# Joculator

Joculator is a genus of minute sea snails, marine gastropod molluscs in the family Cerithiopsidae. This genus was described by Hedley in 1909.

## Species

Species in the genus Joculator include:
* Joculator abbreviatus (Thiele, 1930)
* Joculator acuminatus Cecalupo & Perugia, 2012
* Joculator aeneus Cecalupo & Perugia, 2017
* Joculator albatus Cecalupo & Perugia, 2023
* Joculator albordina Laseron, 1956
* Joculator albulus (Thiele, 1930)
* Joculator alligatus Cecalupo & Perugia, 2012
* Joculator anconae Cecalupo & Perugia, 2017
* Joculator angustus Cecalupo & Perugia, 2017
* Joculator antonioi Cecalupo & Perugia, 2012
* Joculator aoreensis Cecalupo & Perugia, 2013
* Joculator arduinii Cecalupo & Perugia, 2012
* Joculator arenaceus Cecalupo & Perugia, 2013
* Joculator ariadnae Cecalupo & Perugia, 2018
* Joculator ater Cecalupo & Perugia, 2012
* Joculator aterrufus Cecalupo & Perugia, 2019
* Joculator autumnus Cecalupo & Perugia, 2016
* Joculator bangkaensis Cecalupo & Perugia, 2019
* Joculator baraggiai Cecalupo & Perugia, 2013
* Joculator barteri Cecalupo & Perugia, 2018
* Joculator barteri Cecalupo & Perugia, 2018
* Joculator bauquisi Cecalupo & Perugia, 2017
* Joculator berberiani Cecalupo & Perugia, 2018
* Joculator bicinctus Cecalupo & Perugia, 2012
* Joculator boguszae Cecalupo & Perugia, 2018
* Joculator bourgeoisae Cecalupo & Perugia, 2013
* Joculator boguszae Cecalupo & Perugia, 2018
* Joculator bouteti Cecalupo & Perugia, 2014
* Joculator brabantae Cecalupo & Perugia, 2013
* Joculator breviculus Cecalupo & Perugia, 2017
* Joculator brevis Cecalupo & Perugia, 2012
* Joculator brisseti Cecalupo & Perugia, 2021
* Joculator brucei (Melvill & Standen, 1912)
* Joculator brusonii Cecalupo & Perugia, 2013
* Joculator caliginosus Cecalupo & Perugia, 2012
* Joculator caloi Cecalupo & Perugia, 2013
* Joculator carpatinus Cecalupo & Perugia, 2012
* Joculator castaneus Cecalupo & Perugia, 2013
* Joculator cereus Cecalupo & Perugia, 2012
* Joculator christiaensi Jay & Drivas, 2002
* Joculator cinctus Cecalupo & Perugia, 2012
* Joculator citrinus Cecalupo & Perugia, 2013
* Joculator coffeus Cecalupo & Perugia, 2013
* Joculator conformis Cecalupo & Perugia, 2013
* Joculator continens Laseron, 1956
* Joculator corneliae Cecalupo & Perugia, 2018
* Joculator cossignanii Cecalupo & Perugia, 2012
* Joculator cristinae Cecalupo & Perugia, 2013
* Joculator czubai Cecalupo & Perugia, 2018
* Joculator dakoi Cecalupo & Perugia, 2018
* † Joculator decapitatus Grant-Mackie & Chapman-Smith, 1971
* Joculator desmieri Cecalupo & Perugia, 2018
* Joculator diversus Cecalupo & Perugia, 2019
* Joculator drivasi Cecalupo & Perugia, 2012
* Joculator dupouxae Cecalupo & Perugia, 2018
* Joculator emidioi Cecalupo & Perugia, 2014
* Joculator erianae Cecalupo & Perugia, 2017
* Joculator eudeli Jay & Drivas, 2002
* Joculator faninozi Cecalupo & Perugia, 2018
* Joculator fedosovi Cecalupo & Perugia, 2018
* Joculator ferrii Cecalupo & Perugia, 2013
* Joculator ferrugineus Cecalupo & Perugia, 2012
* Joculator filum Cecalupo & Perugia, 2017
* Joculator fischeri Jay & Drivas, 2002
* Joculator flindersi Cotton, 1951
* Joculator flavicans Cecalupo & Perugia, 2014
* Joculator frequens Cecalupo & Perugia, 2012
* Joculator fulvaster Cecalupo & Perugia, 2019
* Joculator furvus Cecalupo & Perugia, 2012
* Joculator fuscostriatus Cecalupo & Perugia, 2013
* Joculator fuscus Cecalupo & Perugia, 2012
* Joculator gagulae Cecalupo & Perugia, 2018
* Joculator ganui Cecalupo & Perugia, 2018
* Joculator garianii Cecalupo & Perugia, 2013
* Joculator gemmae Cecalupo & Perugia, 2012
* Joculator giovanolii Cecalupo & Perugia, 2013
* Joculator gorini Cecalupo & Perugia, 2014
* Joculator goubini Cecalupo & Perugia, 2017
* Joculator gracilis Laseron, 1951
* Joculator granatus Kay, 1979
* Joculator granulosus Cecalupo & Perugia, 2013
* Joculator grimalae Cecalupo & Perugia, 2021
* Joculator hadfieldae Cecalupo & Perugia, 2017
* Joculator hasegawai Cecalupo & Perugia, 2019
* Joculator hedleyi Laseron, 1951
* Joculator herosae Cecalupo & Perugia, 2012
* Joculator humilis Cecalupo & Perugia, 2012
* Joculator ianthinus Cecalupo & Perugia, 2013
* Joculator incisus Cecalupo & Perugia, 2012
* Joculator inflatus Cecalupo & Perugia, 2012
* Joculator introspectus Cotton, 1951
* Joculator iopuei Cecalupo & Perugia, 2017
* Joculator itiensis Cecalupo & Perugia, 2014
* Joculator jeffkinchi Cecalupo & Perugia, 2018
* Joculator johnkasui Cecalupo & Perugia, 2018
* Joculator juliuschani Cecalupo & Perugia, 2018
* Joculator kaipuae Cecalupo & Perugia, 2018
* Joculator kanoi Cecalupo & Perugia, 2013
* Joculator kenganziki Cecalupo & Perugia, 2018
* Joculator keratochroma Jay & Drivas, 2002
* Joculator keszlerae Cecalupo & Perugia, 2021
* Joculator koluae Cecalupo & Perugia, 2021
* Joculator kuboi Cecalupo & Perugia, 2019
* Joculator laguncula (Cecalupo & Perugia, 2012)
* Joculator laregnereensis Cecalupo & Perugia, 2017
* Joculator laseroni Jay & Drivas, 2002
* Joculator latrechei Cecalupo & Perugia, 2021
* Joculator lazzarii Cecalupo & Perugia, 2013
* Joculator legallae Cecalupo & Perugia, 2014
* Joculator legoffi Cecalupo & Perugia, 2017
* Joculator lazzarii Cecalupo & Perugia, 2013
* Joculator leguyaderi Cecalupo & Perugia, 2013
* Joculator lividus Cecalupo & Perugia, 2012
* Joculator lokaroensis Cecalupo & Perugia, 2014
* Joculator lozoueti Jay & Drivas, 2002
* Joculator luteolus Cecalupo & Perugia, 2012
* † Joculator maestratii Lozouet, Lesport & Renard, 2001
* Joculator magninensis Cecalupo & Perugia, 2017
* Joculator maloensis Cecalupo & Perugia, 2013
* Joculator manneensis Cecalupo & Perugia, 2018
* Joculator maranii Cecalupo & Perugia, 2014
* Joculator marinae Cecalupo & Perugia, 2014
* Joculator massimilianoi Cecalupo & Perugia, 2012
* Joculator mauberti Cecalupo & Perugia, 2018
* Joculator mbereensis Cecalupo & Perugia, 2017
* Joculator meanii Cecalupo & Perugia, 2014
* Joculator megacephala Jay & Drivas, 2002
* Joculator melania Laseron, 1956
* Joculator melanoraphis Jay & Drivas, 2002
* Joculator micalii Cecalupo & Perugia, 2012
* Joculator michoni Cecalupo & Perugia, 2018
* Joculator minimului Cecalupo & Perugia, 2018
* Joculator minimus Laseron, 1956
* Joculator minor Laseron, 1956
* Joculator minutissimus (Thiele, 1925)
* Joculator minutus Cecalupo & Perugia, 2012
* Joculator mitsuoi Cecalupo & Perugia, 2019
* Joculator modestus Cecalupo & Perugia, 2012
* Joculator morenae Cecalupo & Perugia, 2013
* Joculator morigii Cecalupo & Perugia, 2017
* Joculator murciai Cecalupo & Perugia, 2014
* Joculator mygaki Jay & Drivas, 2002
* Joculator myia Jay & Drivas, 2002
* Joculator nitidus Cecalupo & Perugia, 2012
* Joculator nucleus Cecalupo & Perugia, 2019
* Joculator obscurus Cecalupo & Perugia, 2012
* Joculator obsoletus Cecalupo & Perugia, 2012
* Joculator occultus Cecalupo & Perugia, 2012
* Joculator olivoideus Cecalupo & Perugia, 2018
* Joculator onnaensis Cecalupo & Perugia, 2019
* Joculator opulentus Cecalupo & Perugia, 2013
* Joculator ovatus Laseron, 1956
* Joculator pallidus Cecalupo & Perugia, 2012
* Joculator parvulus Cecalupo & Perugia, 2012
* Joculator pauxillus Cecalupo & Perugia, 2012
* Joculator perforatus Cecalupo & Perugia, 2013
* Joculator perlucidus Cecalupo & Perugia, 2012
* Joculator phtyr Jay & Drivas, 2002
* Joculator pierresamueli Cecalupo & Perugia, 2021
* Joculator pinguis Cecalupo & Perugia, 2012
* Joculator pizzinii Cecalupo & Perugia, 2013
* Joculator priorai Cecalupo & Perugia, 2012
* Joculator problematicus Albano & Steger, 2021
* Joculator prunus Cecalupo & Perugia, 2013
* Joculator psyllos Jay & Drivas, 2002
* Joculator pulvis (Issel, 1869)
* Joculator pumilus Cecalupo & Perugia, 2018
* Joculator pupiformis Cecalupo & Perugia, 2012
* Joculator pygmaeus Cecalupo & Perugia, 2012
* Joculator quaggiottoi Cecalupo & Perugia, 2012
* Joculator ralijanoai Cecalupo & Perugia, 2014
* Joculator ralphmanai Cecalupo & Perugia, 2018
* Joculator rambellii Cecalupo & Perugia, 2018
* Joculator recisus Cecalupo & Perugia, 2012
* Joculator ridiculus (Watson, 1886)
* Joculator rolani Cecalupo & Perugia, 2012
* Joculator rubus Cecalupo & Perugia, 2013
* Joculator sabrinae Cecalupo & Perugia, 2012
* Joculator saguili Cecalupo & Perugia, 2013
* Joculator salvati Jay & Drivas, 2002
* Joculator sarahpezetae Cecalupo & Perugia, 2018
* Joculator savellii Cecalupo & Perugia, 2021
* Joculator sbranai Cecalupo & Perugia, 2021
* Joculator schiaparellii Cecalupo & Perugia, 2017
* Joculator sekensis Cecalupo & Perugia, 2018
* Joculator semiperlucidus Cecalupo & Perugia, 2013
* Joculator simonsili Cecalupo & Perugia, 2018
* Joculator simulans Cecalupo & Perugia, 2012
* Joculator siwisikae Cecalupo & Perugia, 2018
* Joculator steykerae Cecalupo & Perugia, 2013
* Joculator stramineus Cecalupo & Perugia, 2013
* Joculator subconicus Cecalupo & Perugia, 2012
* Joculator subdolus Cecalupo & Perugia, 2012
* Joculator subglobosus Cecalupo & Perugia, 2013
* Joculator sublima Marshall, 1978
* Joculator subula Laseron, 1956
* Joculator succineus Cecalupo & Perugia, 2013
* Joculator sylviakinchae Cecalupo & Perugia, 2018
* Joculator tagliaferroi Cecalupo & Perugia, 2017
* Joculator testaceus Cecalupo & Perugia, 2013
* Joculator testii Cecalupo & Perugia, 2017
* Joculator theresae Cecalupo & Perugia, 2018
* Joculator thielei Jay & Drivas, 2002
* Joculator tieensis Cecalupo & Perugia, 2017
* Joculator tomacula Laseron, 1956
* Joculator tribulationis (Hedley, 1909)
* Joculator tsiriveloi Cecalupo & Perugia, 2014
* Joculator unicolor Cecalupo & Perugia, 2012
* Joculator uveanus (Melvill & Standen, 1896)
* Joculator vandelae Cecalupo & Perugia, 2014
* Joculator variabilis Cecalupo & Perugia, 2012
* Joculator varians Laseron, 1956
* Joculator vassardi Cecalupo & Perugia, 2014
* Joculator vazzanai Cecalupo & Perugia, 2018
* Joculator venustus Cecalupo & Perugia, 2013
* Joculator vignali Jay & Drivas, 2002
* Joculator violaceus Cecalupo & Perugia, 2012
* Joculator voncoseli Cecalupo & Perugia, 2012
* Joculator websterae Cecalupo & Perugia, 2014
* Joculator zahariasi Cecalupo & Perugia, 2021
* Joculator ziliolii Cecalupo & Perugia, 2012
* Joculator zucconi Cecalupo & Perugia, 2021

### Taxa inquirenda

* Joculator aelomitres (Melvill & Standen, 1896)
* Joculator marileutes (Melvill & Standen, 1896)

### Species brought into synonymy

* Joculator albocinctum (Melvill & Standen, 1896): synonym of Costulopsis albocincta (Melvill & Standen, 1896)
* Joculator albula (Thiele, 1930): synonym of Joculator albulus (Thiele, 1930)
* Joculator caelata Powell, 1930: synonym of Synthopsis caelata (Powell, 1930)
* Joculator granata Kay, 1979: synonym of Nanopsis granata (Kay, 1979)
* Joculator introspecta Cotton, 1951: synonym of Joculator introspectus Cotton, 1951
* Joculator iohannae Cecalupo & Perugia, 2012: synonym of Synthopsis iohannae (Cecalupo & Perugia, 2012)
* Joculator minima Laseron, 1956: synonym of Joculator minimus Laseron, 1956
* Joculator minutissima (Thiele, 1925): synonym of Joculator minutissimus (Thiele, 1925)
* Joculator myia Jay & Drivas, 2002: synonym of Nanopsis myia (Jay & Drivas, 2002)
* Joculator nanus Laseron, 1951: synonym of Potenatomus nanus (Laseron, 1951)
* Joculator ovata Laseron, 1956: synonym of Joculator ovatus Laseron, 1956
* Joculator ridicula: synonym of Joculator ridiculus (Watson, 1886)
* Joculator semipicta (Gould, 1861): synonym of Joculator semipictus (Gould, 1861)
* Joculator semipictus (Gould, 1861): synonym of Horologica semipicta (Gould, 1861)
* Joculator skolix Jay & Drivas, 2002: synonym of Costulopsis skolix (Jay & Drivas, 2002) (original combination)
* Joculator turriger (Watson, 1886): synonym of Horologica turrigera (Watson, 1886)
* Joculator turrigera (Watson, 1886): synonym of Joculator turriger (Watson, 1886)
* Joculator uveanum (Melvill & Standen, 1896): synonym of Joculator uveanus (Melvill & Standen, 1896)