# List of the Paleozoic life of Pennsylvania

This list of the Paleozoic life of Pennsylvania contains the various prehistoric life-forms whose fossilized remains have been reported from within the US state of Pennsylvania and are between 538.8 and 252.17 million years of age.

## A

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* H
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* K
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* M
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* U
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* X
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* †Acanthopecten
** †Acanthopecten carboniferous
* †Acidaspis
** †Acidaspis cincinnatiensis
* †Acrothele
** †Acrothele decipiens
** †Acrothele yorkensis
* †Acrotreta
* †Actinomylacris
** †Actinomylacris similis – type locality for species
* †Actinopteria
** †Actinopteria boydi
** †Actinopteria boydii
** †Actinopteria decussata
* †Adelphobolbina
** †Adelphobolbina medialis
* †Adiantites
** †Adiantites mirabilis
** †Adiantites spectabilis – type locality for species
** †Adiantites ungeri – type locality for species
* †Adiatites
** †Adiatites cyclopteroides – type locality for species
** †Adiatites spectabilis – type locality for species
* †Aechmina
* †Agassizocrinus
* †Ageleodus
** †Ageleodus pectinatus
* †Aglaoglypta
** †Aglaoglypta maera
** †Aglaoglypta nactoides
* †Agnostus
* †Alcicornopteris
** †Alcicornopteris altoonensis – type locality for species
** †Alcicornopteris anthracitica – type locality for species
* Fossilized fronds of the Carboniferous-Early Cretaceous seed fern Alethopteris †Alethopteris
** †Alethopteris densinervosa
** †Alethopteris ingbertensis
** †Alethopteris lonchitifolia
** †Alethopteris missouriensis
** †Alethopteris serli
** †Alethopteris westphalensis
* †Allanella
** †Allanella tullius
* †Alokistocare
** †Alokistocare aoris – type locality for species
** †Alokistocare nasutum – type locality for species
* †Ambocoelia
** †Ambocoelia gregaria
** †Ambocoelia umbonata
* †Amboneura – type locality for genus
** †Amboneura klosei – type locality for species
* †Americaspis
* †Amphiscapha
** †Amphiscapha catilloides
* †Amphissites
* †Anastomopora
* †Ancyrospora
* †Aneurospora
** †Aneurospora greggsii
* †Anisotrypa
* †Ankoura
** †Ankoura apicalis
** †Ankoura orbiculata – or unidentified comparable form
** †Ankoura sublettensis
* Restorative model of the Cambrian arthropod Anomalocaris †Anomalocaris
** †Anomalocaris lineata
* †Anomomylacris – type locality for genus
** †Anomomylacris cubitalis – type locality for species
* †Anoplea
** †Anoplea nucleata
* †Anoplotheca
* †Anthracospirifer
* †Aparchites
** †Aparchites subrotunda
* †Aphthoroblattina
** †Aphthoroblattina fascigera – type locality for species
* †Aphyllum
** †Aphyllum fascicularium
* †Archaeopteridium
** †Archaeopteridium bellasylviana – type locality for species
* †Archaeopteris
** †Archaeopteris halliana
** †Archaeopteris latifolia
** Life restoration of the Late Devonian-Carboniferous tree Archaeopteris macilenta with insets detailing its anatomy †Archaeopteris macilenta
** †Archaeopteris minor
** †Archaeopteris roemeriana – or unidentified comparable form
** †Archaeopteris rogersi
* †Archaeosperma
** †Archaeosperma arnoldii
* †Archaeotriletes
** †Archaeotriletes senticosus
* †Archeopteris
** †Archeopteris latifolia
* †Archimylacris
** †Archimylacris parallelum
* †Archinacella
* †Armathyraster
** †Armathyraster paradoxis
* †Arthrophycus
* †Articulatae
* Fossil of the Cambrian trilobite Asaphiscus †Asaphiscus
** †Asaphiscus glaber
* †Asemoblatta – type locality for genus
** †Asemoblatta pennsylvanica – type locality for species
* †Astartella
** †Astartella vera
* †Atalotaenia – type locality for genus
** †Atalotaenia adela – type locality for species
* †Ateleocystites
** †Ateleocystites huxleyi
* †Athyris
** †Athyris angelica
** †Athyris spiriferiodes
** †Athyris spiriferoides
* †Atimoblatta – type locality for genus
** †Atimoblatta curvipennis – type locality for species
** †Atimoblatta flexuosa – type locality for species
** †Atimoblatta reducta – type locality for species
* †Atlanticocoelia
** †Atlanticocoelia acutiplicata
* †Atrypa
** †Atrypa recticularis
** Fossilized shell of the Silurian-Late Devonian brachiopod Atrypa reticularis †Atrypa reticularis
* †Aulocystis
* †Auroraspora
** †Auroraspora hyalina
** †Auroraspora macra
** †Auroraspora solisorta
** †Auroraspora torquata
* †Aviculopecten
** †Aviculopecten bellus
** †Aviculopecten equilatera
** †Aviculopecten occidentalis

## B

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* B
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* I
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* M
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* †Bactrites
* †Barbclabornia
** †Barbclabornia luedersensis
* †Barinophyton
** †Barinophyton citrulliforme
** †Barinophyton obscurum
* †Bathyuriscus
* †Batostoma
* Fossilized shell of the Silurian-Early Triassic mollusc Bellerophon †Bellerophon
** †Bellerophon clarki
** †Bellerophon nactus
* †Bembexia
** †Bembexia laevis
* Berenicea
* †Beyrichoceratoides
** †Beyrichoceratoides lunatus
* †Bisporangiostrobus
** †Bisporangiostrobus harrisii
* †Blainia
** †Blainia buttsi – type locality for species
* †Blairella
** †Blairella triangularis – type locality for species
* †Blattoidea
** †Blattoidea schucherti – type locality for species
* †Blountia
** †Blountia prolifica – type locality for species
** †Blountia waddlensis
** †Blountia warriorensis – type locality for species
* †Bollia
** †Bollia hama
** †Bollia hindei
** †Bollia planofibra
** †Bollia spinomuralis
** †Bollia ungula
* †Bolllia
** †Bolllia ungula
* †Bonneterrina
** †Bonneterrina aspinosa – type locality for species
* †Bonnia
** †Bonnia bubaris
** †Bonnia capito
** †Bonnia senecta – tentative report
* †Bonniella
** †Bonniella yorkensis
* Fossilized bony armor of the Late Devonian placoderm fish Bothriolepis Life restoration of the Late Devonian placoderm fish Bothriolepis †Bothriolepis
** †Bothriolepis nitida
** †Bothriolepis nitidus
* †Brachycycloceras
** †Brachycycloceras curtum
* †Brachymylacris
** †Brachymylacris bassleri – type locality for species
** †Brachymylacris elongata – type locality for species
** †Brachymylacris rotundata – type locality for species
* †Bucanopsis
* †Bufina
** †Bufina bicornuta
* †Burnetiella
** †Burnetiella ectypa
** †Burnetiella urania
* †Buttisia
** †Buttisia drabensis
* †Buttsia
** †Buttsia completa
** †Buttsia drabensis
* †Bynumia
** †Bynumia terrenda
* †Bynumina
** †Bynumina terrenda
** †Bynumina type locality for species – informal
* †Bynuminia
** †Bynuminia terrenda
* †Byronia
* †Byssonychia – tentative report
** †Byssonychia vera – or unidentified comparable form
* Bythocypris

## C

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* †Calathiops
** †Calathiops pottsvillensis – type locality for species
* †Calathospongia
** †Calathospongia carceralis
** †Calathospongia carlli
** †Calathospongia tidioutensis
* †Callixylon
* †Caloneurella – type locality for genus
** †Caloneurella carbonaria – type locality for species
* †Calvibembexia
** †Calvibembexia sulcomarginata
* †Camaraspis
** †Camaraspis convexa
* †Camarotoechia
** †Camarotoechia constricta
** †Camarotoechia prolifica
** †Camarotoechia saxatilis
* †Camerella
* Life restoration of the Cambrian echinoderm Camptostroma †Camptostroma
** †Camptostroma roddyi
* †Cardiocarpon
** †Cardiocarpon phillipsi – type locality for species
* †Cariniferella
** †Cariniferella carinata
** †Cariniferella tioga
* †Carydium
** †Carydium bellastriatum
* †Cavellina
** †Cavellina planoprocliva
* †Cavusgnathus
* †Centronella
* †Ceramopora
** †Ceramopora incondita
* †Ceratopsis
** †Ceratopsis chambersi
* †Ceriocrinus
* †Chalepomylacris – type locality for genus
** †Chalepomylacris pulchra – type locality for species
* †Chancelloria
** †Chancelloria yorkensis
* †Chancia
* †Cheilocephalus
** †Cheilocephalus quadratus – type locality for species
* †Chonetes
** †Chonetes buttsi
** †Chonetes hemisphericus
* †Chonetinella
** †Chonetinella verneuilana
* †Cincinnaticrinus
* Assemblage of fossilized shells of the Ordovician brachiopod Cincinnetina †Cincinnetina
** †Cincinnetina multisecta
* †Cladopora
** †Cladopora seriata
* †Cleiothyridina
* †Cliffia
** †Cliffia lataegenae
** †Cliffia latagenae
* †Climacograptus
** †Climacograptus typicalis
* †Cobaloblatta – type locality for genus
** †Cobaloblatta simulans – type locality for species
* Coccosteus cuspidatus restoration, shown attacking multiple Palaeospondylus' †Coccosteus
** †Coccosteus macromus – type locality for species
* †Coelacanthus
* †Coleolus
* †Colpodexylon
** †Colpodexylon deatsii
* †Colpomya
** †Colpomya faba
* †Comanchia
** †Comanchia amplooculata
* †Composita
** †Composita subtilita
* †Conaspis
* †Conotheca
** †Conotheca australiensis – or unidentified comparable form
* †Conularia
** †Conularia trentonensis – tentative report
* †Convolutispora
** †Convolutispora opressa – or unidentified comparable form
* †Coosella
** †Coosella convexa – type locality for species
** †Coosella vulgaris – type locality for species
* †Coosia
** †Coosia pustulata – type locality for species
* †Corbulispora
** †Corbulispora cancellata
* Fossilized foliage of the Carboniferous-Permian conifer relative Cordaites †Cordaites
* †Cornellites
** †Cornellites flabella
* †Cornulites
** †Cornulites flexuosus
** †Cornulites progressus – or unidentified comparable form
* †Corticospongia – type locality for genus
** †Corticospongia bradfordensis – type locality for species
* †Cranaena
* †Craniops
** †Craniops hamiltoniae
** †Craniops subtruncata
* †Crenistriella
** †Crenistriella crenistria
* †Crepicephalus
* †Crurithyris
** †Crurithyris planoconvexa
* †Cryptolithus
** †Cryptolithus bellulus
** †Cryptolithus recurvus
* †Cryptozoon
** †Cryptozoon undulatum
* †Ctenerpeton – type locality for genus
** †Ctenerpeton remex – type locality for species
* †Ctenoloculina
** †Ctenoloculina cicatricosa
* Fossils and life restoration of the Silurian eurypterid ("sea scorpion") Ctenopterus †Ctenopterus
** †Ctenopterus cestrotus
* †Cuneamya
* †Cupularostrum
** †Cupularostrum congregata
** †Cupularostrum contracta
** †Cupularostrum exima
** †Cupularostrum orbicularis
* †Cyclonema
* †Cyclopteris
* †Cymatospira
** †Cymatospira montfortianus
* †Cypricardella
** †Cypricardella bellastriata
** †Cypricardella bellistriata
** †Cypricardella tenuistriata
* †Cypricardina
** †Cypricardina indenta
* †Cypricardinia
** †Cypricardinia indenta
* †Cyrtina
** †Cyrtina hamiltonensis
* Fossilized shells of the Middle-Late Devonian brachiopod Cyrtospirifer †Cyrtospirifer
** †Cyrtospirifer disjunctus
* †Cyrtospora
** †Cyrtospora cristifera – or unidentified comparable form
* †Cystihalysites

## D

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* F
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* H
* I
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* †Dawsonia
** †Dawsonia parkeri
* †Deadwoodia
** †Deadwoodia duris
* †Dechenella
* †Deckera
** †Deckera completa
* †Dekayella
* †Dellea
** †Dellea glabellamersa
** †Dellea rogersi – type locality for species
** †Dellea saratogensis
* †Delthyris
* †Deltodus
** †Deltodus angularis
* †Densignathus – type locality for genus
** †Densignathus rowei – type locality for species
* †Derbyia
** †Derbyia crassa
** †Derbyia robusta
* †Devonochonetes
** †Devonochonetes coronatus
** †Devonochonetes scitulus
** †Devonochonetes syrtalis
* †Diacanthaspis
* †Diaphanospora
* †Diaphragmus
* †Dictyomylacris
* †Dictyotomaria
** †Dictyotomaria capillaria
* †Dictyotriletes
** †Dictyotriletes nefandus
* †Dieconeurites – type locality for genus
** †Dieconeurites rigidus – type locality for species
* †Diplocaulus – or unidentified comparable form
* Life restoration of the Permian amphibian Diploceraspis †Diploceraspis
** †Diploceraspis burkei
* †Diplograptus
** †Diplograptus nexus
* †Ditomopyge
** †Ditomopyge scitulus
* †Dizygopleura
* †Dodecaactinella
** †Dodecaactinella spicules
* †Dokimocephalus – tentative report
** †Dokimocephalus intermedius
* †Douvillina
** †Douvillina cayuta
** †Douvillina inaequistriata
* †Drabia
** †Drabia acroccipita
** †Drabia acrocclpita
** †Drabia acrocclptia
** †Drabia menusa
** †Drabia typica
* †Dunbarella
** †Dunbarella striata

## E

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* †Echinoconchus
** †Echinoconchus punctatus
* †Ectenocrinus
** †Ectenocrinus simplex
* Life restoration of the Permian synapsid (mammal precursor) Edaphosaurus †Edaphosaurus
** †Edaphosaurus raymondi – type locality for species
* †Edmondia
* †Eldredgeops
** †Eldredgeops rana
* †Elita
** †Elita fimbriata
* †Elrathia
** †Elrathia grazierensis – type locality for species
* †Elrathina
* †Elvinia
** †Elvinia roemeri
* †Embolophyllum
** †Embolophyllum clarki
** †Embolophyllum coralliferum
** †Embolophyllum schucherti
* †Emmanuella
** †Emmanuella subumbona
* †Emphanisporites
** †Emphanisporites annulatus – or unidentified comparable form
** †Emphanisporites pantagiatus – or unidentified comparable form
** †Emphanisporites rotatus
* †Endolobus
* †Endothyra
* †Endothyranella
** †Endothyranella nitida – or unidentified comparable form
** †Endothyranella stormi – or unidentified comparable form
* †Entellophylloides
** †Entellophylloides inequalils
* †Eoagnostus
** †Eoagnostus roddyi
* †Eoobolus
** †Eoobolus priscus
* †Eoorthis
** †Eoorthis indianola
* †Eoserpeton
** †Eoserpeton tenuicorne – type locality for species
* †Eothele
** †Eothele tubulus
* Life restoration of the Silurian-Early Devonian eurypterid ("sea scorpion") Erettopterus †Erettopterus
** †Erettopterus globiceps
* †Eryops
** †Eryops avinoffi
* †Eshelmania
* †Esmeraldina
** †Esmeraldina macer
* †Euglyphella
** †Euglyphella numismoides
* †Eukloedenella
* †Eumetria
* †Euomphalus
* †Eupachycrinus
* †Euphemites
** †Euphemites carbonarius

## F

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* †Fabreciella – type locality for genus
** †Fabreciella pennsylvanica – type locality for species
* Fossil of the Late Ordovician-Permian tabulate coral Favosites †Favosites
** †Favosites niagarensis
* †Fedexia
* †Fenestella
** †Fenestella emaciata
** †Fenestella sinuosa
* †Fimbrispirifer
* †Finkelnburgia
** †Finkelnburgia bridgei – or unidentified comparable form
* †Fistuliporella
** †Fistuliporella minima
* †Fletcherina
** †Fletcherina incognita – or unidentified comparable form
* †Floweria
** †Floweria chemungensis

## G

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* †Ganorhynchus
** †Ganorhynchus oblongus – type locality for species
* †Geminospora
* Fossil of the Cambrian trilobite Genevievella †Genevievella
** †Genevievella campbellina – type locality for species
** †Genevievella pennstatensis – type locality for species
** †Genevievella plesiochielus – type locality for species
* †Gigantocharinus – type locality for genus
** †Gigantocharinus szatmaryi – type locality for species
* †Girtya
** †Girtya pennslvanica
** †Girtya pennsylvanica – type locality for species
* †Glabrocingulum
** †Glabrocingulum grayvillense
* †Glaphyraspis
** †Glaphyraspis ovata
* †Glaukerpeton – type locality for genus
** †Glaukerpeton avinoffi – type locality for species
* †Glyptocrinus
* †Glyptodesma
** †Glyptodesma erectum – tentative report
* †Gnathodus
* †Goniomylacris – type locality for genus
** †Goniomylacris pauper – type locality for species
* †Goniophora
** †Goniophora hamiltonensis
* †Grandispora
** †Grandispora cornuta
** †Grandispora upensis
* Fossils of the Middle Ordovician-Early Devonian trilobite Gravicalymene †Gravicalymene
* †Greenops
** †Greenops boothi

## H

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* †Hadrotreta
* Life restoration of the Late Devonian eurypterid ("sea scorpion") Hallipterus †Hallipterus
** †Hallipterus lacoanus
* †Hapaloptera – type locality for genus
** †Hapaloptera gracilis – type locality for species
* †Hardieopterus
** †Hardieopterus myops
* †Hazelia
** †Hazelia walcotti
* †Healdia
* †Helcionella
* †Hemimylacrella
** †Hemimylacrella mammothi – type locality for species
* †Hemimylacris
** †Hemimylacris ramificata – type locality for species
* †Herrmannina
* †Heslerodus
** †Heslerodus divergens
* †Hexactinellid
** †Hexactinellid spicules
* †Hibbardella
* †Hibbardia
** †Hibbardia lacrimosa
* †Hindeodus
* †Holcacephalus
** †Holcacephalus tunda
* †Hollinella
** †Hollinella sella – or unidentified comparable form
** †Hollinella tricollina
* Life restoration of the Middle-Late Devonian placoderm fish Holonema †Holonema
** †Holonema horrida – type locality for species
** †Holonema rugosa
* †Holopea
* †Holoptychius
** †Holoptychius americanus
** †Holoptychius filosus – type locality for species
** †Holoptychius giganteus
* †Holoptychus
** †Holoptychus flabellatus – type locality for species
** †Holoptychus latus – type locality for species
** †Holoptychus serrulatus – type locality for species
* †Homacanthus
** †Homacanthus acinaciformis – type locality for species
* †Homilodonta
** †Homilodonta filistriata
* †Hostimella
** †Hostimella crispa
* †Housia
** †Housia vacuna
* †Hudsonaster
** †Hudsonaster lancolatus – or unidentified comparable form
* †Hughmilleria
** †Hughmilleria shawangunk
* †Hymenocaris
* †Hymenozonotriletes
** †Hymenozonotriletes explanatus
** †Hymenozonotriletes granulatus
* Life restoration of the Late Devonian four-limbed animal precursor Hynerpeton †Hynerpeton – type locality for genus
** †Hynerpeton bassetti – type locality for species
* †Hyolithellus
** †Hyolithellus micans
* †Hyolithes
** †Hyolithes wanneri
* †Hyolithid
* †Hypselentoma
** †Hypselentoma perhumerosa
* †Hystricosporites
** †Hystricosporites multifurcatus
** †Hystricosporites porcatus
* †Hystriculina
** †Hystriculina wabashensis

## I

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* †Ianthinopsis
* †Iddingsia
** †Iddingsia anatina
* †Idiomylacris – type locality for genus
** †Idiomylacris gracilis – type locality for species
* †Imlerella
** †Imlerella praecipita – type locality for species
* †Indiana – tentative report
* †Inflatia
* †Inocaulis
* †Iocrinus
** †Iocrinus subcrassus
* †Irvingella
** †Irvingella major
* Life restoration of the Permian amphibian Isodectes †Isodectes
** †Isodectes obtusus – type locality for species
* †Isotelus
** †Isotelus gigas
* †Ithycephalus

## K

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* †Katabuporhynchus
** †Katabuporhynchus mesacostalis
* Fossilized heads of the Silurian eurypterid ("sea scorpion") Kiaeropterus (specimens 1-8) †Kiaeropterus
** †Kiaeropterus otisius
* †Kindbladia
** †Kindbladia wichitaensis
* †Kingstonia
** †Kingstonia apion
** †Kingstonia ara
** †Kingstonia cheilusis – type locality for species
** †Kingstonia trapezoidia – type locality for species
** †Kingstonia ulrichi – type locality for species
** †Kingstonia walcotti
* †Kinklidoblatta
** †Kinklidoblatta lesquereuxi – type locality for species
* †Kinocera – tentative report
* †Kinzercystis
** †Kinzercystis durhami
* †Kinzeria – type locality for genus
** †Kinzeria crinita – type locality for species
* †Kladognathus
* †Kloedinia
* †Knoxisporites
** †Knoxisporites dedaleus
* †Kochiella
* Fossil of the Cambrian trilobite Kootenia †Kootenia
** †Kootenia marcoui
* †Kraeuselisporites
* †Kutorgina
** †Kutorgina cingulata
** †Kutorgina edsoni
* †Kyphocephalus

## L

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Lacoea
** †Lacoea seriata – type locality for species
* †Lagenospermum
** †Lagenospermum imparirameum
* †Lambdagnathus
* †Lancastria
* †Lecanospira
** †Lecanospira compacta
** †Lecanospira salteri – or unidentified comparable form
* †Leiorhynchus
* †Leiozonotriletes
* †Lepdidodendropsis
* †Leperditia
* †Lepidocystis
** †Lepidocystis wanneri
* †Lepidodendropsis
** †Lepidodendropsis scobiniformis
** †Lepidodendropsis stigillariodes
** †Lepidodendropsis stigillaroides
** †Lepidodendropsis vandergrachti
* †Lepidostrobus
** †Lepidostrobus gallowayi
* †Leptaena
** †Leptaena rhomboidalis
** †Leptaena richmondensis – or unidentified comparable form
* †Leptodesma
** †Leptodesma cadmus
** †Leptodesma extenuatum
** †Leptodesma hector
** †Leptodesma laevis
** †Leptodesma lichas
** †Leptodesma maclurii
** †Leptodesma mortoni
** †Leptodesma naviforme
** †Leptodesma nereus
** †Leptodesma nitida
** †Leptodesma orus
** †Leptodesma patulum
** †Leptodesma pelops
** †Leptodesma phaon
** †Leptodesma sayi
** †Leptodesma sociale
** †Leptodesma spinerigum
* †Leptomitus
* †Lichenocrinus
* Lingula
** †Lingula carbonaria
** †Lingula delia
** †Lingula ligea
* Illustration of a fossilized shell of the Cambrian-Late Ordovician brachiopod Lingulella †Lingulella
** †Lingulella marcia – or unidentified comparable form
* †Lingulepis
** †Lingulepis acuminata
* †Linoproductus
** †Linoproductus cora
* †Liospira
** †Liospira americana
* †Llanoaspidella – type locality for genus
** †Llanoaspidella bigenusia – type locality for species
** †Llanoaspidella warriorsmarkensis – type locality for species
* †Llanoaspis – tentative report
* †Lonchocephalus
** †Lonchocephalus planus – type locality for species
** †Lonchocephalus swartzi – type locality for species
** †Lonchocephalus waddlei – type locality for species
* †Longispina
** †Longispina mucronatus
* †Lophophyllidium
** †Lophophyllidium profundum
* †Lophospira
* †Lophozonotriletes
** †Lophozonotriletes excisus
** †Lophozonotriletes tuberosus
* †Loxonema
* Life restoration of the Permian snake-like amphibian Lysorophus showing speculative egg-coiling behavior †Lysorophus
** †Lysorophus dunkardensis

## M

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Macluritella – tentative report
** †Macluritella multiseptarius
* †Maeneceras
** †Maeneceras milleri
* †Margaretia
* †Mariopteris
* †Martinia
* †Mediospirifer
** †Mediospirifer audaculus
** †Mediospirifer belliplicata
** †Mediospirifer bellitropis
** †Mediospirifer dromgoldi
* †Megakozlowskiella
** †Megakozlowskiella sculptilis
* Life restoration of the Late Ordovician eurypterid ("sea scorpion") Megalograptus †Megalograptus – tentative report
* †Megamolgophis
** †Megamolgophis agostinii
* †Megastrophia
* †Menomonia
** †Menomonia calymenoides
* †Mesitoblatta
** †Mesitoblatta emersoni – type locality for species
* †Mesotaphraspis
* †Metacoceras
** †Metacoceras cornutum
* †Metaxyblatta – type locality for genus
** †Metaxyblatta hadroptera – type locality for species
* †Metropator – type locality for genus
** †Metropator pusillus – type locality for species
* †Michelinoceras
* †Micromitra
** †Micromitra alabamensis
* †Mimagoniatites – tentative report
* †Modiella
** †Modiella pygmaea
* †Modiolopsis
* †Modiomorpha
** †Modiomorpha concentrica
* †Modocia
** †Modocia benorensis – type locality for species
* †Monotrypa
** †Monotrypa benjamini
* Fossilized shell of the Devonian brachiopod Mucrospirifer †Mucrospirifer
** †Mucrospirifer mucronatus
* †Mylacridae
** †Mylacridae carbonina
* †Mylacris
** †Mylacris angusta – type locality for species
** †Mylacris anthracophila – type locality for species
** †Mylacris dubia – type locality for species
** †Mylacris lucifuga – type locality for species
** †Mylacris ovalis – type locality for species
** †Mylacris pennsylvanica – type locality for species
** †Mylacris truncatula – type locality for species

## N

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Nalivkinella
** †Nalivkinella echoense
* †Nanahughmilleria
** †Nanahughmilleria clarkei
* †Naticonema
* Fossilized shell of the Early Devonian – Triassic sea snail Naticopsis †Naticopsis
* †Necymylacris
** †Necymylacris heros – type locality for species
** †Necymylacris lacoana – type locality for species
* †Neoblairella – type locality for genus
** †Neoblairella crassimarginata – type locality for species
* †Neochonetes
** †Neochonetes granulifer
* †Neomylacris
** †Neomylacris major – type locality for species
** †Neomylacris pulla – type locality for species
* †Neoprionodus
* †Neoraistrickia
* †Neosimplicius – type locality for genus
** †Neosimplicius medialis – type locality for species
* †Neospirifer
** †Neospirifer cameratus
* †Neriopteris
** †Neriopteris lanceolata
* †Nervostrophia
** †Nervostrophia nervosa
* Fronds of the Carboniferous seed fern Neuropteris †Neuropteris
* †Nisusia
** †Nisusia festinata
* Nucula
** †Nucula bellistriata
** †Nucula lirata
* Nuculana
* †Nuculites
** †Nuculites oblongatus
** †Nuculites planulatus
** †Nuculites triqueter
* †Nuculopsis
** †Nuculopsis girtyi

## O

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Octonaria
** †Octonaria bifurcata
** †Octonaria dicristata
** †Octonaria quadricostata
* †Ogygopsis
** †Ogygopsis klotzi
* †Olenellus
** †Olenellus alius
** †Olenellus similaris
** †Olenellus wanneri
* Fossil of the Cambrian trilobite Olenoides †Olenoides
** †Olenoides buttsi – type locality for species
** †Olenoides pennsylvanicus – type locality for species
** †Olenoides serratus
* †Olethroblatta
** †Olethroblatta americana – type locality for species
* †Orbiculoidea
** †Orbiculoidea tullia
* †Orsadesmus – type locality for genus
** †Orsadesmus rubecollus – type locality for species
* Life restoration of the Carboniferous-Permian freshwater shark Orthacanthus †Orthacanthus
** †Orthacanthus compressus
** †Orthacanthus platypternus
** †Orthacanthus texensis
* †Orthoceras
* †Orthograptus
** †Orthograptus quadrimucronatus
* †Orthomylacris
** †Orthomylacris alutacea – type locality for species
** †Orthomylacris analis – type locality for species
** †Orthomylacris cordiformis – type locality for species
** †Orthomylacris mansfieldi – type locality for species
** †Orthomylacris pauperata – type locality for species
** †Orthomylacris pennsylvaniae – type locality for species
** †Orthomylacris pittstoniana – type locality for species
** †Orthomylacris pluteus – type locality for species
* †Orthonota
** †Orthonota parvula – tentative report
** †Orthonota undulata
* †Orthonychia
** †Orthonychia parva
* †Orthotetes
* †Orthotheca
** †Orthotheca shriveri – type locality for species
* †Oryctocephalus
* Restoration †Osteolepis
* †Oulodus
* †Ovatia
* †Oxynoblatta
** †Oxynoblatta vicina – type locality for species
* †Ozarkodina

## P

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Paedeumias
** †Paedeumias eboracense
** †Paedeumias glabrum
** †Paedeumias yorkense
* Fossil of the Cambrian trilobite Pagetia †Pagetia
* †Paladin
* †Palaeoneilo
** †Palaeoneilo constricta
** †Palaeoneilo fecunda
** †Palaeoneilo muta
** †Palaeoneilo oweni
** †Palaeoneilo plana
** †Palaeoneilo tenuistriata
* †Palaeotherates
** †Palaeotherates pennsylvanicus – type locality for species
* †Palaeozygopleura
** †Palaeozygopleura delphicola
* †Panenka
* †Parabolbina
** †Parabolbina cardocornella
** †Parabolbina obliqua
* †Parabolinella
** †Parabolinella occidentalis
* †Parabolinoides
* †Paracyclas
** †Paracyclas lirata
* †Parahaplophlebium
** †Parahaplophlebium longipenne – type locality for species
* †Parahughmilleria
** †Parahughmilleria maria
* †Parajuresania
** †Parajuresania nebrascensis
* †Parallelodon
** †Parallelodon obsoletus
* †Paranacystites
* Assemblage of the Early-Middle Devonian brachiopod Paraspirifer †Paraspirifer
** †Paraspirifer acuminatus
* †Parvohallopora
** †Parvohallopora dalei – or unidentified comparable form
* †Paterina
** †Paterina bella
* †Pelagiella
* †Pemphigaspis
** †Pemphigaspis bullata
** †Pemphigaspis intermedia – type locality for species
** †Pemphigaspis matutina
* †Penniretepora
* Fossilized theca of the Carboniferous blastoid echinoderm ("sea bud") Pentremites †Pentremites
* †Periomella
** †Periomella roddyi
** †Periomella yorkensis
* †Peronopsis
* †Petalodus
** †Petalodus ohioensis
* †Petrocrania
* Fossil of the Devonian trilobite Phacops rana †Phacops
* †Pharkidonotus
** †Pharkidonotus percarinatus
* †Phestia
* †Philhedra
** †Philhedra laelia – or unidentified comparable form
* †Phoberoblatta – type locality for genus
** †Phoberoblatta grandis – type locality for species
** †Phoberoblatta reticulata – type locality for species
* †Pholidops
* †Pholidostrophia
** †Pholidostrophia pennsylvanica
* †Phragmactis
* †Phricodothyris
* †Phthinomylacris
** †Phthinomylacris medialis – type locality for species
** †Phthinomylacris pauper – type locality for species
* †Phthonia
** †Phthonia sectifrons
* †Phyloblatta
** †Phyloblatta indecisa – type locality for species
** †Phyloblatta prior – type locality for species
* †Pinctus
** †Pinctus latus
* Pinna
* †Pionoceras
** †Pionoceras swartzi
* †Plagioblatta
** †Plagioblatta campbelli – type locality for species
** †Plagioblatta cockerelli – type locality for species
* †Plagioglypta
** †Plagioglypta meekiana
* †Planoendothyra – tentative report
** †Planoendothyra associata
* †Planutenia
** †Planutenia flectata
* †Plasiochiton – type locality for genus
** †Plasiochiton curiosus – type locality for species
* Fossilized shell of the Silurian-Early Triassic sea snail Platyceras †Platyceras
** †Platyceras euomphaloides
** †Platyceras lineatum
* †Platylichas
* †Plectorthis
** †Plectorthis plicatella
* †Pleurodictyum
** †Pleurodictyum styloporum
* †Polidevcia
** †Polidevcia bellistriata
* †Poliella
** †Poliella bala
* †Polypora
* †Ponderodictya
** †Ponderodictya favulosa
** †Ponderodictya punctulifera
* †Poroblattina
** †Poroblattina parvula – type locality for species
* †Praewaagenoconcha
** †Praewaagenoconcha speciosa
* †Primitiella
** †Primitiella unicornis
* †Prismopora
* †Productella
** †Productella rectispina
* †Productus
* Restoration of the Silurian trilobite Proetus †Proetus
** †Proetus haldemani – tentative report
** †Proetus parviusculus
* †Prolepidodendron
** †Prolepidodendron breviinternodium
* †Promopalaeaster
** †Promopalaeaster bellulus
** †Promopalaeaster pricei
* †Promytilus
* †Protobarinophyton
** †Protobarinophyton pennsylvanicum
* Illustration of a fossil of the Cambrian crustacean Protocaris †Protocaris
* †Protocycloceras
** †Protocycloceras rushense
* †Protoleptostrophia
** †Protoleptostrophia perplana
* †Protomya – or unidentified comparable form
* †Protospongia – tentative report
* †Prozacanthoides
* †Pseudoatrypa
** †Pseudoatrypa devoniana
* †Pseudolingula
** †Pseudolingula rectilateralis
* †Pseudopaolia
** †Pseudopaolia lacoana – type locality for species
* †Pseudopolyernus
** †Pseudopolyernus laminarum – type locality for species
* †Pseudosaratogia
** †Pseudosaratogia bulbosa
** †Pseudosaratogia lata – tentative report
** †Pseudosaratogia magna
* †Pseudotealliocaris
** †Pseudotealliocaris palincsari
* †Pteridomylacris – type locality for genus
** †Pteridomylacris paradoxa – type locality for species
* †Pterinea
* †Pterinopecten
** †Pterinopecten hermes
** †Pterinopecten princeps
* †Pterochaenia
* †Pterygometopus
** †Pterygometopus achates
* †Ptilograptus
* †Ptilomylacris – type locality for genus
** †Ptilomylacris medialis – type locality for species
* †Ptilopora
* †Ptomatis
* †Ptychoglyptus
** †Ptychoglyptus virginensis – or unidentified comparable form
* †Ptychoparella
** †Ptychoparella lancastra
* †Ptychopteria
** †Ptychopteria chemungensis
** †Ptychopteria flabella
** †Ptychopteria salamanca
* †Ptylopora
* †Pugnax
** †Pugnax pugnus
* †Pustula
** †Pustula pertenuis
* †Pustulatia
** †Pustulatia pustulosa

## Q

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Quasillites

## R

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Rafinesquina
** †Rafinesquina alternata
* †Retichonetes
** †Retichonetes marylandicus
** †Retichonetes vicinus
* †Retispira
** †Retispira marcouianus
* †Retusotriletes
** †Retusotriletes phillipsii
* †Rhacophyton
* †Rhacopteris
* †Rhinidictya
* Fossil of the Silurian eurypterid ("sea scorpion") Rhinocarcinosoma †Rhinocarcinosoma
** †Rhinocarcinosoma cicerops
* †Rhipidomella
** †Rhipidomella leucosia
** †Rhipidomella penelope
** †Rhipidomella vanuxemi
* †Rhodea
** †Rhodea alleghanensis – type locality for species
** †Rhodea tionestana – type locality for species
** †Rhodea vespertina – type locality for species
* †Rhombopora
* †Rhytimya
** †Rhytimya radiata
* †Ribeiria
** †Ribeiria parva – or unidentified comparable form
* †Roddyia
* †Rodea (genus)
** †Rodea vespertina – type locality for species
* †Ruedemannipterus
** †Ruedemannipterus stylonuroides
* †Rugospora
** †Rugospora flexuosa

## S

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* Illustration of fossilized shells of the mysterious Cambrian organism Salterella †Salterella
** †Salterella acervulosa
** †Salterella conulata
** †Salterella mccullochi
** †Salterella pulchella
* †Sanguinolites
* †Sapphicorhynchus
** †Sapphicorhynchus sappho
* †Saratogia
** †Saratogia type locality for species – informal
* Fossilized fin of the Devonian lobe-finned fish Sauripterus †Sauripterus
** †Sauripterus taylori
* †Schaefferia (Paleozoic plant) – or unidentified comparable form
* †Schizoblatta
** †Schizoblatta pennsylvanica – type locality for species
* †Schizodiscus
** †Schizodiscus capsa
* †Schizodus
** †Schizodus cuneatus
* †Schizophoria
** †Schizophoria amanaensis
** †Schizophoria impressa
** †Schizophoria striatula
* †Schuchertella
** †Schuchertella parva
** †Schuchertella variabilis
* †Schuchertia
** †Schuchertia laxata
* †Sciadophyton
* †Secarisporites
* †Selkirkia
* †Septopora
* †Serracaris
* †Shansiella
** †Shansiella carbonaria
* Fossils of the Cambrian arthropod Sidneyia †Sidneyia
* †Sigillaria
* †Sinochonetes
** †Sinochonetes lepidus
* †Sinuites
** †Sinuites cancellatus
* †Soleniscus
** †Soleniscus primogenia
** †Soleniscus typicus
* †Solenochilus
** †Solenochilus collectum
* †Solenopleurella
** †Solenopleurella elatifrons – type locality for species
** †Solenopleurella transversa – type locality for species
* †Solutotherates
** †Solutotherates analis – type locality for species
* †Sowerbyella
** †Sowerbyella plicatellus
** †Sowerbyella rugosa
* †Spathognathodus
* †Spelaeotriletes
** †Spelaeotriletes crustalus
** †Spelaeotriletes resolutus
* †Sphenomylacris
** †Sphenomylacris singularis – type locality for species
* †Spinatrypa
** †Spinatrypa hystrix
** †Spinatrypa spinosa
* †Spinocyrtia
** †Spinocyrtia girtyi
** †Spinocyrtia granulosa
** †Spinocyrtia macbridei
* †Spinulicosta
** †Spinulicosta spinulicosta
* Fossilized shell of the Late Ordovician-Late Triassic brachiopod Spirifer †Spirifer
** †Spirifer disjunctus
** †Spirifer varicosa
* †Spongophylloides
* †Spyroceras
** †Spyroceras nuntium
* †Stauromatidium
** †Stauromatidium marylandicum
* †Stemtonoceras
** †Stemtonoceras elongatum
* †Stenomylacris – type locality for genus
** †Stenomylacris elegans – type locality for species
* †Stereolasma
* †Stittaspis
** †Stittaspis loria – type locality for species
* †Straparollus
* †Striacoceras
** †Striacoceras typum – tentative report
* †Strobeus
** †Strobeus paludinaeformis
* †Strophalosia
** †Strophalosia truncata
* †Stropheodonta
** †Stropheodonta demissa
* Fossilized shell of the Ordovician-Silurian brachiopod Strophomena †Strophomena
* †Styliolina
** †Styliolina bellistriata
** †Styliolina fissurella
* †Subrensselandia
** †Subrensselandia claypolii
* †Sulcocephalus
** †Sulcocephalus candidus
** †Sulcocephalus granulosus
* †Sulcoretepora
** †Sulcoretepora incisurata
** †Sulcoretepora sinuosa – or unidentified comparable form
* †Swataria – type locality for genus
** †Swataria derstleri – type locality for species
* †Syntrophina
* †Syscioblatta
** †Syscioblatta allegheniensis – type locality for species
** †Syscioblatta pennsylvanica – type locality for species

## T

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Tabulipora
* †Taeniaster
** †Taeniaster maximus
* †Taenicephalus
** †Taenicephalus shumardi
* Fossils of the Devonian plant stem morphogenus Taeniocrada †Taeniocrada
* †Taeniopora
** †Taeniopora exigua
* †Taonurus
* †Tellinopsis
* †Temnocheilus
** †Temnocheilus crassus
* †Tentaculites
** †Tentaculites attenuatus
* †Tetradella
** †Tetradella multipustulosa
* †Tetraxylopteris
* †Thlipsura
** †Thlipsura ultimata
* †Thlipsurella
** †Thlipsurella semipunctata
* †Titanodictya
** †Titanodictya jucunda – type locality for species
* †Titusvillia – type locality for genus
** †Titusvillia drakei – type locality for species
* †Trepospira
** †Trepospira illinoisensis
* Restoration showing the top (left) and underside of the Late Ordovician trilobite Triarthrus †Triarthrus
** †Triarthrus becki
* †Tricrepicephalus
** †Tricrepicephalus arcuatus – type locality for species
* †Trimerus
** †Trimerus dekayi
* †Triphyllopteris
** †Triphyllopteris biloba – type locality for species
** †Triphyllopteris latilobata – type locality for species
** †Triphyllopteris latilobatus
** †Triphyllopteris lescuriana
* †Trocophyllum
** †Trocophyllum breviinternodium
* †Tropidoleptus
** †Tropidoleptus carinatus
* †Truncalosia
** †Truncalosia truncata
* †Truncatiramus – now regarded as a jr. synonym of Erettopterus
* †Tryplasma
* †Tubulella
* †Tuzoia
* †Tylothyris
** †Tylothyris mesacostalis
** †Tylothyris pauliformis

## U

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Ulrichia
** †Ulrichia elongata
** †Ulrichia spinifera
* Fossil of the Ordovician-Devonian sea star Urasterella †Urasterella
** †Urasterella pulchella

## V

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Vallatisporites
** †Vallatisporites pusilites
* †Vernonaspis
* †Verrucosisporites
** †Verrucosisporites nitidus
** †Verrucosisporites scurrus
* †Vinella

## W

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* Fossil of the Cambrian trilobite Wanneria †Wanneria
** †Wanneria walcottana
* †Warrenella
** †Warrenella laevis
* †Welleraspis
** †Welleraspis newfoundlandensis – or unidentified comparable form
* †Whidbornella
** †Whidbornella lachrymosa
* †Wilkingia
** †Wilkingia terminale
* †Worthenia
** †Worthenia tabulata

## X

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Xenoblatta
** †Xenoblatta pennsylvanica – type locality for species
* †Xenocheilos
** †Xenocheilos spineum

## Y

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* Life restoration of the probable Cambrian mollusc Yochelcionella †Yochelcionella
** †Yochelcionella americana – type locality for species
** †Yochelcionella chinensis
** †Yochelcionella greenlandica
* †Yorkia
** †Yorkia wanneri

## Z

* A
* B
* C
* D
* E
* F
* G
* H
* I
* J
* K
* L
* M
* N
* O
* P
* Q
* R
* S
* T
* U
* V
* W
* X
* Y
* Z
* †Zaphrenthis
* †Zelophyllia
* †Zelophyllum
* †Zygospira