# List of the prehistoric life of Pennsylvania

This list of the prehistoric life of Pennsylvania contains the various prehistoric life-forms whose fossilized remains have been reported from within the US state of Pennsylvania.

## Precambrian

The Paleobiology Database records no known occurrences of Precambrian fossils in Pennsylvania.

## Paleozoic
### Selected Paleozoic taxa of Pennsylvania

* †Acidaspis
* †Agnostus
* †Alethopteris
* †Amphiscapha
* †Aphthoroblattina
* †Archaeopteris
** †Archaeopteris halliana
** †Archaeopteris macilenta
* †Archimylacris
* †Asaphiscus
* †Athyris
** †Athyris spiriferoides
* †Atrypa
** †Atrypa reticularis
* †Aviculopecten
** †Aviculopecten occidentalis
* †Barinophyton
* †Bathyuriscus
* †Bellerophon
* †Bembexia
* †Beyrichoceratoides
* †Blattoidea
* †Bonneterrina
* Life restoration of the Late Devonian placoderm fish Bothriolepis †Bothriolepis
* †Byronia
* †Callixylon
* †Camarotoechia
* †Camptostroma
* †Cavusgnathus
* †Ceramopora
* †Ceratopsis
* †Chancelloria
* †Chancia
* †Chonetes
* †Cincinnetina
** †Cincinnetina multisecta
* †Cleiothyridina
* †Climacograptus
* Coccosteus cuspidatus restoration, shown attacking multiple Palaeospondylus' †Coccosteus
* †Coelacanthus
* †Composita
** †Composita subtilita
* †Conotheca
* †Coosella
* †Coosia
* †Cordaites
* †Cornulites
** †Cornulites flexuosus
* †Craniops
* †Crepicephalus
* †Ctenerpeton – type locality for genus
* †Ctenopterus
* †Cyclonema
* Fossilized foliage from the Carboniferous seed fern Cyclopteris †Cyclopteris
* †Cypricardinia
** †Cypricardinia indenta
* †Cyrtospirifer
* †Dawsonia
* †Deckera
* †Densignathus – type locality for genus
* †Diplocaulus – or unidentified comparable form
* †Diploceraspis
** †Diploceraspis burkei
* †Diplograptus
* Life restoration of the Permian synapsid (mammal precursor) Edaphosaurus †Edaphosaurus
* †Edmondia
* †Eldredgeops
** †Eldredgeops rana
* †Elita
* †Elrathia
* †Elrathina
* †Endolobus
* †Eoagnostus
* †Erettopterus
* †Eryops
* †Esmeraldina
* †Euomphalus
* Life restoration of the Devonian lobe-finned fish Eusthenodon†Eusthenodon
** E. bourdoni – type locality for species
* Fossil of the Late Ordovician-Permian tabulate coral Favosites †Favosites
** †Favosites niagarensis
* †Fenestella
* †Ganorhynchus
* †Genevievella
* †Glaukerpeton – type locality for genus
* †Glyptocrinus
* †Gnathodus
* †Gravicalymene
* †Greenops
* †Hallipterus
* †Hardieopterus
* †Hazelia
* †Helcionella
* †Hexactinellid
* †Hindeodus
* †Holonema
* †Holopea
* †Holoptychius
* †Holoptychus
* †Hughmilleria
* †Hymenocaris
* Life restoration of the Late Devonian four-limbed animal precursor Hynerpeton †Hynerpeton – type locality for genus
** †Hynerpeton bassetti – type locality for species
* Life restoration of the Late Devonian lobe-finned fish Hyneria†Hyneria – type locality for genus
** †Hyneria lindae – type locality for species
* †Hyolithellus
* †Hyolithes
* †Hyolithid
* †Indiana – tentative report
* †Iocrinus
* †Irvingella
* †Isodectes
* Fossil of the Middle-Late Ordovician giant trilobite Isotelus. †Isotelus
** †Isotelus gigas
* †Kiaeropterus
* †Kingstonia
* †Kladognathus
* †Kootenia
* †Kutorgina
* †Lancastria
* †Langlieria
** †L. radiatus – type locality for species
** †L. smalingi – type locality for species
* †Lepidostrobus
* †Leptomitus
* †Lenisicaris
** Appendage of the Cambrian radiodont Lenisicaris†L. pennsylvanica – type locality for species
* Lingula
* †Lingulella
* †Llanoaspidella – type locality for genus
* †Llanoaspis – tentative report
* †Lonchocephalus
* Life restoration of the Permian snake-like amphibian Lysorophus showing speculative egg-coiling behavior †Lysorophus
* †Margaretia
* †Martinia
* †Megalograptus – tentative report
* †Megamolgophis
* †Metacoceras
* †Michelinoceras
* †Micromitra
* †Mimagoniatites – tentative report
* †Mucrospirifer
** †Mucrospirifer mucronatus
* †Nanahughmilleria
* †Naticopsis
* Fossilized shell of the Carboniferous-Permian brachiopod Neospirifer †Neospirifer
** †Neospirifer cameratus
* †Neuropteris
* †Nisusia
* Nucula
* †Ogygopsis
* †Olenellus
* †Olenoides
** †Olenoides serratus
* †Orthacanthus
* †Orthoceras
* †Orthotheca
* †Oryctocephalus
* Restoration †Osteolepis
* †Oulodus
* †Ozarkodina
* †Paedeumias
* †Pagetia
* †Paladin
* †Panenka
* †Parahughmilleria
* †Paraspirifer
* †Paterina
* †Pelagiella
* †Pemphigaspis
* †Pentremites
* †Peronopsis
* †Petalodus
* Fossil of the Devonian trilobite Phacops rana †Phacops
* Pinna
* †Platyceras
* †Pleurodictyum
* †Proetus
* †Protobarinophyton
* †Protocaris
* †Protospongia – tentative report
* †Pugnax
* †Quasillites
* †Rhinocarcinosoma
* †Rhodea
* †Ribeiria
* †Ruedemannipterus
* †Salterella
* Fossilized fin of the Devonian lobe-finned fish Sauripterus †Sauripterus
** †Sauripterus taylori
* †Schaefferia (Paleozoic plant) – or unidentified comparable form
* †Schizodiscus
* †Sciadophyton
* †Selkirkia
* †Sidneyia
* †Sigillaria
* †Solenochilus
* †Sowerbyella
* †Spathognathodus
* †Spirifer
* †Spyroceras
* †Strophomena
* †Taeniocrada
* †Tentaculites
* Diagram of the branches of the Middle-Late Devonian vascular plant Tetraxylopteris †Tetraxylopteris
* †Titusvillia – type locality for genus
* †Triarthrus
* †Tricrepicephalus
* †Trimerus
* †Tubulella
* †Tuzoia
* †Urasterella
* †Vinella
* †Wanneria
* †Wilkingia
* †Worthenia
* †Yochelcionella

## Mesozoic

* †Anchisauripus
** †Anchisauripus gwyneddensis – type locality for species
** †Anchisauripus poolei – type locality for species
** †Anchisauripus sillimani
* Fossil (right) of the Early Jurassic dinosaur footprint ichnogenus Anomoepus †Anomoepus
** †Anomoepus gracillimus
* †Apatopus
* †Atreipus
** †Atreipus milfordensis
* †Batrachopus
** †Batrachopus gracilis
* †Belodon
** †Belodon priscus
* †Brachychirotherium
* †Brontozoum
** †Brontozoum sillimanium
* †Calamops – type locality for genus
** †Calamops paludosus – type locality for species
* †Chirotherium
** †Chirotherium lulli
** †Chirotherium parvum
* Fossil of the Late Triassic shark Carinacanthus†Carinacanthus – type locality for genus
** †Carinacanthus jepseni – type locality for species
* †Cionichthys
* †Clepsysaurus – type locality for genus
** †Clepsysaurus pennsylvanicus – type locality for species
** †Clepsysaurus veatleianus
* †Dinophyton
* Fossil of the Late Triassic coelacanth Diplurus†Diplurus
** †Diplurus longicaudatus
* †Dipterus
** †Dipterus redfieldi
* †Eubrontes
** †Eubrontes giganteus
* †Grallator
** †Grallator cuneatus
** †Grallator tenuis
* †Gwyneddichnium – type locality for genus
** †Gwyneddichnium majore – type locality for species
* †Gwyneddosaurus – type locality for genus
** †Gwyneddosaurus erici – type locality for species
* †Koskinonodon
** †Koskinonodon perfectus
* Fossilized skeleton of the Middle Triassic amphibian Mastodonsaurus †Mastodonsaurus
** †Mastodonsaurus durus – type locality for species
* †Orthodactylus
** †Orthodactylus howelli – type locality for species
* †Otozoum – tentative report
** †Otozoum minus
* †Pagiophyllum
* †Palaeoctonus
** †Palaeoctonus appalachianus
* †Palaeosaurus
** †Palaeosaurus fraserianus – type locality for species
* †Platypterna
** †Platypterna lockatong – type locality for species
* †Pterodactylus
** †Pterodactylus longispinis – type locality for species
* †Revueltosaurus
* †Rhynchosauroides
** †Rhynchosauroides brunswickii
** †Rhynchosauroides hyperbates
* Life restoration of the Late Triassic phytosaur Rutiodon †Rutiodon
** †Rutiodon carolinensis
* †Scoyenia
* †Sphodrosaurus – type locality for genus
** †Sphodrosaurus pennsylvanicus – type locality for species
* †Suchoprion
** †Suchoprion aulacodus
** †Suchoprion cyphodon
* †Synorichthys
** †S. cf. stewarti
* †Turseodus

## Cenozoic

* Acidota
** †Acidota crenata
** †Acidota quadrata
* Illustration of an Aegialia dung beetle Aegialia
* Agonum
** †Agonum affine
** †Agonum consimile
** †Agonum gratiosum
** †Agonum quinquepunctatum
** †Agonum simile
* Amara
* Apalone
** †Apalone mutica – tentative report
* Aphodius
** †Aphodius precursor – type locality for species
* Apion
* Restoration of an Arctodus, or short-faced bear, with a human to scale †Arctodus
** †Arctodus pristinus – type locality for species
* Arpedium
** †Arpedium cribratum
* †Ateuchus – tentative report
* Bembidion
** †Bembidion anguliferum group informal
** †Bembidion fortestriatum
** †Bembidion frontale
** †Bembidion incertum group informal
** †Bembidion morulum
** †Bembidion mutatum
** †Bembidion pseudocautum
** †Bembidion versicolor – or unidentified comparable form
* Blarina
** A living Blarina brevicauda, or northern short-tailed shrew †Blarina brevicauda – type locality for species
** †Blarina carolinensis
* Bledius
* Blethisa
** †Blethisa catenaria
* Boreaphilus
** †Boreaphilus henningianus
* †Brachyprotoma
** †Brachyprotoma obtusata – type locality for species
* Bufo
** †Bufo americanus
* Canis
** †Canis armbrusteri
** Modern mounted skeleton of Canis lupus, the grey wolf, to scale with a fossilized skeleton of the Pleistocene wolf Canis dirus, or dire wolf †Canis dirus
* Carabus
** †Carabus maeander
* †Cariacus
** †Cariacus laevicornis – type locality for species
* Carphoborus
** †Carphoborus andersoni
* Castor
** †Castor canadensis
* †Castoroides
** †Castoroides ohioensis
* Catops
* †Cervalces
* Chlaenius
** †Chlaenius punctulatus – type locality for species
* Colinus
** †Colinus virginianus
* Coluber
** †Coluber acuminatus – type locality for species
** †Coluber constrictor
* Colymbetes
* A living Crocuta crocuta, or spotted hyena Crocuta
** †Crocuta inexpectatus – type locality for species
* Crotalus
** †Crotalus horridus
* Curimopsis
** †Curimopsis moosilauke – or unidentified comparable form
* †Cychrus
** †Cychrus minor – type locality for species
** †Cychrus wheatleyi – type locality for species
* Cymindis
** †Cymindis aurora – type locality for species
** †Cymindis cribricollis
* Cytilus
** †Cytilus alternatus
* A preserved Diacheila ground beetle Diacheila
** †Diacheila arctica
* Dicaelus
** †Dicaelus alutaceus – type locality for species
* Donacia
* Dyschirius
** †Dyschirius integer group informal
** †Dyschirius melancholicus
* Elaphe
** †Elaphe vulpina
* Elaphrus
** †Elaphrus americanus
** †Elaphrus clairvillei
** †Elaphrus lapponicus
* Emydoidea
** A living Emys blandingii, or Blanding's turtle †Emydoidea blandingii
* Enochrus
* Equus
** †Equus complicatus
** †Equus pectinatus
* Erethizon
** †Erethizon dorsatum
* Eucnecosum
** †Eucnecosum brunnescens
* Glyptemys
** †Glyptemys insculpta
* Gulo
** A living Gulo gulo, or wolverine †Gulo gulo
* Gymnusa
* Harpalus
* Helophorus
** †Helophorus sempervarians
** †Helophorus tuberculatus
* †Hesperotestudo
** †Hesperotestudo percrassa – type locality for species
* Heterodon
** †Heterodon platyrhinos
* Holoboreaphilus
** †Holoboreaphilus nordenskioldi
* Homo
** †Homo sapiens
* Hydraena
* Hydroporus
* Hyla
** A living Pseudacris crucifer (formerly Hyla crucifer), or spring peeper †Hyla crucifer
* Lampropeltis
** †Lampropeltis triangulum
* Lasiopodomys – tentative report
* Lesteva
** †Lesteva pallipes
* Lontra
** †Lontra canadensis
* Loricera
** †Loricera pilicornis
* Lynx
** †Lynx calcaratus – type locality for species
* †Mammut
** †Mammut americanum
* †Mammuthus
** Life restorations of a Mammut americanum, or American mastodon (right), and a Mammuthus primigenius, or wooly mammoth (left) †Mammuthus primigenius
* Mantura
* Marmota
** †Marmota monax – or unidentified comparable form
* †Megalonyx
** †Megalonyx wheatleyi
* Meleagris
** †Meleagris gallopavo
* Mephitis
** †Mephitis mephitis
* Microedus
** †Microedus austinianus
* Microtus
** †Microtus guildayi – or unidentified comparable form
* Restoration of the Pliocene-Pleistocene Miracinonyx, or American cheetah †Miracinonyx
** †Miracinonyx inexpectatus – or unidentified comparable form
* Mustela
* Mycetoporus
* †Mylohyus
** †Mylohyus fossilis
* Myotis
* Napaeozapus
** †Napaeozapus insignis
* Natrix
* A living Neofiber, or round-tailed muskrat Neofiber
** †Neofiber diluvianus – type locality for species
* Nerodia
** †Nerodia sipedon
* Nitidotachinus
** †Nitidotachinus tachyporoides
* Notaris
** †Notaris aethiops
* Notiophilus
* Ochthebius
* Odocoileus
** A living Odocoileus virginianus, or white-tailed deer †Odocoileus virginianus
* Olophrum
** †Olophrum boreale
** †Olophrum consimile
** †Olophrum latum
** †Olophrum rotundicolle
* Ondatra
** †Ondatra idahoensis – type locality for species
* Opheodrys
** †Opheodrys vernalis
* †Osmotherium – type locality for genus
** †Osmotherium spelaeum – type locality for species
* Oxytelus
* Panthera
** †Panthera onca
* Fossilized skeleton of the Pliocene-Pleistocene ground sloth Paramylodon †Paramylodon
** †Paramylodon harlani – tentative report
* Parascalops
** †Parascalops breweri
* Patrobus
** †Patrobus foveocollis
** †Patrobus septentrionis
** †Patrobus stygicus
* Pekania
** †Pekania diluviana
* Pelenomus
* Peromyscus
** †Peromyscus leucopus – tentative report
* †Phanaeus
** †Phanaeus antiquus – type locality for species
* Philonthus
* Phloeotribus
** †Phloeotribus piceae
* Pitymys
** †Pitymys cumberlandensis – or unidentified comparable form
* Pityophthorus
* Restoration of a herd of alarmed Miocene-Pleistocene peccaries of the genus Platygonus. Charles R. Knight (1922). †Platygonus
** †Platygonus vetus – type locality for species
* Platynus
** †Platynus hypolithos
* Podabrus
* Polygraphus
** †Polygraphus rufipennis
* Pterostichus
** †Pterostichus laevigatus – type locality for species
** †Pterostichus patruelis
** †Pterostichus punctatissimus
* Pycnoglypta
** †Pycnoglypta aptera
** †Pycnoglypta lurida
* Quedius
* †Rana
** †Rana pipiens – or unidentified comparable form
* Scalopus
* Simplocaria
* Life restoration of the Pleistocene-Holocene saber-tooth cat Smilodon †Smilodon
** †Smilodon gracilis – type locality for species
* Sorex
* Sphaeroderus
** †Sphaeroderus nitidicollis – or unidentified comparable form
* Stenus
* Stereocerus
** †Stereocerus haematopus
* Sylvilagus
** †Sylvilagus floridanus
* Synaptomys
** †Synaptomys cooperi
* Tachinus
** †Tachinus luridus
* Tamias
** †Tamias striatus – or unidentified comparable form
* A living Tapirus, or tapir Tapirus
** †Tapirus haysii
** †Tapirus veroensis
* Taxidea
** †Taxidea taxus
* Terrapene
** †Terrapene carolina
* Thamnophis
* Thanatophilus
* Thomomys
** †Thomomys potomacensis
* Trechus
** †Trechus crassiscapus
* †Uncia
** †Uncia mercerii – type locality for species
* Urocyon
** A living Urocyon cinereoargenteus, or gray fox †Urocyon cinereoargenteus
* Ursus
** †Ursus americanus
* Vulpes
* Zapus
** †Zapus hudsonius