# 2020 in paleobotany

This article records new taxa of fossil plants that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2020.

## Flowering plants
### Alismatales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Limnobiophyllum pedunculatum | Sp. nov | Valid | Low, Su & Xing in Low et al. | Late Oligocene | | China | A member of the family Araceae. | |

### Apiales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Paleopanax puryearensis | Sp. nov | Valid | Na, Blanchard & Wang | Middle Eocene | Cockfield Formation | United States ( Tennessee) | A member of the family Araliaceae. | |

### Arecales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Echimonocolpites chicxulubensis | Sp. nov | Valid | Smith et al. | Eocene (Ypresian) | | Mexico | Pollen of a flowering plant, probably a member of the family Arecaceae. |
| Palmoxylon ceroxyloides | Sp. nov | In press | Khan, Hazra & Bera in Khan et al. | Late Cretaceous (Maastrichtian)-Paleocene (Danian) | Deccan Intertrappean Beds | India | A petrified palm stem of a member of the subfamily Ceroxyloideae. |
| Palmoxylon dindoriensis | Sp. nov | Valid | Khan, Roy & Bera in Khan et al. | Late Cretaceous (Maastrichtian)-Paleocene (Danian) | Deccan Intertrappean Beds | India | A petrified palm stem. |
| Sabalites dawsonii | Sp. nov | Valid | Greenwood & Conran | Eocene | | Canada ( British Columbia) | |
| Sabalites karondiensis | Sp. nov | In press | Roy, Hazra & Khan in Roy et al. | Late Cretaceous-Paleocene (latest Maastrichtian-earliest Danian) | Deccan Intertrappean Beds | India | A palm frond . |
| Spinizonocolpites riochiquensis | Sp. nov | Valid | Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal | Late Cretaceous (Maastrichtian) | Lago Colhué Huapí Formation | Argentina | A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020. |

### Buxales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Pachysandra europaea | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A species of Pachysandra. Announced in 2019; the final version of the article naming it was published in 2020. | |

### Caryophyllales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Gomphrenipollis garciae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant, possibly produced by members of the family Amaranthaceae. Announced in 2020; the final version of the article naming it was published in 2021. | |

### Chloranthales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Sarcandraxylon | Gen. et sp. nov | Valid | Pipo, Iglesias & Bodnar | Late Cretaceous (early–middle Campanian) | Santa Marta Formation | Antarctica | A member of the family Chloranthaceae. Genus includes new species S. sanjosense. | |

### Cornales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Amersinia littletonensis | Sp. nov | Valid | Huegele & Manchester | Early Paleocene | Denver Formation | United States ( Colorado) | |
| Langtonia parva | Sp. nov | Valid | Huegele & Manchester | Early Paleocene | Denver Formation | United States ( Colorado) | A member of the family Mastixiaceae. |
| Mastixicarpum hoodii | Sp. nov | Valid | Huegele & Manchester | Early Paleocene | Denver Formation | United States ( Colorado) | A member of the family Mastixiaceae. |
| Nyssa gergoei | Sp. nov | Valid | Hably | Miocene | | Hungary | A tupelo. |
| Nyssa gyoergyi | Sp. nov | Valid | Hably | Miocene | | Hungary | A tupelo. |
| Platycrater iljinskajae | Sp. nov | In press | Denk et al. | Late Oligocene | | Russia | |
| Portnallia alexanderi | Sp. nov | Valid | Huegele & Manchester | Early Paleocene | Denver Formation | United States ( Colorado) | A member of the family Mastixiaceae. |

### Crossosomatales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Staphylea woodworthensis | Sp. nov | In press | Zhu & Manchester | Oligocene | Renova Formation | United States ( Montana) | A species of Staphylea. | |

### Cucurbitales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Coriaripites goodii | Sp. nov | Valid | Barreda, Palazzesi & Tellería in Renner et al. | Late Cretaceous (Campanian–early Maastrichtian) | Lopez de Bertodano Formation Santa Marta Formation Snow Hill Island Formation | Antarctica | Pollen grains similar to those of extant members of the genus Coriaria. | |
| Echitriporites jolyi | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the genus Cayaponia. Announced in 2020; the final version of the article naming it was published in 2021. | |

### Ericales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Andrewsiocarpon puryearensis | Sp. nov | Valid | Na, Blanchard & Wang | Middle Eocene | Cockfield Formation | United States ( Tennessee) | A member of the family Theaceae. | |
| Anubiscarpon | Gen. et sp. nov | Valid | Smith & Manchester | Middle Eocene | Clarno Formation | United States ( Oregon) | A member of the family Theaceae. Genus includes new species A. andersonae. | |

### Fabales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Cercis zekuensis | Sp. nov | Valid | Li et al. | Early Miocene | | China | A species of Cercis. Announced in 2020; the final version of the article naming it was published in 2021. |
| Gleditsia pliocaenica | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A species of Gleditsia. Announced in 2019; the final version of the article naming it was published in 2020. |
| Menendoxylon lutzi | Sp. nov | In press | Baez & Crisafulli | Miocene | Chiquimil | Argentina | Fossil wood of a member of the family Fabaceae. |
| Parkiidites marileae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the genus Parkia. Announced in 2020; the final version of the article naming it was published in 2021. |
| Prioria martineziorum | Sp. nov | Valid | Rodríguez-Reyes & Estrada-Ruiz | Oligocene-Miocene | Santiago Formation | Panama | A species of Prioria. |
| Psilastephanocolporites deoliverae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the family Polygalaceae. Announced in 2020; the final version of the article naming it was published in 2021. |
| Psilastephanocolporites endoporatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Announced in 2020; the final version of the article naming it was published in 2021. |
| Striatopollis grahamii | Sp. nov | Valid | Smith et al. | Eocene (Ypresian) | | Mexico | Pollen of an eudicot, probably a member of the family Fabaceae. |

### Fagales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Alnus chaybulakensis | Sp. nov | Valid | Averyanova & Xing | Paleogene | | Kazakhstan | An alder. |
| Berryophyllum hainanensis | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | A member of the family Fagaceae. |
| Carpinus asymmetrica | Sp. nov | In press | Xue & Jia in Xue et al. | Early Miocene | Maguan | China | A species of Carpinus. |
| Carpinus symmetrica | Sp. nov | In press | Xue & Jia in Xue et al. | Early Miocene | Maguan Basin | China | A species of Carpinus |
| Carya pipecreekensis | Sp. nov | In press | Swinehart & Farlow | Late Neogene | Pipe Creek Sinkhole | United States ( Indiana) | A hickory. |
| Castaneophyllum hainanensis | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | A member of the family Fagaceae. |
| Castaneophyllum lanceolata | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | A member of the family Fagaceae. |
| Castanopsis bulgarica | Sp. nov | Valid | Mantzouka, Ivanov & Bozukov | Late Miocene–early Pliocene (late Messinian–early Zanclean) | Pokrovnik | Bulgaria | A species of Castanopsis. Announced in 2020; the final version of the article naming it was published in 2021. |
| Lithocarpus changchangensis | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | A species of Lithocarpus. |
| Quercus aimeeana | Sp. nov | Valid | Sadowski, Schmidt & Denk | Eocene | Baltic amber | Europe (Baltic Sea region) | An oak. |
| Quercus borissovii | Sp. nov | Valid | Averyanova & Xing | Paleogene | | Kazakhstan | An oak. |
| Quercus campanulata | Sp. nov | Valid | Sadowski, Schmidt & Denk | Eocene | Baltic amber | Europe (Baltic Sea region) | An oak. |
| Quercus casparyi | Sp. nov | Valid | Sadowski, Schmidt & Denk | Eocene | Baltic amber | Europe (Baltic Sea region) | An oak. |
| Quercus changchangensis | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | An oak. |
| Quercus conwentzii | Sp. nov | Valid | Sadowski, Schmidt & Denk | Eocene | Baltic amber | Europe (Baltic Sea region) | An oak. |
| Quercus longianthera | Sp. nov | Valid | Sadowski, Schmidt & Denk | Eocene | Baltic amber | Europe (Baltic Sea region) | An oak. |
| Quercus multipilosa | Sp. nov | Valid | Sadowski, Schmidt & Denk | Eocene | Baltic amber | Europe (Baltic Sea region) | An oak. |
| Quercus paleoargyrotricha | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | An oak. |
| Quercus paleohypargyrea | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | An oak. |
| Quercus paleolamellosa | Sp. nov | Valid | Liu & Jin in Liu, Song & Jin | Eocene | Changchang Formation | China | An oak. |

### Garryales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Eucommia szaferi | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A species of Eucommia. Announced in 2019; the final version of the article naming it was published in 2020. | |

### Gentianales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Asclepiadospermum | Gen. et 2 sp. nov | Valid | Del Rio et al. | Early Eocene | Niubao Formation | China | An asclepiadoid Apocynaceae genus. Included species A. marginatum and A. ellipticum. | |
| Margocolporites carinae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the genus Rauvolfia. Announced in 2020; the final version of the article naming it was published in 2021. | |

### Icacinales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Iodes elliptica | Sp. nov | In press | Del Rio et al. | Early Oligocene | Wenshan Basin | China | A member of the family Icacinaceae. |
| Iodes passiciensis | Sp. nov | Valid | Del Rio & De Franceschi | Early Eocene | | France | A member of the family Icacinaceae. |
| Manchesteria | Gen. et sp. nov | In press | Stull & Rozefelds in Rozefelds et al. | Cenozoic (mid-Miocene or, more likely, middle Eocene) | | Australia | A member of the family Icacinaceae. Genus includes new species M. australis. |
| Pyrenacantha simonsii | Sp. nov | Valid | Stull et al. | Early Oligocene | | Egypt | A species of Pyrenacantha. |

### Lamiales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Echitricolpites cruziae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the genus Aegiphila. Announced in 2020; the final version of the article naming it was published in 2021. |
| Multiareolites? reticulatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the family Acanthaceae. Announced in 2020; the final version of the article naming it was published in 2021. |
| Retistephanocolpites curvimuratus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Announced in 2020; the final version of the article naming it was published in 2021. |
| Retistephanocolpites pardoi | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant, possibly produced by members of the genus Amphilophium. Announced in 2020; the final version of the article naming it was published in 2021. |

### Laurales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Actinodaphnoxylon | Gen. et sp. nov | In press | Akkemik et al. | Eocene (Lutetian) | | Turkey | A member of the family Lauraceae. Genus includes new species A. zileensis. |
| Mezilaurinoxylon oleiferum | Sp. nov | Valid | Ruiz, Brea & Pujana in Ruiz et al. | Paleocene (Danian) | Salamanca Formation | Argentina | A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Patagonoxylon | Gen. et sp. nov | Valid | Ruiz, Brea & Pujana in Ruiz et al. | Paleocene (Danian) | Salamanca Formation | Argentina | A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Thymolepis | Gen. et sp. nov | Valid | Chambers & Poinar | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra. |
| Valviloculus | Gen. et sp. nov | Valid | Poinar et al. | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | Possibly a member of Laurales related to the families Monimiaceae and Atherospermataceae. Genus includes new species V. pleristaminis. |

### Liliales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Smilax fujianensis | Sp. nov | Valid | Dong et al. | Middle Miocene | | China | A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021. | |
| Smilax zhangpuensis | Sp. nov | Valid | Dong et al. | Middle Miocene | | China | A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021. | |

### Magnoliales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Magnolia nanningensis | Sp. nov | Valid | Huang et al. | Late Oligocene | Nanning Basin | China | A species of Magnolia. |
| Magnolia waltheri | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A species of Magnolia. Announced in 2019; the final version of the article naming it was published in 2020. |
| Melloniflora | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous | Potomac Group | United States ( Virginia) | A relative of extant early-diverging members of the Magnoliales. Genus includes new species M. virginiensis. |

### Malpighiales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Salix palaeofutura | Sp. nov | Valid | Narita et al. | Miocene | Bifuka Formation | Japan | A willow. | |

### Malvales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Bastardioxylon | Gen. et sp. nov | In press | Baez & Crisafulli | Miocene | Chiquimil | Argentina | Fossil wood of a member of the family Malvaceae. Genus includes new species B. antiqua. |
| Dipterocarpus dindoriensis | Sp. nov | Valid | Khan, Spicer & Bera in Khan et al. | Late Cretaceous (Maastrichtian) | Deccan Intertrappean Beds | India | A species of Dipterocarpus. |
| Echiperiporites germeraadii | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Announced in 2020; the final version of the article naming it was published in 2021. |
| Echiperiporites jaramilloi | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the genus Hibiscus. Announced in 2020; the final version of the article naming it was published in 2021. |
| Echiperiporites titanicus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the genus Malachra. Announced in 2020; the final version of the article naming it was published in 2021. |
| Retistephanocolporites elizabeteae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the genus Ceiba. Announced in 2020; the final version of the article naming it was published in 2021. |
| Veraguasoxylon | Gen. et sp. nov | Valid | Rodríguez-Reyes & Estrada-Ruiz | Oligocene-Miocene | Santiago Formation | Panama | A member of the family Malvaceae. Genus includes new species V. panamense. |

### Myrtales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Eucalyptus xoshemium | Sp. nov | Valid | Gandolfo & Zamaloa in Zamaloa, Gandolfo & Nixon | Eocene (Ypresian) | Huitrera Formation | Argentina | A species of Eucalyptus. |
| Mangroveoxylon | Gen. et comb. nov | In press | Moya & Brea | Late Miocene? | Ituzaingó Formation | Argentina | A member of the family Combretaceae; a new genus for "Menendoxylon" areniensis Lutz (1979). |
| Myrceugenellites grandiporosum | Sp. nov | Valid | Ruiz, Brea & Pujana in Ruiz et al. | Paleocene (Danian) | Salamanca Formation | Argentina | A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Primotrapa | Gen. et sp. nov | Valid | Li & Li in Li et al. | Late Eocene to early Miocene | Hannuoba | China  Czech Republic  France  Germany | A member of Trapoideae. Genus includes new species P. weichangensis, as well as "Carpolithus" pomelii Saporta (1878) and "Hemitrapa" alpina Su & Zhou in Su et al. (2018). |

### Nymphaeales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Praenymphaeapollenites | Gen. et sp. nov | Valid | Barrón, Peris & Labandeira in Peris et al. | Cenomanian | Burmese amber | Myanmar | Pollen of a member of Nymphaeaceae. Genus includes new species P. cenomaniensis. | |

### Oxalidales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Cunoniantha | Gen. et sp. nov | Valid | Jud & Gandolfo | Paleocene (early Danian) | Salamanca Formation | Argentina | A member of the family Cunoniaceae. Genus includes new species C. bicarpellata. Announced in 2020; the final version of the article naming it was published in 2021. |
| Elaeocarpus nanningensis | Sp. nov | In press | Liu et al. | Late Oligocene | Yongning Formation | China | A species of Elaeocarpus. |
| Elaeocarpus prelacunosus | Sp. nov | In press | Liu et al. | Late Miocene | Foluo Formation | China | A species of Elaeocarpus. |
| Elaeocarpus preprunifolioides | Sp. nov | In press | Liu et al. | Late Miocene | Foluo Formation | China | A species of Elaeocarpus. |
| Elaeocarpus prerugosus | Sp. nov | In press | Liu et al. | Late Miocene | Foluo Formation | China | A species of Elaeocarpus. |
| Elaeocarpus preserratus | Sp. nov | In press | Liu et al. | Late Miocene | Foluo Formation | China | A species of Elaeocarpus. |
| Elaeocarpus presikkimensis | Sp. nov | In press | Liu et al. | Miocene | Erzitang Formation | China | A species of Elaeocarpus. |

### Poales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Monoporopollenites scabratus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen produced by members of the family Poaceae. Announced in 2020; the final version of the article naming it was published in 2021. | |
| Rhizomatites | Gen. et sp. nov | In press | Robledo & Anzótegui in Robledo et al. | Miocene-Pliocene | | Argentina | A member of Cyperaceae. Genus includes new species R. cyperoides. | |

### Proteales

| Name | Novelty | Status | Authors | Age | Location | Notes |
| Banksia microphylla | Sp. nov | Valid | Carpenter in Carpenter & Milne | Late Eocene | Australia | A species of Banksia. |
| Banksieaeidites zanthus | Sp. nov | Valid | Milne in Carpenter & Milne | Late Eocene | Australia | A Banksia-like pollen. |
| Platanus emryi | Sp. nov | Valid | Huegele, Spielbauer & Manchester | Miocene | United States | A species of Platanus. |

### Ranunculales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Cissampelos defranceschii | Sp. nov | Valid | Del Rio & Su in Del Rio et al. | Middle Eocene | Niubao Formation | China | A species of Cissampelos. Announced in 2020; the final version of the article naming it was published in 2021. |
| Clematis csabae | Sp. nov | Valid | Hably | Miocene | | Hungary | A species of Clematis. |
| Diploclisia praeaffinis | Sp. nov | Valid | Jia et al. | Late Miocene | | China | A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021. |
| Menispermites bangorensis | Sp. nov | Valid | Huang in Del Rio et al. | Middle Eocene | Niubao Formation | China | A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021. |
| Menispermites haominae | Sp. nov | Valid | Huang in Del Rio et al. | Middle Eocene | Niubao Formation | China | A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021. |
| Menispermites tibetica | Sp. nov | Valid | Huang in Del Rio et al. | Middle Eocene | Niubao Formation | China | A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021. |
| Paleoorbicarpum | Gen. et sp. nov | Valid | Han et al. | Paleocene | Sanshui Basin | China | A member of the family Menispermaceae. Genus includes new species P. parvum. |
| Stephania bangorensis | Sp. nov | Valid | Del Rio & Su in Del Rio et al. | Middle Eocene | Niubao Formation | China | A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021. |
| Stephania geniculata | Sp. nov | Valid | Han et al. | Paleocene | Sanshui Basin | China | A species of Stephania |
| Stephania ornamenta | Sp. nov | Valid | Han et al. | Paleocene | Sanshui Basin | China | A species of Stephania |
| Stephania shuangxingii | Sp. nov | Valid | Del Rio & Su in Del Rio et al. | Middle Eocene | Niubao Formation | China | A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021. |

### Rosales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Berhamniphyllum junrongii | Sp. nov | Valid | Zhou, Wang & Huang in Zhou et al. | Late Eocene | Markam Basin | China | A member of the family Rhamnaceae |
| Crataegus pentagynoides | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A species of Crataegus. Announced in 2019; the final version of the article naming it was published in 2020. |
| Hemiptelea kryshtofovichii | Sp. nov | Valid | Averyanova & Xing | Paleogene | | Kazakhstan | A member of the family Ulmaceae. |
| Scabrastephanoporites | Gen. et sp. nov | Valid | Smith et al. | Eocene (Ypresian) | | Mexico | Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis. |

### Sapindales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Acer dombeyopsis | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A maple. Announced in 2019; the final version of the article naming it was published in 2020. |
| Acer viburnoides | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A maple. Announced in 2019; the final version of the article naming it was published in 2020. |
| Acer vitiforme | Sp. nov | Valid | Kvaček, Teodoridis & Denk | Pliocene | | Germany | A maple. Announced in 2019; the final version of the article naming it was published in 2020. |
| Brosipollis reticulatus | Sp. nov | Valid | Smith et al. | Eocene (Ypresian) | | Mexico | Pollen of a flowering plant, probably a member of the family Burseraceae. |
| Choerospondias fujianensis | Sp. nov | In press | Wang et al. | Miocene | | China | A species of Choerospondias. |
| Llanodelacruzoxylon | Gen. et sp. nov | | Rodríguez-Reyes, Estrada-Ruiz & Gasson | Oligocene–Miocene | Santiago Formation | Panama | A member of the family Anacardiaceae. Genus includes new species L. sandovalii. |
| Manchestercarpa | Gen. et sp. nov | Valid | Atkinson | Late Cretaceous (Campanian) | | Canada ( British Columbia) | A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis. |
| Parametopioxylon | Gen. et sp. nov | Valid | Franco et al. | Miocene | Ituzaingó Formation | Argentina | A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum. |
| Psilastephanocolporites hammenii | Sp. nov | Valid | Smith et al. | Eocene (Ypresian) | | Mexico | Pollen of a flowering plant, probably a member of the family Meliaceae |
| Quinquala | Gen. et sp. nov | Valid | Manchester & Disney in Manchester, Disney & Pham | Eocene | Clarno Formation Tepee Trail Formation | United States ( Oregon Wyoming) | A fossil fruit with affinities with the Rutaceae. Genus includes new species Q. obovata. |
| Rousea cavitata | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Announced in 2020; the final version of the article naming it was published in 2021. |

### Saxifragales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Corylopsis grisea | Sp. nov | Valid | Quirk & Hermsen | Early Pliocene | Gray Fossil Site | United States ( Tennessee) | A species of Corylopsis. Announced in 2020; the final version of the article naming it was published in 2021. |
| Itea polyneura | Sp. nov | In press | Huang & Tian in Tian et al. | Oligocene | Huazhige Formation | China | A species of Itea. |
| Protoaltingia | Gen. et sp. nov | Valid | Scharfstein, Stockey & Rothwell | Late Cretaceous (Coniacian) | | Canada ( British Columbia) | A member of the family Altingiaceae. Genus includes new species P. comoxense. |

### Solanales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Physalis hunickenii | Sp. nov | Valid | Deanna, Wilf & Gandolfo | Early Eocene | Laguna del Hunco Formation | Argentina | A species of Physalis. | |

### Trochodendrales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Eotrochion | Gen. et sp. nov | Valid | Manchester, Kvaček & Judd | Paleocene | | United States ( Wyoming) | A member of the family Trochodendraceae. Genus includes new species E. polystylum. Announced in 2020; the final version of the article naming it was published in 2021. |
| Paraconcavistylon | Gen. et comb. nov | Valid | Manchester, Kvaček & Judd | Ypresian | Klondike Mountain Formation | United States ( Washington) | A Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018). Announced in 2020; the final version of the article naming it was published in 2021. |
| Trochodendron infernense | Sp. nov | Valid | Manchester, Kvaček & Judd | Paleocene | | United States ( Wyoming) | A species of Trochodendron. Announced in 2020; the final version of the article naming it was published in 2021. |

### Vitales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Yua jiangxiensis | Sp. nov | Valid | He & Wang | Miocene | Toupi Formation | China | A species of Yua. Announced in 2020; the final version of the article naming it was published in 2021. | |

### Other angiosperms

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Aextoxicoxylon kawasianus | Sp. nov | Valid | Vera et al. | Late Cretaceous | Puntudo Chico Formation | Argentina | A fossil dicot wood |
| Atlantocarpus | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (Albian) | Potomac Group | Portugal United States ( Virginia) | An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species A. virginiensis. |
| Carpolithes gergoei | Sp. nov | Valid | Hably & Erdei in Hably | Miocene | | Hungary | A fossil fruit of a flowering plant of uncertain phylogenetic placement. |
| Catanthus | Gen. et sp. nov | In press | Friis, Crane & Pedersen | Early Cretaceous | | Portugal | An early flowering plant. Genus includes new species C. dolichostemon. |
| Cavilignum | Gen. et sp. nov | Valid | Siegert & Hermsen | Early Pliocene | Gray Fossil Site | United States ( Tennessee) | A flowering plant of uncertain phylogenetic placement, described on the basis of fossil endocarps. Genus includes new species C. pratchettii. |
| Chainandra | Gen. et sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | Genus includes new species C. zeugostylus. |
| Cichoreacidites? flammulatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Crotonoidaepollenites echinatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Cyathitepala | Gen. et sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | Genus includes new species C. papillosa. |
| Dasykothon | Gen. et sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus. |
| Dinganthus | Gen. et sp. nov | Valid | Liu et al. | Miocene | Dominican amber | Dominican Republic | A eudicot of uncertain phylogenetic placement. Genus includes new species D. pentamera. |
| Echistephanoporites annulatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Eofructus | Gen. et sp. nov | Valid | Han & Wang | Early Cretaceous | Yixian Formation | China | An infructescence including a central axis and five fruits resembling Liaoningfructus. Genus includes new species E. liutiaogouensis. |
| Foveotricolporites crassus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Inaperturopollenites microechinatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Ladakhipollenites carmoi | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Lambertiflora | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (Albian) | Potomac Group | United States ( Virginia) | An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species L. elegans. |
| Malvacipolloides diversus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Malvacipolloides echibaculatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Malvacipolloides romeroae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Menatanthus | Gen. et sp. nov | Valid | Uhl, Paudayal & El Atfy in Uhl et al. | Paleocene (Thanetian) | | France | A eudicot of uncertain phylogenetic placement. Genus includes new species M mosbruggeri. Announced in 2020; the final version of the article naming it was published in 2021. |
| Mugideiriflora | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (Aptian-early Albian) | Almargem Formation | Portugal | An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species M. portugallica. |
| Phantophlebia | Gen. et sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla. |
| Platanites willigeri | Sp. nov | Valid | Halamski & Kvaček in Halamski et al. | Late Cretaceous (Santonian) | Czerna Formation | Poland | Trifoliolate platanoid leaves. |
| Psilaperiporites delicatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Psilaperiporites lunaris | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Ranunculacidites reticulatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Rasenganus | Gen. et sp. nov | Valid | Xing & Gu | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A possible epizoochorous fruit. Genus includes new species R. auricularus. |
| Retibrevitricolpites microreticulatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Retibrevitricolporites costaporus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Retibrevitricolporites? toigoae | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Retipollenites solimoensis | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Retitriporites crassoreticulatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Rhoipites alfredii | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021. |
| Singpuria | Gen. et sp. nov | Valid | Ramteke, Manchester & Nagrale in Ramteke et al. | Late Cretaceous (Maastrichtian) | Deccan Intertrappean Beds | India | A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species S. kapgatei. |
| Sinoherba | Gen. et sp. nov | Valid | Liu & Wang in Liu, Chen & Wang | Early Cretaceous (Barremian–Aptian) | Yixian Formation | China | An early monocot. Genus includes new species S. ningchengensis. Announced in 2020; the final version of the article naming it was published in 2021. |
| Varifructus | Gen. et sp. nov | In press | Liu et al. | Early Cretaceous | Yixian Formation | China | An early flowering plant. Genus includes new species V. lingyuanensis. |
| Wireroadia | Gen. et sp. et comb. nov | Valid | Zhang et al. | Late Cretaceous (Cenomanian to Santonian) | | United States ( Alabama Massachusetts New Jersey New York) | A winged fruit of a eudicot of uncertain phylogenetic placement. Genus includes new species W. viccallii, as well as W. major (Hollick). |

## Pinales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Agathis ledongensis | Sp. nov | Valid | Oskolski et al. | Late Oligocene–early Miocene | Qiutangling Formation | China | A species of Agathis |
| Agathoxylon cozzoi | Sp. nov | Valid | Gnaedinger & Zavattieri | Late Triassic | Chihuido Formation | Argentina | |
| Araucaria cuneoi | Sp. nov | Valid | Noll & Kunzmann | Middle Jurassic | | Argentina | A species of Araucaria. |
| Araucaria famii | Sp. nov | Valid | Stockey & Rothwell | Late Cretaceous (Campanian) | | Canada ( British Columbia) | A species of Araucaria. |
| Araucaria fildesensis | Sp. nov | Valid | Shi et al. | Eocene | Fossil Hill Formation | Antarctica (King George Island) | A species of Araucaria |
| Araucaria huncoensis | Sp. nov | Valid | Rossetto-Harris in Rossetto-Harris et al. | Early Eocene | Laguna del Hunco Formation | Argentina | A species of Araucaria. |
| Araucaria stockeyana | Sp. nov | Valid | Noll & Kunzmann | Middle Jurassic | | Argentina | A species of Araucaria. |
| Araucarites pachacuteci | Sp. nov | In press | Martínez in Martínez et al. | Early Cretaceous (Berriasian–Valanginian) | Huancané Formation | Peru | A member of the family Araucariaceae |
| Brachyoxylon zhouii | Sp. nov | Valid | Jiang et al. | Early Cretaceous | Guantou Formation | China | A conifer wood. Announced in 2020; the final version of the article naming it was published in 2021. |
| Brachyphyllum sattlerae | Sp. nov | Valid | Batista et al. | Early Cretaceous (Aptian) | Crato Formation | Brazil | A member of the family Pinidae. |
| Callialastrobus | Gen. et sp. nov | In press | Kvaček & Mendes | Early Cretaceous (late Aptian–early Albian) | Lusitanian Basin | Portugal | A pollen cone of a member of Araucariaceae. Genus includes new species C. sousai. |
| Cedrus anatolica | Sp. nov | Valid | Akkemik | Early Miocene | Hançili Formation | Turkey | A species of Cedrus. Announced in 2020; the final version of the article naming it was published in 2021. |
| Circoporoxylon tibetense | Sp. nov | In press | Xia et al. | Middle Jurassic (Callovian) | Xiali Formation | China | Possibly a member of the family Podocarpaceae. |
| Comoxostrobus | Gen. et sp. nov | Valid | Stockey, Rothwell & Atkinson | Late Cretaceous (early Coniacian) | | Canada ( British Columbia) | A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species C. rossii. |
| Cupressinoxylon klimovii | Nom. nov | Valid | Blokhina | Miocene | | Russia | A member of the family Cupressaceae; a replacement name for Cupressinoxylon biotoides Blokhina (1989). |
| Cupressinoxylon llantenesense | Sp. nov | Valid | Gnaedinger & Zavattieri | Late Triassic | Llantenes Formation | Argentina | |
| Cupressinoxylon manuelii | Sp. nov | In press | Ríos-Santos, Cevallos-Ferriz & Pujana | Late Cretaceous (Campanian-Maastrichtian) | Cabullona Group | Mexico | |
| Ductoagathoxylon wangii | Sp. nov | In press | Gou & Feng in Gou et al. | Middle Jurassic | Xishanyao Formation | China | A conifer stem. |
| Friisia | Gen. et sp. nov | In press | Mendes & Kvaček | Early Cretaceous (late Aptian – early Albian) | Lusitanian Basin | Portugal | An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica. |
| Juniperoxylon acarcaea | Sp. nov | In press | Akkemik | Early Miocene | Hançili Formation | Turkey | A member of the family Cupressaceae. |
| Lesbosoxylon kemaliyensis | Sp. nov | Valid | Akkemik & Mantzouka in Akkemik, Mantzouka & Kıran Yıldırım | Miocene | Divriği Formation | Turkey | A member of the family Pinaceae. |
| Marskea cuspidata | Sp. nov | Valid | Frolov & Mashchuk | Middle Jurassic | Prisayan Formation | Russia | A member of the family Taxaceae. |
| Mukawastrobus | Gen. et sp. nov | Valid | Stockey, Nishida & Rothwell | Late Cretaceous (late Campanian—early Maastrichtian) | | Japan | A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species M. satoi. |
| Piceoxylon yumeniense | Sp. nov | Valid | Zhou, Peng, Deng, Zhang & Yang in Zhou et al. | Early Cretaceous | Xiagou Formation | China | Fossil wood of a member of the family Pinaceae. Announced in 2020; the final version of the article naming it was published in 2021. |
| Pinuxylon selmeierianum | Sp. nov | Valid | Dolezych & Reinhardt | Paleogene | Eureka Sound Group | Canada ( Nunavut) | A member of the family Pinaceae described on the basis of fossil wood |
| Protophyllocladoxylon chijinense | Sp. nov | Valid | Zhou, Peng, Deng, Zhang & Yang in Zhou et al. | Early Cretaceous | Xiagou Formation | China | Announced in 2020; the final version of the article naming it was published in 2021. |
| Tsuga asiatica | Sp. nov | Valid | Wu & Zhou in Wu et al. | Late Paleogene | | China | A species of Tsuga. Announced in 2019; the final version of the article naming it was published in 2020. |

## Other seed plants

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Amyelon turpanense | Sp. nov | In press | Shi, Yu & Sun | Permian (Lopingian) | Wutonggou | China | A root of a member of Cordaitales |
| Androstrobus obovatus | Sp. nov | In press | Bodnar et al. | Late Triassic | Potrerillos Formation | Argentina | A member of Cycadales. |
| Araripestrobus | Gen. et sp. nov | Valid | Seyfullah, Roberts, Schmidt & Kunzmann in Seyfullah et al. | Early Cretaceous (Aptian-Albian) | Crato | Brazil | A seed plant belonging to the group Erdtmanithecales. Genus includes new species A. resinosus. |
| Archaeopetalanthus | Gen. et sp. nov | Valid | Naugolnykh | Carboniferous | Listvjanskaya | Russia | A member of Pinophyta belonging to the group Vojnovskyales. Genus includes new species A. progressus. |
| Battenispermum | Gen. et sp. nov | In press | Mendes, Pedersen & Friis | Early Cretaceous | | Portugal | A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species B. hirsutum. |
| Carpolithus volantus | Sp. nov | Valid | Gómez et al. | Early Cretaceous (Aptian) | La Cantera Formation | Argentina | A fossil seed, possibly produced by a member of Gnetales. |
| Ductolobatopitys | Gen. et sp. nov | Valid | Conceição & Crisafulli in Conceição et al. | Permian (Cisuralian) | Pedra de Fogo Formation | Brazil | A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae. |
| Filigigantopteris | Gen. et 2 sp. nov | In press | Zhou et al. | Late Permian | Nayixiong Formation | China | A gigantopterid. Genus includes new species F. asymmetrica and F. dahaia. |
| Ginkgo pediculata | Sp. nov | In press | Deng, Yang & Zhou | Early Cretaceous | | China | A species of Ginkgo. |
| Jianchangia | Gen. et sp. nov | Valid | Yang, Wang & Ferguson | Early Cretaceous (Aptian) | Jiufotang Formation | China | A member of Ephedraceae. Genus includes new species J. verticillata. |
| Johniphyllum | Gen. et sp. nov | Valid | Looy & Duijnstee | Permian (Guadalupian) | | United States ( Texas) | A member of Voltziales. Genus includes new species J. multinerve. |
| Jordaniopteris | Gen. et comb. nov | Valid | Anderson in Anderson et al. | Permian (possibly Lopingian) | Um Irna Formation | Jordan | A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017). |
| Kaokoxylon brasiliensis | Sp. nov | Valid | Conceição, Neregato & Iannuzzi in Conceição et al. | Permian (Cisuralian) | Pedra de Fogo Formation | Brazil | A conifer described on the basis of fossil wood. |
| Nilssonia mirovanae | Sp. nov | Valid | Čepičková & Kvaček | Late Cretaceous (Cenomanian) | Peruc-Korycany Formation | Czech Republic | A cycad. |
| Novaiorquepitys | Gen. et sp. nov | In press | Conceição & Crisafulli in Conceição et al. | Permian (Cisuralian) | Pedra de Fogo Formation | Brazil | A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi. |
| Ovalocarpus butmanii | Sp. nov | Valid | Naugolnykh & Linkevich | Permian (Artinskian) | | Russia ( Sverdlovsk Oblast) | A member of Ginkgoales belonging to the family Cheirocladaceae. |
| Palaeocupressinoxylon | Gen. et sp. nov | In press | Wan, Yang & Wang | Late Permian | Turpan–Hami Basin | China | A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale. |
| Phoenicopsis anadyrensis | Sp. nov | Valid | Nosova in Zolina et al. | Late Cretaceous–Paleocene (Maastrichtian–Danian) | Rarytkin | Russia ( Chukotka Autonomous Okrug) | A member of Czekanowskiales |
| Pseudovoltzia sapflorensis | Sp. nov | Valid | Looy & Duijnstee | Permian (Guadalupian) | | United States ( Texas) | A member of Voltziales |
| Pterostoma neehoffii | Sp. nov | Valid | Conran et al. | Middle Miocene | | New Zealand | A cycad |
| Pteruchus frenguellii | Sp. nov | In press | Blomenkemper et al. | Late Permian | Umm Irna Formation | Jordan | A pollen organ of a seed fern |
| Pteruchus lepidus | Sp. nov | In press | Blomenkemper et al. | Late Permian | Umm Irna Formation | Jordan | A pollen organ of a seed fern |
| Samaropsis jinchangensis | Sp. nov | Valid | Hua & Sun in Hua et al. | Early Permian | | China | A seed fossil. Announced in 2019; the final version of the article naming was published in 2020. |
| Umaltolepis involuta | Sp. nov | In press | Nosova | Middle Jurassic | Angren Formation | Uzbekistan | |
| Umaltolepis sogdianica | Sp. nov | In press | Nosova | Middle Jurassic | Angren Formation | Uzbekistan | |
| Umkomasia aequatorialis | Sp. nov | In press | Blomenkemper et al. | Late Permian | Umm Irna Formation | Jordan | A cupulate structure of a seed fern |
| Wantus | Gen. et sp. nov | Valid | Looy & Duijnstee | Permian (Guadalupian) | | United States ( Texas) | A member of Voltziales. Genus includes new species W. acaulis. |
| Wudaeophyton | Gen. et sp. nov | Valid | Pšenička et al. | Early Permian | Taiyuan Formation | China | A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii. |
| Yangopteris | Gen. et comb. nov | In press | Zhou et al. | Permian (Asselian) | | China | A seed fern; a new genus for "Alethopteris" ascendens Halle. |
| Yvyrapitys | Gen. et sp. nov | In press | Conceição & Crisafulli in Conceição et al. | Permian (Cisuralian) | Pedra de Fogo Formation | Brazil | A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi. |

## Other plants

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Annularia paisii | Sp. nov | Valid | Correia et al. | Carboniferous (Gzhelian) | | Portugal | |
| Blasiites huolinhensis | Sp. nov | In press | Li et al. | Early Cretaceous | Huolinhe Formation | China | A liverwort belonging to the family Blasiaceae. |
| Botryopteris multifolia | Sp. nov | In press | He et al. | Permian (Lopingian) | Junlian Formation | China | A fern |
| Birisia mandshurica | Sp. nov | Valid | Golovneva, Grabovskiy & Zolina | Early Cretaceous (Albian) | Frentsevka Formation | Russia ( Primorsky Krai) | A fern belonging to the family Dicksoniaceae. |
| Calamites cambrensis | Sp. nov | Valid | Thomas | Carboniferous (Pennsylvanian) | | United Kingdom | |
| Catenuporella | Gen. et sp. nov | Valid | Zhang et al. | Late Ordovician | Ordos Basin | China | A green alga belonging to the group Dasycladales. Genus includes new species C. gigantia. |
| Charaxis spicatus | emend. nov. | Valid | Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al. | Barremian | Maestrat Basin | Spain | Thallus of Echinochara lazarii. Both taxa has been firstly found anatomically attached |
| Chlamydomonas hanublikanus | Sp. nov | | Vršanská & Hinkelman | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A species of Chlamydomonas |
| Circinites | Gen. et sp. nov | In press | Robledo & Anzótegui in Robledo et al. | Miocene-Pliocene | | Argentina | A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides. |
| Clavatisporites cenomaniana | Sp. nov | Valid | Santamarina in Santamarina et al. | Late Cretaceous (Cenomanian) | Mata Amarilla Formation | Argentina | Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Clavator calcitrapus var. jiangluoensis | comb. nov | Valid | Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al. | Barremian | Maestrat Basin | Spain | Clavatoracean species. |
| Collarecodium? nezpercae | Sp. nov | Valid | Bucur & Rigaud in Bucur et al. | Late Triassic (Norian) | | United States ( Idaho) | A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae. |
| Collarisporites minor | Sp. nov | Valid | Santamarina in Santamarina et al. | Late Cretaceous (Cenomanian) | Mata Amarilla Formation | Argentina | Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Coniopteris sandaolingensis | Sp. nov | Valid | Yuan & Sun in Yuan et al. | Middle Jurassic | Xishanyao Formation | China | |
| Dimicheleodendron | Gen. et comb. nov | Valid | Thomas & Cleal | Carboniferous | | United Kingdom | A lycophyte; a new genus for "Lepidodendron" hickii. |
| Drynaria diplosticha | Sp. nov | Valid | Yu & Xie in Yu et al. | Late Miocene | Bangmai Formation | China | A fern belonging to the family Polypodiaceae. |
| Echinochara lazarii | comb. nov | Valid | Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al. | Barremian | Maestrat Basin | Spain | A member of Clavatoraceans. |
| Equicalastrobus pusillus | Sp. nov | Valid | Zhang & Yan in Zhang et al. | Late Triassic | Baojishan Basin | China | A member of Equisetales. Announced in 2020; the final version of the article naming was published in 2021. |
| Equisetum yenbaiense | Sp. nov | Valid | Aung et al. | Late Miocene | | Vietnam | A species of Equisetum |
| Equisetum yongpingense | Sp. nov | Valid | Aung et al. | Late Pliocene | Sanying Formation | Vietnam | A species of Equisetum |
| Filippoporella | Gen. et sp. nov | In press | Sokač & Grgasović | Early Paleocene | | Croatia | A green alga belonging to the group Dasycladales. Genus includes new species F. barattoloi. |
| Frullania partita | Sp. nov | Valid | Li et al. | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A liverwort, a species of Frullania |
| Frullania vanae | Sp. nov | Valid | Mamontov et al. | Eocene | Rovno amber | Ukraine | A liverwort, a species of Frullania |
| Gippslandites | Gen. et sp. nov | Valid | McSweeney, Shimeta & Buckeridge | Late Silurian–early Devonian | | Australia | A member of Zosterophyllaceae. Genus includes new species G. minutus. |
| Gmujij | Gen. et sp. nov | Valid | Pfeiler & Tomescu | Devonian (Emsian) | Battery Point | Canada ( Quebec) | An early euphyllophyte. Genus includes new species G. tetraxylopteroides. Announced in 2020; the final version of the article naming it was published in 2021. |
| Griphoporella minuta | Sp. nov | Valid | Bucur & Peybernes in Bucur et al. | Late Triassic | | Japan | A green alga belonging to the group Dasycladales and the family Triploporellaceae. |
| Hansopteris | Gen. et sp. nov | In press | Zhou et al. | Permian (Asselian) | | China | An anachoropterid fern. Genus includes new species H. uncinatus. |
| Holosporella magna | Sp. nov | Valid | Bucur & Fucelli in Bucur et al. | Late Triassic (Norian) | | United States ( Nevada) | A green alga belonging to the group Dasycladales and the family Triploporellaceae. |
| Holosporella? rossanae | Sp. nov | Valid | Bucur & Del Piero in Bucur et al. | Late Triassic (Norian) | | Canada ( Yukon) | A green alga belonging to the group Dasycladales and the family Triploporellaceae. |
| Inocladus | Gen. et comb. nov | Valid | LoDuca et al. | Silurian | | United States | An alga related to the group Bryopsidales. Genus includes "Buthotrephis" divaricata White (1901), "B." newlini White (1901), "B." lesquereuxi Grote & Pitt (1876) and "Chondrites" verus Ruedemann (1925). Announced in 2020; the final version of the article naming it was published in 2021. |
| Intermurella ordosensis | Sp. nov | Valid | Zhang et al. | Late Ordovician | Ordos Basin | China | A green alga belonging to the group Dasycladales. |
| Jurafructus | Gen. et sp. nov | Valid | Chen et al. | Middle−Late Jurassic | Jiulongshan Formation | China | A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis. |
| Keraphyton | Gen. et sp. nov | Valid | Champreux, Meyer-Berthaud & Decombeix | Devonian (Famennian) | Mandowa Mudstone Formation | Australia | A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae. |
| Khasurtya | Gen. et sp. nov | Valid | Mamontov in Kopylov et al. | Early Cretaceous | | Russia | A liverwort belonging to the group Marchantiidae. Genus includes new species K. ginkgoides. |
| Lepidodendron demkinae | Sp. nov | Valid | Mosseichik | Carboniferous (Viséan) | | Russia | |
| Lobatannularia linjiaensis | Sp. nov | Valid | Xu et al. | Middle Triassic | Linjia Formation | China | A member of Equisetales. |
| Lygodium sanshuiense | Sp. nov | Valid | Naugolnykh et al. | Paleocene | Buxin Formation | China | A species of Lygodium. Announced in 2019; the final version of the article naming it was published in 2020. |
| Metzgeriites kujiensis | Sp. nov | Valid | Katagiri in Katagiri & Shinden | Late Cretaceous (Santonian) | Tamagawa Formation | Japan | A liverwort. |
| Munieria martinclosasi | comb. nov. | Valid | Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al. | Barremian | Lebanon | Lebanon | Clavatoracean thallus. Formerly known as Charaxis martinclosasi |
| Neoarthropitys | Gen. et sp. nov | In press | Gnaedinger et al. | Middle Triassic | Quebrada de los Fósiles | Argentina | A member of Equisetales. Genus includes new species N. gondwanaensis. |
| Osmundacaulis asiatica | Sp. nov | Valid | Cheng et al. | Cretaceous | | China | A member of the family Osmundaceae |
| Osmundacaulis sinica | Sp. nov | Valid | Cheng et al. | Cretaceous | | China | A member of the family Osmundaceae |
| Ovoidites circumplicatus | Sp. nov | Valid | Zavattieri, Gutiérrez & Monti | Middle Triassic | Quebrada de los Fósiles Formation | Argentina | A green alga belonging to the group Zygnematales. |
| Ovoidites tripartitus | Sp. nov | Valid | Zavattieri, Gutiérrez & Monti | Middle Triassic | Quebrada de los Fósiles Formation | Argentina | A green alga belonging to the group Zygnematales. |
| Palaeostachya guanglongii | Sp. nov | In press | Liu et al. | Permian (Asselian) | Taiyuan Formation | China | A member of the family Calamitaceae. |
| Parazosterophyllum | Gen. et sp. nov | Valid | McSweeney, Shimeta & Buckeridge | Late Silurian–early Devonian | | Australia | A member of Zosterophyllaceae. Genus includes new species P. timsiae. |
| Patruliuspora oregonica | Sp. nov | Valid | Bucur & Rigaud in Bucur et al. | Late Triassic (Norian) | | United States ( Oregon) | A green alga belonging to the group Dasycladales and the family Polyphysaceae. |
| Patruliuspora pacifica | Sp. nov | Valid | Bucur, Del Piero & Peyrotty in Bucur et al. | Late Triassic (Norian) | | Canada ( Yukon) | A green alga belonging to the group Dasycladales and the family Polyphysaceae. |
| Pellites hamiensis | Sp. nov | Valid | Li et al. | Middle Jurassic | Xishanyao | China | A liverwort belonging to the family Pelliaceae. |
| Plenasium (Aurealcaulis) elegans | Sp. nov | In press | Hiller et al. | Eocene | Na Duong Formation | Vietnam | A member of Osmundaceae |
| Pleuromeia shaolinii | Sp. nov | Valid | Zhang & Wang in Zhang et al. | Middle Triassic | Linjia | China | |
| Polycingulatisporites multiverrucata | Sp. nov | Valid | Santamarina in Santamarina et al. | Late Cretaceous (Cenomanian) | Mata Amarilla | Argentina | Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it was published in 2020. |
| Polypodiisporites minutiverrucatus | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021. |
| Polysporia baetica | Sp. nov | In press | Álvarez-Vázquez, Bek & Drábková | Carboniferous (Pennsylvanian) | Peñarroya-Belmez-Espiel Coalfield | Spain | A member of Isoetales |
| Polystichum pacltovae | Sp. nov | Valid | Kvaček in Kvaček & Teodoridis | Oligocene | | Czech Republic | A fern, a species of Polystichum |
| Proodontosoria | Gen. et sp. nov | Valid | Li et al. | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis. |
| Proterocladus antiquus | Sp. nov | Valid | Tang et al. | Mesoproterozoic circa 1 Ga | Nanfen | China | An early siphonocladalean chlorophyte |
| Psilatriletes cozzuolii | Sp. nov | Valid | Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito | Miocene | Solimões Basin | Brazil | Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021. |
| Psilochara monevaensis | Sp. nov | Valid | Sanjuan & Soulié-Märsche | Middle Miocene | | Spain | A charophyte. |
| Qianshouia | Gen. et sp. nov | Valid | Huang et al. | Late Devonian | Wutong Formation | China | A plant of uncertain phylogenetic placement, possibly a lycopsid or a sphenopsid. Genus includes new species Q. mira. |
| Scolecopteris minuta | Sp. nov | In press | Wan et al. | Early Permian | Taiyuan Formation | China | A fern belonging to the group Marattiales. |
| Sigillaria pfefferkornii | Sp. nov | In press | D'Antonio, Boyce & Wang | Permian (Asselian) | | China | |
| Sigillaria wudensis | Sp. nov | In press | D'Antonio, Boyce & Wang | Permian (Asselian) | | China | |
| Sphaerochara miocenica | Sp. nov | Valid | Sanjuan & Soulié-Märsche | Miocene | | Lebanon  Spain | A charophyte. |
| Thyrsopteris cretacea | Sp. nov | Valid | Li et al. | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A species of Thyrsopteris |
| Tomiostrobus sinensis | Sp. nov | Valid | Feng in Feng et al. | Early Triassic (Induan) | Kayitou Formation | China | A member of the family Isoetaceae. |
| Tumidopteris astra | Sp. nov | Valid | Naugolnykh | Permian (Roadian) | Pechora coal basin | Russia | A fern belonging to the family Gleicheniaceae. |
| Ufadendron elongatum | Sp. nov | Valid | Tang et al. | Late Permian | Linxi | China | A lycopsid belonging to the family Tomiodendraceae |
| Uzhurodendron | Gen. et sp. nov | Valid | Mosseichik & Filimonov | Carboniferous (Tournaisian) | Bystrianskaya | Russia ( Krasnoyarsk Krai) | A member of Lycopodiopsida. Genus includes new species U. asiaticum. |
| Zeilleria fosteri | Sp. nov | Valid | Thomas et al. | Carboniferous (Bashkirian) | | United Kingdom | A fern |

## General research

* A study on the evolutionary history of green plants is published by Nie et al. (2020).
* Description of new fossil material of Yurtusia uniformis from the Cambrian Yanjiahe Formation (China) and a study on the phylogenetic relationships and possible life cycle of this organism is published by Shang et al. (2020), who consider Y. uniformis to be a likely green microalga.
* A study on the phylogenetic relationships of extant and fossil complex thalloid liverworts (Marchantiidae) is published by Flores et al. (2020).
* Evidence of development of dichotomous roots in euphyllophytes that were extant during the Devonian and Carboniferous periods is presented by Hetherington, Berry & Dolan (2020), who interpret their findings as indicating that dichotomous root branching evolved in both lycophytes and euphyllophytes.
* An early land plant producing multiple spore size classes is described from the Lower Devonian Campbellton Formation (Canada) by Bonacorsi et al. (2020).
* A study on the impact of the appearance and evolution of herbivorous tetrapods on the evolution of land plants from the Carboniferous to the Early Triassic is published by Brocklehurst, Kammerer & Benson (2020).
* A study on the production of periderm in Late Paleozoic arborescent lycopsids is published by D'Antonio & Boyce (2020), who argue that these lycopsids did not grow from sporelings into large trees through the production of a periderm cylinder, because physiological limitations would have prohibited the production of thick periderm.
* A study on the architecture and development of the Carboniferous arborescent lycopsid Paralycopodites is published by DiMichele & Bateman (2020).
* New information on the anatomy of Weichselia reticulata is presented by Blanco-Moreno, Decombeix & Prestianni (2020).
* A study on the phylogenetic placement of the extinct fern genus Coniopteris is published by Li et al. (2020).
* New information on the morphology of Paleoazolla patagonica is presented by Benedetti et al. (2020), who evaluate the implications of this taxon for the knowledge of the evolution of water ferns.
* A study aiming to determine which ferns were most likely to be the producers of Cyathidites spores from earliest Paleocene plant localities across western North America, and were most likely to be among the first plants in western North America to thrive in the immediate aftermath of the Cretaceous–Paleogene extinction event, is published by Berry (2020).
* A study on the morphology and development of Genomosperma, and on its implications for the knowledge of the evolutionary origins of seed development, is published by Meade, Plackett & Hilton (2020).
* A pollen organ resembling seed fern pollen organs Dictyothalamus and Melissiotheca is described from the Lopingian Umm Irna Formation (Jordan) by Zavialova et al. (2020), who interpret this finding as evidence of persistence of lyginopterid seed ferns until the late Permian.
* Evidence of increasing atmospheric CO$_2$ concentration at the onset of the end-Triassic extinction event, based on data from fossil leaves of the seed fern Lepidopteris ottonis from southern Sweden, is presented by Slodownik, Vajda & Steinthorsdottir (2020), who confirm L. ottonis as a valid proxy for pCO$_2$ reconstructions.
* A study on the anatomy of the seed cone scales of Krassilovia mongolica is published by Herrera et al. (2020), who argue that K. mongolica and Podozamites harrisii are the seed cones and leaves of the same extinct plant, and name a new family Krassiloviaceae within the order Voltziales.
* A study on the microscopic wood anatomy of a fossil tree trunk of Agathoxylon arizonicum with the characteristic external features of a fire scar from the Upper Triassic Chinle Formation (Petrified Forest National Park, Arizona, United States) is published by Byers et al. (2020), who evaluate the implications of this specimen for the knowledge of the evolution of fire-adapted plant traits.
* A putative bamboo "Chusquea" oxyphylla from the early Eocene Laguna del Hunco biota (Argentina) is reinterpreted as a conifer by Wilf (2020), who transfers this species to the genus Retrophyllum.
* A study on evolutionary history of conifers as indicated by fossil and molecular data, aiming to determine whether the rise of angiosperms drove the decline of conifers and other gymnosperms, is published by Condamine et al. (2020).
* Presence of secretory tissues is reported in extinct flowers from the Cretaceous amber from Myanmar and Cenozoic Dominican amber (including specimens preserved while in the process of emitting compounds) by Poinar & Poinar (2020).
* Fossil pollen of flowering plants is reported from the Aptian and Albian of Australia by Korasidis & Wagstaff (2020), who evaluate the implications of their findings for the knowledge of the timing of the appearance and diversification of the flowering plants in the high-latitude southern basins of Australia.
* A study on the morphology of palm and palm-like pollen from the Eocene Yaw Formation (Myanmar), and on the implications of these fossils for the knowledge of distribution and diversity of Eocene palms across the globe, is published by Huang et al. (2020).
* Fossils fruits of Illigera eocenica, representing the second fossil occurrence of Illigera worldwide and the first in Asia, are described from the Eocene Niubao Formation (central Tibetan Plateau) by Wang et al. (2020), who evaluate the implications of this finding for the knowledge of the climate in the central Tibetan Plateau during the early middle Eocene, and for the knowledge of the floristic links between Asia and North America during the Paleogene.
* A study on the morphology and phylogenetic relationships of Montsechia vidalii is published by Gomez et al. (2020).
* Eocene leaves of members of the family Urticaceae with stinging trichomes are described from the Okanogan Highlands (British Columbia, Canada) by DeVore et al. (2020).
* A revision of the fossil record of the family Nothofagaceae from South America is published by Pujana et al. (2020).
* A study on the extinction of plants from south polar terrestrial ecosystems during the Permian–Triassic extinction event and on their recovery after this extinction event, based on data from the Sydney Basin (Australia), is published by Mays et al. (2020).
* A study on the impact of ecological disturbances around the Permian–Triassic boundary (from the Wuchiapingian to Ladinian) on land plant communities is published by Nowak, Vérard & Kustatscher (2020).
* A study on the age of the Paleogene Kanaka Creek fossil flora (Huntingdon Formation; British Columbia, Canada) and on its implications for reconstructions of the contemporaneous paleoclimate and paleoenvironment is published by Mathewes, Greenwood & Love (2020).
* Evidence from Eocene plant fossils from the Bangong-Nujiang suture indicating that the Tibetan Plateau area hosted a diverse subtropical ecosystem approximately 47 million years ago and that this area was both low and humid at the time is presented by Su et al. (2020), who also report that the composition of this flora is similar to Early-Middle Eocene floras in both North America and Europe, but shows little affinity to Eocene floras from the Indian Plate.
* A study aiming to estimate leaf dry mass per area in fossil plants from 22 western North American sites spanning the Eocene–Oligocene transition is published online by Butrim & Royer (2020), who evaluate the implications of their findings for the knowledge of the impact of the environmental changes occurring during the Eocene–Oligocene transition on leaf-economic strategies of plants.
* A study on the Neogene paleobotanical record and climate in the northernmost part of the Central Andean Plateau, based on data from the Descanso Formation (Peru), is published by Martínez et al. (2020), who report the earliest evidence of a puna-like ecosystem in the Pliocene and a montane ecosystem without modern analogs in the Miocene.
* Fossil fruits (mericarps) of the neoendemic Apiaceae Melanoselinum (≡ Daucus) decipiens were reported from the lacustrine and fluvial sediments of Porto da Cruz, Madeira, dated 1.3 Ma, by Góis-Marques et al. 2020. This paper not only reports the oldest Daucus s.l. fossil known to date but also the first fossil evidence of a plant with insular woodiness (see Island gigantism).
* The leaf fossil Mesodescolea plicata from the Early Cretaceous of Patagonia, first interpreted as a cycad with affinities with extant Stangeria, is reinterpreted as an angiosperm leaf with affinities with Austrobaileyales or Chloranthales by Coiro et al. 2020, with implications for the evolution of leaf shape in the early radiation of the angiosperms.
* A study on the phylogenetic relationships of 10 Cretaceous flower taxa (Chloranthistemon endressii, Dakotanthus cordiformis, Kajanthus lusitanicus, Mauldinia mirabilis, Microvictoria svitkoana, Paleoclusia chevalieri, Paradinandra suecica, Spanomera mauldiniensis, Tylerianthus crossmanensis and Virginianthus calycanthoides) is published by Schönenberger et al. (2020).