# Emarginula

Emarginula is a genus of small keyhole limpets, marine gastropod molluscs in the family Fissurellidae.

## Description

The oval shell is obliquely conical with a backward-pointing recurved apex. There is a deep incision in the anterior margin. A distinct anal fasciole extends upwards from this incision. The surface is latticed, sculptured with radial ribs. There is no septum inside.

### Radula

The radula has a dental formula of [many].1.4.1.4.1.[many], with a wide, rectangular, acuspate rachidian tooth. The various genera have a few dozen rows of teeth.

## Species

According to the World Register of Marine Species (WoRMS), species within the genus Emarginula include:
* Emarginula adamsiana Sowerby in Adams & Sowerby, 1863
* Emarginula adriatica Costa O.G., 1829
* Emarginula agulhasensis Thiele, 1925
* Emarginula alba Watson, 1897
* Emarginula amyda Shikama, 1962
* Emarginula angusta McLean, 1970
* Emarginula annielangleitae Poppe & Tagaro, 2020
* Emarginula apolonia Pacaud, 2015
* † Emarginula auversiensis Deshayes, 1861
* Emarginula bajula Hedley, 1913
* Emarginula balicasagensis Poppe & Tagaro, 2020
* Emarginula bellula A. Adams, 1852
* Emarginula bicancellata Montrouzier, 1860
* Emarginula bonfittoi Smriglio & Mariottini, 2001
* Emarginula boucheti Poppe & Tagaro, 2020
* † Emarginula brebioni Ceulemans, Van Dingenen & Landau, 2016
* Emarginula camilla Melvill & Standen, 1903
* Emarginula candida Adams, 1852
* Emarginula capuloidea G. Nevill & H. Nevill, 1869
* † Emarginula carezi de Laubrière, 1881
* Emarginula choristes Dall, 1925
* Emarginula christiaensi Piani, 1985
* Emarginula circumalbum Poppe & Tagaro, 2020
* † Emarginula clathrata Deshayes, 1824
* † Emarginula clathrataeformis Eichwald, 1830
* Emarginula colipanoae Poppe & Tagaro, 2020
* Emarginula compta Habe, 1953
* Emarginula concinna Adams, 1852
* Emarginula connectens Thiele, 1915
* Emarginula connelli Kilburn, 1978
* Emarginula convexa Hedley, 1907
* † Emarginula costata Lamarck, 1803
* Emarginula costulata Deshayes, 1863
* Emarginula crassa Sowerby, 1812
* Emarginula crassicostata Sowerby in Adams & Sowerby, 1863
* Emarginula cucullata A. Adams, 1852
* Emarginula curvamen Iredale, 1925
* Emarginula curvata Schepman, 1908
* Emarginula cuvieri Audouin, 1827 (species inquirenda)
* Emarginula decorata Deshayes, 1863
* † Emarginula delicatissima Chapman & Gabriel, 1923
* Emarginula delonguevilleae Poppe & Tagaro, 2020
* † Emarginula dennanti Chapman & Gabriel, 1923
* Emarginula devota Thiele, 1915
* Emarginula dictya McLean, 1970
* † Emarginula didactica Ludbrook, 1956
* † Emarginula dilatoria Ludbrook, 1956
* Emarginula dilecta Adams, 1852
* Emarginula divae (van Aartsen & Carrozza, 1995)
* Emarginula dubia Schepman, 1908 (accepted > unreplaced junior homonym, non Emarginula dubia Defrance, 1819)
* Emarginula dujardini Dollfus & Dautzenberg, 1886 †
* † Emarginula espibosensis Lozouet, 1999
* † Emarginula exigua Trautschold, 1866
* Emarginula fenestrella Deshayes, 1863
* Emarginula fissura (Linnaeus, 1758) - slit limpet
* † Emarginula fossorensis Lozouet, 1999
* Emarginula foveolata Schepman, 1908
** Emarginula foveolata fugitai Habe, 1953
* † Emarginula foveolata Gerasimov, 1955 (accepted > unreplaced junior homonym)
* Emarginula fragilis Yokoyama, 1920
* Emarginula fuliginea Adams, 1851
* Emarginula gabensis (Gabriel, 1961)
* Emarginula galericulata A. Adams, 1852
* † Emarginula galeriformis Marwick, 1928
* † Emarginula gigantea Poppe, 2008
* †Emarginula guerangeri A. d'Orbigny, 1843
* Emarginula gustavi Poppe & Tagaro, 2020
* Emarginula hataii Habe, 1953
* Emarginula hawaiiensis Dall, 1895
* † Emarginula haweraensis Powell, 1931
* Emarginula hosoyai Habe, 1953
* Emarginula huzardii (Payraudeau, 1826)
* Emarginula icosisculpta Simone & Cunha, 2014
* Emarginula imaizumi Dall, 1926
* Emarginula imella Dall, 1926
* Emarginula janae Cossignani, 2020
* Emarginula japonica Sowerby in Adams & Sowerby, 1863
* Emarginula jungcheni K.-Y. Lai, 2019
* † Emarginula kaiparica Laws, 1939
* † Emarginula kaiwarensis Bandel, Gründel & P. A. Maxwell, 2000 †
* † Emarginula karuna Das, Bardhan & Lahiri, 1999
* Emarginula kashimaensis Shikama, 1962
* † Emarginula komitica Laws, 1939
* Emarginula koon Kilburn, 1978
* † Emarginula ladowae Eichman, 1955
* Emarginula lata J.R.C. Quoy & J.P. Gaimard, 1834 ((uncertain, status in recent literature not researched)
* Emarginula lentiginosa D. G. Herbert, 2024
* Emarginula liuzzii Poppe & Tagaro, 2020
* Emarginula longifissa Sowerby, 1866
* Emarginula lorenzoi Giusti & Micali, 2019
* Emarginula macclurgi Kilburn, 1978
* Emarginula mactanensis Poppe & Tagaro, 2020
* Emarginula maculata Adams in Adams & Sowerby, 1863
* Emarginula margarita Poppe & Tagaro, 2020
* † Emarginula mariae Cossmann, 1888
* † Emarginula maudensis Chapman & Gabriel, 1923
* Emarginula micans Adams, 1852
* Emarginula montrouzieri Souverbie, 1872
* Emarginula multisquamosa Schepman, 1908
* Emarginula multistriata Jeffreys, 1882
* Emarginula natalensis Barnard, 1963
* Emarginula nodulicostata D. G. Herbert, 2024
* Emarginula nordica Pérez Farfante, 1947 - northern emarginula
* † Emarginula oblonga Sandberger, 1859
* Emarginula obovata Adams, 1852
* Emarginula octaviana Coen, 1939
* Emarginula oppressa Barnard, 1963
* Emarginula paivana (Crosse, 1867)
* Emarginula papilionacea G. Nevill & H. Nevill, 1869
* Emarginula patula Cotton, 1930
* † Emarginula paucicostata Laws, 1936
* Emarginula paucipunctata Schepman, 1908
* † Emarginula pelagica Passy, 1832
* Emarginula philippinensis Poppe & Tagaro, 2020
* Emarginula phrixodes Dall, 1927 - ruffled emarginula
* Emarginula pittensis Marwick, 1928
* Emarginula poppeorum Romani & Crocetta, 2017
* † Emarginula praesicula Lozouet, 1999
* Emarginula puncticulata Adams, 1852
* Emarginula punctulum Piani, 1980
* † Emarginula punctura S. Wood, 1848
* Emarginula pustula Thiele in Küster, 1913
* Emarginula quadrata Poppe & Tagaro, 2020
* † Emarginula radiocostata Kiel & Bandel, 2000
* Emarginula regia Habe, 1953
* Emarginula retecosa Adams, 1851
* Emarginula retrogyra D. G. Herbert, 2024
* Emarginula rollandi P. Fischer, 1857 (uncertain: status in recent literature not researched)
* Emarginula rosea Bell, 1824
* Emarginula salebrosa D. G. Herbert, 2024
* † Emarginula sanctaecaatharine Passy, 1832
* Emarginula scabricostata A. Adams, 1852
* Emarginula scabriuscula Adams, 1852
* Emarginula sicula Gray, 1825 - dagger emarginula
* Emarginula sinica Lu, 1986
* Emarginula solidula Costa, 1829
* Emarginula souverbiana Pilsbry, 1890
* Emarginula spinosa Deshayes, 1863
* † Emarginula squammata Grateloup, 1836
* † Emarginula striatissima Traub, 1979
* Emarginula striatula Quoy and Gaimard, 1834 - dagger emarginula
* Emarginula sublaevis Schepman, 1908
* Emarginula subtilitexta Verco, 1908
* Emarginula superba Hedley, 1906
* Emarginula suspira Simone & Cunha, 2014
* Emarginula tenera Locard, 1892
* † Emarginula teneraeformis Lozouet, 1999
* Emarginula tenuicostata Sowerby, 1863
* Emarginula teramachii Habe, 1953
* Emarginula textilis Gould, 1859 in 1859-61
* Emarginula thomasi Crosse, 1864
* Emarginula thorektes Kilburn, 1978
* Emarginula tosaensis Habe, 1953
* †Emarginula transenna Tenison Woods, 1877
* Emarginula tuberculosa Libassi, 1859 - tuberculate emarginula
* Emarginula undulata Melvill & Standen, 1903
* Emarginula velascoensis Shasky, 1961
* Emarginula velascoi Rehder, 1980
* Emarginula viridibrunneis Poppe & Tagaro, 2020
* Emarginula viridicana Herbert & Kilburn, 1985
* Emarginula viridis Poppe & Tagaro, 2020
* † Emarginula wannonensis G. F. Harris, 1897
* † Emarginula washingtoniana Squires & Goedert, 1994
* Emarginula xishaensis Lu, 1986
* Emarginula yangeorum Poppe & Tagaro, 2020

## Synonyms

* Emarginula altilis Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
* Emarginula amitina Iredale, 1925: synonym of Emarginula dilecta A. Adams, 1852
* Emarginula angustata Thiele, 1915: synonym of Tugali decussata A. Adams, 1852
* Emarginula arabica A. Adams, 1852: synonym of Octomarginula arabica (A. Adams, 1852)
* Emarginula arconatii Issel, 1869: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
* Emarginula australis Quoy & Gaimard, 1834: synonym of Montfortia subemarginata (Blainville, 1819)
* Emarginula bella Gabb, 1865: synonym of Scelidotoma bella (Gabb, 1865)
* Emarginula biangulata Sowerby, 1901: synonym of Emarginella biangulata (G. B. Sowerby III, 1901)
* † Emarginula blainvillii Defrance, 1825: synonym of † Rimula blainvillii (Defrance, 1825)
* Emarginula brevirimata Deshayes, 1862: synonym of Montfortula brevirimata (Deshayes, 1862)
* Emarginula clathrata A. Adams & Reeve, 1850: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
* Emarginula clathrata Pease, 1863: synonym of Emarginula dilecta A. Adams, 1852
* Emarginula conica Lamarck, 1801: synonym of Emarginula fissura (Linnaeus, 1758)
* Emarginula conica Schumacher, 1817 non Lamarck, 1801: synonym of Emarginula rosea Bell, 1824
* Emarginula dentigera Heilprin, 1889: synonym of Hemimarginula dentigera (Heilprin, 1889)
* Emarginula elata Locard, 1898: synonym of Emarginula christiaensi Piani, 1985
* Emarginula elongata O. G. Costa, 1829: synonym of Emarginula octaviana Coen, 1939
* Emarginula eximia A. Adams, 1852: synonym of Emarginella eximia (A. Adams, 1852)
* Emarginula flindersi Cotton, 1930: synonym of Emarginula convexa Hedley, 1907
* Emarginula fuliginea Kuroda, 1941: synonym of Variemarginula variegata (A. Adams, 1852)
* Emarginula guernei Dautzenberg & Fischer, 1896: synonym of Emarginula tuberculosa Libassi, 1859
* Emarginula hedleyi Thiele, 1915: synonym of Emarginula candida A. Adams, 1852
* Emarginula incisula(sic): synonym of Emarginella incisura (A. Adams, 1852)
* Emarginula incisura A. Adams, 1852: synonym of Emarginella incisura (A. Adams, 1852)
* Emarginula kimberti Cotton, 1930: synonym of Laeviemarginula kimberti (Cotton, 1930)
* Emarginula modesta H. Adams, 1872: synonym of Hemimarginula modesta (H. Adams, 1872)
* Emarginula muelleri Forbes & Hanley, 1849: synonym of Emarginula fissura (Linnaeus, 1758)
* Emarginula ostheimerae Abbott, 1958: synonym of Octomarginula ostheimerae (Abbott, 1958)
* Emarginula panhi Quoy & Gaimard, 1834: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
* Emarginula papillosa Risso 1826: synonym of Emarginula huzardii Payraudeau, 1826
* Emarginula parmophoidea Quoy & Gaimard, 1834: synonym of Tugali elegans Gray, 1843
* Emarginula peasei Thiele, 1915: synonym of Emarginula dilecta A. Adams, 1852
* Emarginula phrygium Herbert & Kilburn, 1985: synonym of Agariste phrygium (Herbert & Kilburn, 1986) (original combination)
* Emarginula picta Dunker, 1860: synonym of Montfortula picta (Dunker, 1860)
* Emarginula pileata Gould, 1859: synonym of Variemarginula punctata (A. Adams, 1852)
* Emarginula pileum Heilprin, 1889: synonym of Hemimarginula pileum (Heilprin, 1889)
* Emarginula planulata A. Adams, 1852: synonym of Emarginella planulata (A. Adams, 1852)
* Emarginula polygonalis A. Adams, 1852: synonym of Hemitoma polygonalis (A. Adams, 1852)
* Emarginula pulchreclathrata Tomlin, 1932: synonym of Emarginula agulhasensis Thiele, 1925
* Emarginula pumila A. Adams, 1852: synonym of Hemimarginula pumila (A. Adams, 1852)
* Emarginula punctata A. Adams, 1852: synonym of Variemarginula punctata (A. Adams, 1852)
* Emarginula reticulata Sowerby, 1813: synonym of Emarginula fissura (Linnaeus, 1758)
* Emarginula rugosa Quoy & Gaimard, 1834: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
* Emarginula scutellata Deshayes, 1863: synonym of Octomarginula scutellata (Deshayes, 1863)
* Emarginula sibogae Schepman, 1908: synonym of Emarginella sibogae (Schepman, 1908)
* Emarginula simpla Christiaens, 1987: synonym of Hemimarginula simpla (Christiaens, 1987)
* Emarginula stellata A. Adams, 1852: synonym of Montfortula rugosa (Quoy & Gaimard, 1834)
* Emarginula subclathrata Pilsbry, 1890: synonym of Emarginula dilecta A. Adams, 1852
* Emarginula subemarginata Blainville, 1819: synonym of Montfortia subemarginata (Blainville, 1819)
* Emarginula subrugosa Thiele, 1916: synonym of Hemimarginula subrugosa (Thiele, 1916)
* Emarginula sulcifera A. Adams, 1852: synonym of Montfortulana sulcifera (A. Adams, 1852)
* Emarginula tricarinata Pilsbry, 1890: synonym of Montfortista panhi (Quoy & Gaimard, 1834)
* Emarginula tricostata G.B. Sowerby I, 1823: synonym of Hemitoma octoradiata (Gmelin, 1791)
* Emarginula vadososinuata Yokoyama, 1922: synonym of Tugalina vadososinuata (Yokoyama, 1922)
* Emarginula vadum Barnard, 1963: synonym of Emarginula undulata Melvill & Standen, 1903
* Emarginula variegata A. Adams, 1852: synonym of Variemarginula variegata (A. Adams, 1852)