonocephalum duplegranulatum Gridelli, 1948
* Gonocephalum elderi (Blackburn, 1892)
* Gonocephalum elegans Chatanay, 1917
* Gonocephalum elytrale Kaszab, 1952
* Gonocephalum endroedii Kaszab, 1952
* Gonocephalum ermischi Kaszab, 1960
* Gonocephalum espagnoli Kaszab, 1965
* Gonocephalum fallaciosum Gridelli, 1948
* Gonocephalum feae Gebien, 1920
* Gonocephalum fernandezi Ferrer, 2004
* Gonocephalum ficifolium Chûjô, 1995
* Gonocephalum ficifoliu , M. T. Chûjô, 1994.
* Gonocephalum folini Ferrer, 1995
* Gonocephalum formosanum Gebien, 1913
* Gonocephalum fortidens (Reitter, 1915)
* Gonocephalum foveicolle Kaszab, 1952
* Gonocephalum foveoseriatum Chatanay, 1914
* Gonocephalum frederici Ferrer, 1995
* Gonocephalum freudei Kaszab, 1960
* Gonocephalum freyi Kaszab, 1952
* Gonocephalum fuscosetosum Chatanay, 1917
* Gonocephalum gallagheri Ferrer, 1997
* Gonocephalum gebieni Gridelli, 1945
* Gonocephalum gebienianum Kaszab, 1952
* Gonocephalum geneirotundum Ren, 1998
* Gonocephalum germari Ferrer, 2000
* Gonocephalum gerstaeckeri Ferrer, 1995
* Gonocephalum gestroi Gebien, 1920
* Gonocephalum girardi Ferrer, 1995
* Gonocephalum giraudini Ardoin, 1965
* Gonocephalum gracile (Bates, 1879)
* Gonocephalum granohispidum Kolbe, 1897
* Gonocephalum granosum Gebien, 1921
* Gonocephalum granulatipenne Kaszab, 1978
* Gonocephalum granulatum (Fabricius, 1791)
* Gonocephalum granulipenne Gridelli, 1945
* Gonocephalum greensladei Kaszab, 1980
* Gonocephalum gridellianum Kaszab, 1952
* Gonocephalum gridellii Koch, 1953
* Gonocephalum grimmi Ferrer, 1995
* Gonocephalum griseovittatum Gridelli, 1940
* Gonocephalum griseovittatum lomii, Ferrer, 2000
* Gonocephalum gruveli Ferrer, 2004
* Gonocephalum guerryi Chatanay, 1917
* Gonocephalum guignoti Kaszab, 1960
* Gonocephalum gustafssoni Ferrer, 1995
* Gonocephalum hackeri Carter, 1928
* Gonocephalum hadroide (Fairmaire, 1888)
* Gonocephalum hasticolle Chatanay, 1917
* Gonocephalum hauschildi Kaszab, 1952
* Gonocephalum helaeoides Kaszab, 1952
* Gonocephalum helferi Kaszab, 1952
* Gonocephalum helopioide (Fairmaire, 1894)
* Gonocephalum hiekei Ferrer, 1995
* Gonocephalum himalayense Kaszab, 1952
* Gonocephalum hingstoni Kaszab, 1952
* Gonocephalum hintoni Kaszab, 1952
* Gonocephalum hiranoi Akita et Masomoto, 2003
* Gonocephalum hispidocostatum (Fairmaire, 1883)
* Gonocephalum hispidulum Kaszab, 1952
* Gonocephalum hoffmannseggi (Steven, 1829)
* Gonocephalum horni Kaszab, 1952
* Gonocephalum humeridens (Fairmaire, 1883)
* Gonocephalum imitator Ferrer, 1995
* Gonocephalum impictum (Fairmaire, 1849)
* Gonocephalum impressiceps Kaszab, 1952
* Gonocephalum impressiusculum Gebien, 1920
* Gonocephalum inaequale Gridelli, 1945
* Gonocephalum incisum (Blanchard, 1853)
* Gonocephalum indictum Ferrer, 1995
* Gonocephalum indicum (Kaszab, 1952)
* Gonocephalum inquinatum (Sahlberg, 1923)
* Gonocephalum insulana Ardoin, 1965
* Gonocephalum insulana halsteadi, Ferrer, 2000
* Gonocephalum insulanum (Olliff, 1887)
* Gonocephalum interruptum Gridelli, 1948
* Gonocephalum irroratum Fauvel, 1867
* Gonocephalum japanum Motschulsky, 1860
* Gonocephalum javae Ferrer, 2010
* Gonocephalum javanicum Kaszab, 1952
* Gonocephalum joliveti Ardoin, 1965
* Gonocephalum juanense Ferrer, 2000
* Gonocephalum kalidii Skopin, 1964
* Gonocephalum kandaharicum Kaszab, 1960
* Gonocephalum karakorumense Kaszab, 1961
* Gonocephalum karatalica Skopin, 1966
* Gonocephalum kaszabi Ferrer, 1995
* Gonocephalum kazenasi Skopin, 1961
* Gonocephalum kempsteri Ferrer, 2000
* Gonocephalum kenyense Ardoin, 1967
* Gonocephalum klapperichi Kaszab, 1952
* Gonocephalum kochi Kaszab, 1952
* Gonocephalum kolbei Gridelli, 1945
* Gonocephalum kondoi Masumoto, 1983
* Gonocephalum konoi Kaszab, 1952
* Gonocephalum koreanum Kaszab, 1952
* Gonocephalum kuehnelti Kaszab, 1961
* Gonocephalum kuluanum Kaszab, 1952
* Gonocephalum kulzeri Kaszab, 1952
* Gonocephalum kuntzeni Kaszab, 1952
* Gonocephalum labriquei Ferrer, 2010
* Gonocephalum laosense Kaszab, 1952
* Gonocephalum laticolle Gebien, 1910
* Gonocephalum lefranci (Fairmaire, 1863)
* Gonocephalum lesnei Ferrer, 1995
* Gonocephalum lewisi Blair, 1922
* Gonocephalum litorale Chatanay, 1914
* Gonocephalum lomii n. sp. Gridelli det.") (in litt.) Mojo Atschavo,
* Gonocephalum longitarse Kaszab, 1952
* Gonocephalum longulum Gebien, 1911
* Gonocephalum lucidicolle Ardoin, 1965
* Gonocephalum macleay Blackburn, 1907
* Gonocephalum macrophthalmum Kaszab, 1952
* Gonocephalum madagascariense Chatanay, 1914
* Gonocephalum madurense Kaszab, 1952
* Gonocephalum malayanum Gebien, 1935
* Gonocephalum malcyi Ardoin, 1966
* Gonocephalum marani Kaszab, 1952
* Gonocephalum mariei Chatanay, 1914
* Gonocephalum martensi Kaszab, 1977
* Gonocephalum martini Ferrer, 1995
* Gonocephalum marylinneae Ferrer, 2000
* Gonocephalum massauense Gridelli, 1945
* Gonocephalum mastersi (MacLeay, 1872)
* Gonocephalum medvedevi Kaszab, 1965
* Gonocephalum merensi Uyttenboogaart, 1929
* Gonocephalum merkli Ferrer, 2000
* Gonocephalum mesopotamicum Blair, 1923
* Gonocephalum meyricky Blackburn, 1894
* Gonocephalum micantipenne Fairmaire, 1893
* Gonocephalum michaelseni Gebien, 1920
* Gonocephalum minusculum (Fairmaire, 1894)
* Gonocephalum misellum Blackburn, 1907
* Gonocephalum miyakense Nakane, 1963
* Gonocephalum modestum (Reiche, 1847)
* Gonocephalum moluccanum (Blanchard, 1853)
* Gonocephalum moraguesi Ferrer, 2000
* Gonocephalum muelleri Gridelli, 1945
* Gonocephalum mysorense Kaszab, 1952
* Gonocephalum neoblairi Iwan, 2010
* Gonocephalum nepalicum Kaszab, 1973
* Gonocephalum niloticum Ardoin, 1965
* Gonocephalum novaecaledoniae Kaszab, 1982
* Gonocephalum nyassae Ferrer, 2000
* Gonocephalum obenbergeri Kaszab, 1952
* Gonocephalum oblitum (Wollaston, 1864)
* Gonocephalum oblongum (Fabricius, 1801)
* Gonocephalum obscurum (Küster, 1849)
* Gonocephalum oceanicum Kaszab, 1985
* Gonocephalum oculare Kaszab, 1952
* Gonocephalum okinawanum Chujô, 1963
* Gonocephalum omoense Ardoin, 1967
* Gonocephalum opacum Walker, 1871
* Gonocephalum outreyi Chatanay, 1917
* Gonocephalum papuanum Gebien, 1920
* Gonocephalum papulosum (Fairmaire, 1896)
* Gonocephalum parcesetosum Kaszab, 1952
* Gonocephalum patricium Kaszab, 1952
* Gonocephalum patrizii Gridelli, 1948
* Gonocephalum patruele (Erichson, 1843)
* Gonocephalum peguanum Kaszab, 1952
* Gonocephalum peregrinum Kolbe, 1902
* Gonocephalum perplexum (Lucas, 1849)
* Gonocephalum persimile (Lewis, 1894)
* Gonocephalum philippinense Kaszab, 1952
* Gonocephalum pilosa Ferrer, 2000
* Gonocephalum planicolle Kaszab, 1952
* Gonocephalum platipenne Kaszab, 1960
* Gonocephalum podagrarium (Kaszab, 1939)
* Gonocephalum politum Ferrer, 2000
* Gonocephalum pottsi Kulzer, 1957
* Gonocephalum prolixum (Erichson, 1843)
* Gonocephalum pseudopubens Kaszab, 1952
* Gonocephalum pubens (Marseul, 1876)
* Gonocephalum pubescens (Palisot de Beauvois, 1821)
* Gonocephalum pubiferum Reitter, 1904
* Gonocephalum pulverulentum Olivier, 1811
* Gonocephalum pygmaeum (Steven, 1829)
* Gonocephalum rastellii Ferrer, 1997
* Gonocephalum recticolle Motschulsky, 1866
* Gonocephalum recurvum Gebien, 1920
* Gonocephalum remedellii Gridelli, 1945
* Gonocephalum reticulatum Motschulsky, 1854
* Gonocephalum rileyi Kaszab, 1952
* Gonocephalum rondoni Kaszab, 1972
* Gonocephalum roseni Kaszab, 1952
* Gonocephalum rostratum Bremer et Ferrer, 2002
* Gonocephalum rotundatum Gridelli, 1940
* Gonocephalum royi Ardoin, 1961
* Gonocephalum rugatulum (Fairmaire, 1893)
* Gonocephalum rusticum (Olivier, 1811)
* Gonocephalum sauteri Kaszab, 1952
* Gonocephalum sawadai Masumoto, 1985
* Gonocephalum schelpi Bremer et Ferrer, 1992
* Gonocephalum schneideri Reitter, 1898
* Gonocephalum schusteri Kaszab, 1952
* Gonocephalum sculptithorax Gridelli, 1948
* Gonocephalum scutellare (Perty, 1831)
* Gonocephalum semipatruele Kaszab, 1952
* Gonocephalum senkakuense Chujô, 1973
* Gonocephalum seriatum (Boisduval, 1835)
* Gonocephalum sericeum (Baudi, 1875)
* Gonocephalum setulosum (Faldermann, 1837)
* Gonocephalum sexuale (Marseul, 1876)
* Gonocephalum shimoganum Kaszab, 1952
* Gonocephalum sibuyanum Kaszab, 1952
* Gonocephalum sikkimense Kaszab, 1952
* Gonocephalum simplex (Fabricius, 1801)
* Gonocephalum simplicipes Gridelli, 1948
* Gonocephalum simulatrix (Fairmaire, 1891)
* Gonocephalum sinicum Reichardt, 1936
* Gonocephalum soederbomi Ferrer, 2010
* Gonocephalum somalica (Ardoin, 1965)
* Gonocephalum somalicum Gridelli, 1945
* Gonocephalum soricinum (Reiche et Saulcy, 1857)
* Gonocephalum spangleri Kaszab, 1980
* Gonocephalum spinicolle (Fairmaire, 1896)
* Gonocephalum stevensi Kaszab, 1952
* Gonocephalum stoeckleini Kaszab, 1952
* Gonocephalum strangulatum (Fairmaire, 1888)
* Gonocephalum strigosum (Reiche, 1850)
* Gonocephalum subcontractum Gridelli, 1945
* Gonocephalum subrugulosum Reitter, 1887
* Gonocephalum subsetosum (Kolbe, 1883)
* Gonocephalum subspinosum (Fairmaire, 1894)
* Gonocephalum subsulcatum (Reiche, 1850)
* Gonocephalum swazilandense Ferrer, 1995
* Gonocephalum szekessyi Kaszab, 1952
* Gonocephalum takara Nakane, 1963
* Gonocephalum tellinii Gridelli, 1945
* Gonocephalum tenasserimicum Kaszab, 1952
* Gonocephalum tenuicorne Kaszab, 1952
* Gonocephalum tenuipes Kaszab, 1952
* Gonocephalum terminale Reichardt, 1936
* Gonocephalum thailandicum Kaszab et Chujô, 1966
* Gonocephalum tibetanum Kaszab, 1952
* Gonocephalum titschacki Kaszab, 1952
* Gonocephalum tomentosum (Walker, 1871)
* Gonocephalum tonkinense Kaszab, 1952
* Gonocephalum topali Kaszab 1975
* Gonocephalum torridum (Champion, 1894)
* Gonocephalum triste Ferrer, 2000
* Gonocephalum tschilianum Kaszab, 1952
* Gonocephalum tuberculatum (Hope, 1831)
* Gonocephalum tuberculiferum Ferrer, 1995
* Gonocephalum uhligi Ferrer, 2002
* Gonocephalum ulugurense Ardoin, 1976
* Gonocephalum uniseriatum Kaszab, 1952
* Gonocephalum vagum (Steven, 1829)
* Gonocephalum validum Ferrer, 2000
* Gonocephalum veigae Koch, 1956
* Gonocephalum verrucosum (Fairmaire, 1849)
* Gonocephalum vientianeum Kaszab, 1972
* Gonocephalum villiersi Gridelli, 1948
* Gonocephalum walkeri Champion, 1894
* Gonocephalum wau Kaszab, 1970
* Gonocephalum wittmeri Kaszab, 1975
* Gonocephalum woynarovichi Kaszab, 1973
* Gonocephalum wui Ren, 1995
* Gonocephalum zambesense Ferrer, 1995
* Gonocephalum zoltani Iwan, 2010