# List of penguins

Penguins are birds in the family Spheniscidae in the monotypic order Sphenisciformes. They inhabit high-productivity marine habitats, almost exclusively in the Southern Hemisphere; the only species to occur north of the Equator is the Galapagos penguin. The only group of birds other than the ratites to be entirely flightless, penguins are extremely adapted to their aquatic lifestyle, with a streamlined shape that minimizes drag, countershaded dark-and-white plumage, dense bones, powerful flippers, and insulating feathers that allow them to withstand very low temperatures on land and in water.
There are currently 18 extant species of penguins recognised by the International Ornithologists' Union, distributed among six genera. Many species of fossil penguins are known from the Paleocene onwards; however, their exact number and taxonomy are unsettled due to ongoing discoveries.

## Conventions

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the penguin's range is provided. Ranges are based on the IOC World Bird List for that species unless otherwise noted. Population estimates are of the number of mature individuals and are taken from the IUCN Red List.
This list follows the taxonomic treatment (designation and order of species) and nomenclature (scientific and common names) of version 13.2 of the IOC World Bird List. Where the taxonomy proposed by the IOC World Bird List conflicts with the taxonomy followed by the IUCN or the 2023 edition of The Clements Checklist of Birds of the World, the disagreement is noted next to the species's common name (for nomenclatural disagreements) or scientific name (for taxonomic disagreements).

## Classification

The International Ornithologists' Union (IOU) recognises 18 species of penguins in six genera. This list does not include hybrid species, extinct prehistoric species, or putative species not yet accepted by the IOU.
Family Spheniscidae
* Genus Aptenodytes: two species
* Genus Pygoscelis: three species
* Genus Eudyptula: one species
* Genus Spheniscus: four species
* Genus Megadyptes: one species
* Genus Eudyptes: seven species

## Penguins

Genus Aptenodytes – Miller, J. F., 1778 – 2 species :
| Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
| King penguin | A. patagonicus Miller, J. F., 1778 | Subantarctic islands | LC  1,084,320–1,228,320 breeding pairs |
| Emperor penguin | A. forsteri Gray, G. R., 1884 | Antarctic sea ice | NT  256,500 breeding pairs |
Genus Pygoscelis – Wagler, 1832 – 3 species :
| Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
| Adélie penguin | P. adeliae (Hombron and Jacquinot, 1841) | Antarctica and surrounding islands | LC  1,084,320–1,228,320 breeding pairs |
| Chinstrap penguin | P. antarcticus (Forster, J. R., 1781) | Antarctic Peninsula and Balleny Islands | LC  8,000,000 |
| Gentoo penguin | P. papua (Forster, J. R., 1781) Four subspecies P. p. taeniata P. p. papua P. p. ellsworthi P. p. poncetii | Subantarctic islands and locally in Antarctica | LC  774,000 |
Genus Eudyptula – Bonaparte, 1856 – 1 species :
| Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
| Little penguin | E. minor (Forster, J. R., 1781) Six subspecies E. m. novaehollandiae E. m. iredalei E. m. variabilis E. m. minor E. m. albosignata E. m. chathamensis | Australia and New Zealand | LC  469,760 |
Genus Spheniscus – Brisson, 1760 – 4 species :
| Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
| Galapagos penguin | S. mendiculus Sundevall, 1871 | Galápagos Islands | EN  1,200 |
| Humboldt penguin | S. humboldti Meyen, 1834 | South America | VU  23,800 |
| Magellanic penguin | S. magellanicus (Forster, J. R., 1781) | South America | LC  2,200,000–3,200,000 |
| African penguin | S. demersus (Linnaeus, 1758) | Coasts of Southern Africa | CR  19,800 |
Genus Megadyptes – Milne-Edwards, 1880 – 1 species :
| Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
| Yellow-eyed penguin | M. antipodes (Hombron and Jacquinot, 1841) | New Zealand | EN  2,600–3,000 |
Genus Eudyptes – Vieillot, 1816 – 7 species :
| Common name | Scientific name and subspecies | Range | IUCN status and estimated population |
| Macaroni penguin | E. chrysolophus (Brandt, J. F., 1837) | Antarctic Peninsula, South America, and subantarctic islands in South Atlantic and Indian Oceans | VU  6,300,000 breeding pairs |
| Royal penguin | E. schlegeli Finsch, 1876 | Macquarie Island and nearby islands | LC  1,340,000–1,660,000 |
| Northern rockhopper penguin | E. moseleyi Mathews & Iredale, 1921 | Tristan da Cunha, Amsterdam Island, and St. Paul Island | EN  413,700 |
| Southern rockhopper penguin | E. chrysocome (Forster, J. R., 1781) Two subspecies E. c. filholi E. c. chrysocome | South America and subantarctic islands in the Indian Ocean and New Zealand | VU  2,500,000 |
| Fiordland penguin | E. pachyrhynchus Gray, G. R., 1845 | New Zealand | NT  12,500–50,000 |
| Snares penguin | E. robustus Oliver, 1953 | Snares Islands | VU  63,000 |
| Erect-crested penguin | E. sclateri Buller, 1888 | Bounty and Antipodes Islands | EN  150,000 |

## Fossil species
### Basal sphenisciformes

| Species | Source |
| Waimanu manneringi | Slack et al. 2006 |
| Muriwaimanu tuatahi | Mayr et al. 2017 |
| Sequiwaimanu roseae | Mayr et al. 2017 |
| Crossvallia unienwillia | Jadwiszczak et al. 2013 |
| Crossvallia waiparensis | Mayr et al. 2019 |
| Petradyptes stonehousei | Ksepka et al. 2023 |
| Kaiika maxwelli | Fordyce & Thoman 2011 |
| Kumimanu biceae | Mayr et al. 2017 |
| Kumimanu fordecyi | Ksepka et al. 2023 |
| Kupoupou stilwelli | Blokland et al. 2019 |

### Advanced sphenisciformes

| Species | Notes | Source |
| Anthropornis nordenskjoldii | | Jadwiszczak et al. 2002 |
| Anthropornis grandis | | Jadwiszczak et al. 2002 |
| UCMP 321021 (Anthropornis sp.) | | Ksepka & Clarke, 2010 |
| Palaeeudyptes antarcticus | | |
| Palaeeudyptes marplesi | | |
| Palaeeudyptes klekowskii | | Jadwiszczak et al. 2002 |
| Palaeeudyptes gunnari | May represent the opposite sex of P. klekowskii | Jadwiszczak et al. 2002 |
| Chilean Palaeeudyptes | | |
| Burnside "Palaeeudyptes" | | Ksepka & Clarke, 2010 |
| Duntroon "Palaeeudyptes" | | Ksepka & Clarke, 2010 |
| Archaeospheniscus lowei | | Giovanardi et al. 2021 |
| Archaeospheniscus lopdelli | | Giovanardi et al. 2021 |
| Notodyptes wimani | Formerly Archaeospheniscus | Giovanardi et al. 2021 |
| Delphinornis larseni | | Jadwiszczak et al. 2002 |
| Delphinornis gracilis | | Jadwiszczak et al. 2002 |
| Delphinornis arctowskii | | Jadwiszczak et al. 2002 |
| Mesetaornis polaris | | Jadwiszczak et al. 2002 |
| Marambiornis exilis | | Jadwiszczak et al. 2002 |
| Aprosdokitos mikrotero | | Hospitaleche et al. 2017 |
| Perudyptes devriesi | | Ksepka & Clarke, 2010 |
| Eretiscus tonni | | Hospitaleche & Tambussi, 2008 |
| Palaeospheniscus patagonicus | | Hospitaleche & Tambussi, 2008 |
| Palaeospheniscus bilocular | | Hospitaleche & Tambussi, 2008 |
| Palaeospheniscus bergi | | Hospitaleche & Tambussi, 2008 |
| Paraptenodytes antarcticus | | Hospitaleche & Tambussi, 2008 |
| Arthrodytes andrewsi | | Hospitaleche & Tambussi, 2008 |
| Madrynornis mirandus | | Hospitaleche & Tambussi, 2008 |
| Pachydyptes simpsoni | | Park & Fitzgerald, 2012 |
| Pachydyptes ponderosus | | Giovanardi et al. 2021 |
| Anthropodyptes gilli | | Park & Fitzgerald, 2012 |
| Pseudaptenodytes macraei | | Park & Fitzgerald, 2012 |
| Marambiornopsis sobrali | | Jadwiszczak et al. 2021 |
| Icadyptes salasi | | Clarke et al. 2007 |
| Kairuku waewaeroa | | Giovanardi et al. 2021 |
| Kairuku grebneffi | | Giovanardi et al. 2021 |
| Kairuku waitaki | | Giovanardi et al. 2021 |
| Glen Murray Kairuku sp. | | Giovanardi et al. 2021 |
| Platydyptes novaezealandiae | | Giovanardi et al. 2021 |
| Platydyptes marplesi | | Giovanardi et al. 2021 |
| Inkayacu paracasensis | | Clarke et al. 2010 |
| Pakudyptes hakataramea | | Ando et al. 2024 |

### Extinct species of extant genera

| Species | Source |
| Spheniscus megaramphus | Hospitaleche & Tambussi, 2008 |
| Spheniscus urbinai | Hospitaleche & Tambussi, 2008 |
| Spheniscus chilensis | Hospitaleche & Tambussi, 2008 |
| Spheniscus muizoni | Hospitaleche & Tambussi, 2008 |
| Pygoscelis grandis | Hospitaleche & Tambussi, 2008 |
| Pygoscelis tyreei | |
| Pygoscelis calderensis | Hospitaleche & Tambussi, 2008 |
| Eudyptes atatu | Thomas et al. 2020 |
| Eudyptes warhami | Cole et al. 2019 |
| Eudyptes calauina | Hoffmeister et al. 2014 |
| Eudyptula wilsonae | Thomas et al. 2023 |
| Megadyptes antipodes richdalei | Cole et al. 2019 |
| Megadyptes antipodes waitaha | Cole et al. 2019 |
| Aptenodytes ridgeni | |

### Poorly understood taxa

| Species | Notes | Source |
| Dege hendeyi | | |
| Inguza predemersus | | |
| Duntroonornis parvus | | |
| Nucleornis insolitus | | |
| Marplesornis novaezealandiae | | |
| Korora oliveri | | |
| Paraptenodytes robustus | | |
| Platydyptes amiesi | | |
| "Pakudyptes hakataramea" | Also called the "Hakataramea Penguin" | Ando, 2007 (unpublished) |

#### Invalid taxa

| Species | Notes | Source |
| Tonniornis mesetaensis | Named on an isolated humerus which is unable to be compared to other species present in the same locality. | Ksepka & Clarke, 2010 |
| Tonniornis minimum | Named on an isolated humerus which is unable to be compared to other species present in the same locality. Similar size to a specimen referred to Delphinornis larseni. | Ksepka & Clarke, 2010 |
| Tereingaornis moisleyi | Genus is a nomen dubium due to lack of identifiable traits, but the specimen may belong to a new species (the Te Ringa Falls Penguin) | Thomas et al. 2019 |
| Wimanornis seymourensis | Synonymous with Palaeeudypytes gunnari | |
| Orthopteryx gigas | | Hospitaleche and Reguero, 2010 |
| Ichtyopteryx gracilis | | Hospitaleche and Reguero, 2010 |
| Palaeoapterodytes ictus | | Hospitaleche, 2010 |
| Psuedaptenodytes minor | Lack of diagnostic characteristics on the holotype | Park & Fitzgerald, 2012 |
The Early Oligocene genus Cruschedula was formerly thought to belong to Spheniscidae; however, re-examination of the holotype in 1943 resulted in the genus being placed in Accipitridae. Further examination in 1980 resulted in placement as Aves incertae sedis.