# Taxonomy of the Dermaptera

This taxonomy of the Dermaptera follows Engel & Haas (2007) to the rank of genus.

## Taxonomy

* Order Dermaptera de Geer, 1773
   * Suborder †Archidermaptera Bey-Bienko, 1936
      * Superfamily †Protodiplatyoidea Martynov, 1925a
         * Family †Protodiplatyidae Martynov, 1925a
                           * Genus †Abrderma Xing, Shih, & Ren, 2016
                           * Genus †Aneuroderma Xiong, Engel, & Ren, 2021
                           * Genus †Anisodiplatys Vishniakova, 1980
                           * Genus †Applanatiforceps Yin, Shih, Engel, & Ren, 2021
                           * Genus †Archidermapteron Vishniakova, 1980
                           * Genus †Asiodiplatys Vishniakova, 1980
                           * Genus †Babderma Xing, Shih, & Ren, 2016
                           * Genus †Brevicula Whalley, 1985
                           * Genus †Longicerciata Zhang, 1994
                           * Genus †Microdiplatys Vishniakova, 1980
                           * Genus †Perissoderma Xing, Shih, & Ren, 2016
                           * Genus †Protodiplatys Martynov, 1925
                           * Genus †Sinoprotodiplatys Nel, Aria, Garrouste, & Waller, 2012
         * Family †Dermapteridae Vishniakova, 1980
                           * Genus †Dacryoderma Engel, 2021
                           * Genus †Dimapteron Kelly, Ross, & Jarzembowski, 2016
                           * Genus †Dermapteron Martynov, 1925
                           * Genus †Grammoderma Yin, Shih, Engel, & Ren, 2024
                           * Genus †Jurassimedeola Zhang, 2002
                           * Genus †Palaeodermapteron Zhao, Shih, & Ren, 2011
                           * Genus †Phanerogramma Cockerell, 1915
                           * Genus †Sinoplaeodermata Zhang, 2002
                           * Genus †Valdopteron Kelly, Ross, & Jarzembowski, 2016
         * Family †Turanoviidae Engel, 2003
                           * Genus †Turanovia Vishniakova, 1980
         * Family †Incertae sedis in †Archidermaptera Bey-Bienko, 1936
                           * Genus †Trivenapteron Kelly, Ross, & Jarzembowski, 2016
   * Suborder †Eodermaptera Engel, 2003
      * Superfamily †Bellodermatoidea Zhao, Shih, & Ren, 2010
         * Family †Bellodermatidae Zhao, Shih, & Ren, 2010
                           * Genus †Belloderma Zhao, Shih, & Ren, 2010
      * Superfamily †Semenovioloidea Vishniakova, 1980
         * Family †Semenoviolidae Vishniakova, 1980
               * Subfamily †Semenoviolinae Vishniakova, 1980
                           * Genus †Semenoviola Vishniakova, 1980
                           * Genus †Semenovioloides Vishniakova, 1980
               * Subfamily †Aglyptodermatinae Xiong, Engel, & Ren, 2021
                           * Genus †Aglyptoderma Xiong, Engel, & Ren, 2021
                           * Genus †Ekpagloderma Yin, Shih, Engel, & Ren, 2023
      * Infraorder †Turanodermaptera Engel, 2019
         * Superfamily †Turanodermatoidea Engel, 2003
            * Family †Turanodermatidae Engel, 2003
                           * Genus †Turanoderma Vishniakova, 1980
   * Suborder Neodermaptera Engel, 2003
      * Infraorder Protodermaptera Zacher, 1910
         * Superfamily Diplatyoidea Verhoeff, 1902a
            * Family Diplatyidae Verhoeff, 1902a
                           * Genus †Acanthodiplatys Ren, Zhang, Shih, & Ren, 2018
                           * Genus Circodiplatys Steinmann, 1986
                           * Genus Diplatys Audinet-Serville, 1831
                           * Genus †Hirtidiplatys Ren, Zhang, Shih, & Ren, 2018
                           * Genus Lobodiplatys Steinmann, 1974
                           * Genus Mesodiplatys Steinmann, 1986
                           * Genus Schizodiplatys Steinmann, 1974
                           * Genus †Tytthodiplatys Engel, 2011
            * Family Haplodiplatyidae Engel, 2017
                           * Genus Haplodiplatys Hincks, 1955
                           * Genus †Zeugmadiplatys Peng, Engel, Boderau, Zhuo, & Nel, 2024
         * Superfamily Pygidicranoidea Verhoeff, 1902a
            * Family Pygidicranidae Verhoeff, 1902a
               * Subfamily †Astreptolabidinae Engel, 2011
                           * Genus †Astreptolabis Engel, 2011
               * Subfamily †Burmapygiinae Engel & Grimaldi, 2004
                           * Genus †Burmapygia Engel & Grimaldi, 2004
               * Subfamily Anataeliinae Burr, 1909b
                           * Genus Anataelia Bolivar, 1899
               * Subfamily Blandicinae Burr, 1915
                           * Genus Alloblandex Hincks, 1957
                           * Genus Austroblandex Brindle, 1986
                           * Genus Blandex Burr, 1912
                           * Genus Parablandex Brindle, 1966
               * Subfamily Brindlesiinae Srivastava, 1985b
                           * Genus Brindlesia Srivastava, 1985b
               * Subfamily Challiinae Steinmann, 1973
                           * Genus Challia Burr, 1904
               * Subfamily Cylindrogastrinae Maccagno, 1929
                           * Genus Cylindrogaster Stål, 1855
               * Subfamily Diplatymorphinae Boeseman, 1954
                           * Genus Diplatymorpha Boeseman, 1954
               * Subfamily Echinosomatinae Burr, 1910a
                           * Genus Parapsalis Borelli, 1921
                           * Genus Echinosoma Audinet-Serville, 1831
               * Subfamily Esphalmeninae Burr, 1909a
                           * Genus Esphalmenus Burr, 1909
               * Subfamily Karschiellinae Verhoeff, 1902a
                           * Genus Bormansia Verhoeff, 1902
                           * Genus Karschiella Verhoeff, 1902
               * Subfamily †Prodacnodinae Peng, Engel, Liu, & Nel, 2024
                           * Genus †Prodacnodes Peng, Engel, Liu, & Nel, 2024
               * Subfamily Pygidicraninae Verhoeff, 1902a
                           * Genus Acrania Burr, 1915
                           * Genus Cranopygia Burr, 1908
                           * Genus Dacnodes Burr, 1907
                           * Genus Paracranopygia Steinmann, 1986
                           * Genus Pygidicrana Audinet-Serville, 1831
                           * Genus Tagalina Dohrn, 1863
               * Subfamily Pyragrinae Verhoeff, 1902a
                           * Genus Echinopsalis Bormans, 1893
                           * Genus †Gracilipygia Ren, Zhang, Shih, & Ren, 2017
                           * Genus Pyragra Audinet-Serville, 1831
                           * Genus Pyragropsis Borelli, 1908
               * Subfamily †Stonychopygiinae Engel & Huang, 2017
                           * Genus †Stonychopygia Engel & Huang, 2017
               * Subfamily Incertae sedis in Pygidicranidae Verhoeff, 1902a
                           * Genus †Archaeosoma Zhang, 1994
                           * Genus †Eminepygia Chen & Zhang, 2021
                           * Genus †Gallinympha Perrichot & Engel, 2011
                           * Genus †Geosoma Zhang, 1997
                           * Genus †Robustipygia Ren, Zhang, Shih, & Ren, 2018
      * Infraorder Epidermaptera Engel, 2003
         * Parvorder Paradermaptera Verhoeff, 1902a
            * Superfamily Hemimeroidea Sharp, 1895
               * Family Hemimeridae Sharp, 1895
                           * Genus Araeomerus Maa, 1974
                           * Genus Hemimerus Walker, 1871
            * Superfamily Apachyoidea Verhoeff, 1902a
               * Family Apachyidae Verhoeff, 1902a
                           * Genus Apachys Audinet-Serville, 1831
                           * Genus Dendroiketes Burr, 1909
         * Parvorder Metadermaptera Engel, 2003
            * Superfamily Anisolabidoidea Verhoeff, 1902a
               * Family Anisolabididae Verhoeff, 1902a
                           * Genus Aborolabis Srivastava, 1969
                           * Genus Anisolabis Fieber, 1853
                           * Genus Canarilabis Steinmann, 1985
                           * Genus Carcinophora Scudder, 1876
                           * Genus Epilandex Hebard, 1927
                           * Genus Euborellia Burr, 1909
                           * Genus Flexiolabis Steinmann, 1989
                           * Genus Gonolabis Burr, 1910
                           * Genus Indolabis Steinmann, 1989
                           * Genus Mongolabis Zacher, 1911
                           * Genus Neolabis Brindle, 1981
                           * Genus Ornatolabis Steinmann, 1989
                           * Genus Paraflexiolabis Steinmann, 1989
                           * Genus Placolabis Bey-Bienko, 1959
                           * Genus Socotralabis Kočárek, 2014
                           * Genus Thekalabis Kapoor, 1967
                           * Genus Zacheria Steinmann, 1975
                  * Subfamily Anophthalmolabidinae Steinmann, 1975
                           * Genus Anophthalmolabis Brindle, 1968
                  * Subfamily Antisolabidinae Brindle, 1978a
                           * Genus Antisolabis Burr, 1911
                  * Subfamily Brachylabidinae Burr, 1908a
                           * Genus Brachylabis Dohrn, 1864
                           * Genus Ctenisolabis Verhoeff, 1902
                           * Genus Metisolabis Burr, 1910
                  * Subfamily †Cretolabiinae Engel & Haas, 2007
                           * Genus †Cretolabia Popham, 1990
                  * Subfamily Gonolabininae Popham and Brindle, 1966c
                           * Genus Gonolabina Verhoeff, 1902
                  * Subfamily Idolopsalidinae Steinmann, 1975
                           * Genus Idolopsalis Borelli, 1910
                  * Subfamily Isolabidinae Verhoeff, 1902b
                           * Genus Africolabis Brindle, 1978
                           * Genus Geracodes Hebard, 1917
                           * Genus Isolabis Verhoeff, 1902
                           * Genus Pterolabis Steinmann, 1989
                  * Subfamily Parisolabidinae Verhoeff, 1904
                           * Genus Parisolabis Verhoeff, 1904
                           * Genus Parisopsalis Burr, 1914
                  * Subfamily Platylabiinae Burr, 1911
                           * Genus Platylabia Dohrn, 1867
                  * Subfamily Titanolabidinae Srivastava, 1982
                           * Genus Titanolabis Burr, 1910
                     * Subfamily Incertae sedis in Anisolabididae Verhoeff, 1902a
                           * Genus †Geroncolabis Engel & Chatzimanolis, 2010
                           * Genus †Kotejalabis Engel & Chatzimanolis, 2005
                           * Genus †Palaeocarcinophora Engel & Chatzimanolis, 2010
                           * Genus †Spiladopygia Engel & Chatzimanolis, 2010
                           * Genus †Toxolabis Engel & Grimaldi, 2014
         * Parvorder Eteodermaptera Engel, 2003
            * Nanorder Plesiodermaptera Engel, 2003
               * Superfamily Labiduroidea Verhoeff, 1902a
                  * Family Labiduridae Verhoeff, 1902a
                     * Subfamily Allostethinae Verhoeff, 1904
                           * Genus Allostethella Zacher, 1910
                           * Genus Allostethus Verhoeff, 1904
                           * Genus Gonolabidura Zacher, 1910
                           * Genus Protolabidura Steinmann, 1985
                     * Subfamily Labidurinae Verhoeff, 1902a
                           * Genus Forcipula Bolivar, 1897
                           * Genus Labidura Leach, 1815
                           * Genus Tomopygia Burr, 1904
                     * Subfamily †Metaxylabidinae Peng, Engel, Zhou, & Nel, 2024
                           * Genus †Metaxylabis Peng, Engel, Zhou, & Nel, 2024
                     * Subfamily Nalinae Steinmann, 1975
                           * Genus Nala Zacher, 1910
                     * Subfamily †Zigrasolabidinae Engel, 2022
                           * Genus †Zigrasolabis Engel & Grimaldi, 2014
                           * Genus †Tricholabidura Peng & Engel, 2022
                     * Subfamily Incertae sedis in Labiduridae Verhoeff, 1902a
                           * Genus †Acantholabis Mao, Engel, & Ren, 2020
                           * Genus †Caririlabia Martins-Neto, 1990
                           * Genus †Myrrholabia Engel & Grimaldi, 2004
            * Nanorder Eudermaptera Verhoeff, 1902a
               * Superfamily Forficuloidea Latreille, 1810
                  * Family Arixeniidae Jordan, 1909
                           * Genus Arixenia Jordan, 1909
                           * Genus Xeniaria Maa, 1974
                  * Family Spongiphoridae Verhoeff, 1902a
                     * Subfamily Caecolabiinae Steinmann, 1990
                           * Genus Caecolabia Brindle, 1975
                     * Subfamily Cosmogeracinae Brindle, 1982
                           * Genus Cosmogerax Hebard, 1933
                     * Subfamily Geracinae Brindle, 1971
                           * Genus Apterogerax Engel, 2017
                           * Genus Barygerax Hebard, 1917
                           * Genus Eugerax Hebard, 1917
                           * Genus Gerax Hebard, 1917
                           * Genus †Nesogerax Engel, 2017
                           * Genus Pseudovostox Borelli, 1926
                           * Genus Yepezia Brindle, 1982
                     * Subfamily Isolaboidinae Brindle, 1978b
                           * Genus Isolaboides Hincks, 1958
                           * Genus Paralaboides Steinmann, 1989
                     * Subfamily Isopyginae Hincks, 1951
                           * Genus Isopyge Borelli, 1931
                     * Subfamily Labiinae Burr, 1909b
                           * Genus Chaetolabia Brindle, 1972
                           * Genus Circolabia Steinmann, 1987
                           * Genus Isolabella Verhoeff, 1902
                           * Genus Labia Leach, 1815
                           * Genus Paralabella Steinmann, 1990
                           * Genus Paraspania Steinmann, 1985
                           * Genus Sphingolabis Bormans, 1883
                           * Genus Spirolabia Steinmann, 1987
                     * Subfamily Nesogastrinae Verhoeff, 1902a
                           * Genus Nesogaster Verhoeff, 1902
                     * Subfamily Nesolabiinae Engel, 2017
                           * Genus Nesolabia Hincks, 1957
                     * Subfamily Pericominae Burr, 1911a
                           * Genus Parapericomus Ramamurthi, 1967
                           * Genus Pericomus Burr, 1911
                     * Subfamily Ramamurthiinae Steinmann, 1975
                           * Genus Ramamurthia Steinmann, 1975
                     * Subfamily Rudracinae Srivastava, 1995
                           * Genus Rudrax Srivastava, 1995
                     * Subfamily Sparattinae Verhoeff, 1902a
                        * Tribe Auchenomini Burr, 1909b
                           * Genus Auchenomus Karsch, 1886
                        * Tribe Chaetospaniini Steinmann, 1990
                           * Genus Chaetospania Karsch, 1886
                        * Tribe Sparattini Verhoeff, 1902a
                           * Genus Mecomera Audinet-Serville, 1838
                           * Genus Sparatta Audinet-Serville, 1838
                     * Subfamily Spongiphorinae Verhoeff, 1902a
                           * Genus Filolabia Steinmann, 1989
                           * Genus Formicilabia Rehn & Hebard, 1917
                           * Genus Homotages Burr, 1909
                           * Genus †Ikelus Estrada-Álvarez & Núñez-Bázan, 2023
                           * Genus Irdex Burr, 1911
                           * Genus Marava Burr, 1911
                           * Genus Pseudomarava Steinmann, 1989
                           * Genus Purex Burr, 1911
                           * Genus Spongiphora Audinet-Serville, 1831
                           * Genus Spongovostox Burr, 1911
                           * Genus Vostox Burr, 1911
                     * Subfamily Strongylopsalinae Burr, 1911a
                           * Genus Strongylolabis Steinmann, 1986
                           * Genus Strongylopsalis Burr, 1900
                     * Subfamily Vandicinae Burr, 1911a
                           * Genus Vandex Burr, 1911
                     * Subfamily Incertae sedis in Spongiphoridae Verhoeff, 1902
                           * Genus Paratages Srivastava, 1987
                  * Family Chelisochidae Verhoeff, 1902a
                     * Subfamily Chelisochinae Verhoeff, 1902a
                           * Genus Adiathella Brindle, 1970
                           * Genus Adiathetus Burr, 1907
                           * Genus Chelisochella Verhoeff, 1902
                           * Genus Chelisoches Scudder, 1876
                           * Genus †Chelisoficula Nel, Waller, Albouy, Menier, & de Ploëg, 2003
                           * Genus Euenkrates Rehn, 1927
                           * Genus Exypnus Burr, 1907
                           * Genus Gressitolabis Brindle, 1970
                           * Genus Hamaxas Burr, 1907
                           * Genus Lamprophorella Mjöberg, 1924
                           * Genus Proreus Burr, 1907
                           * Genus Schizochelisoches Steinmann, 1987
                           * Genus Schizoproreus Steinmann, 1987
                           * Genus Solenosoma Burr, 1907
                     * Subfamily Genitalatinae Steinmann, 1987
                           * Genus Genitalata Kapoor, 1974
                     * Subfamily Kinesinae Srivastava, 2003
                           * Genus Kinesis Burr, 1907
                     * Subfamily Incertae sedis in Chelisochidae Verhoeff, 1902a
                           * Genus †Amblypygia Engel, 2016
                           * Genus †Sthenotropis Engel, 2016
                  * Family Forficulidae Latreille, 1810
                     * Subfamily Allodahliinae Verhoeff, 1902a
                           * Genus Allodahlia Verhoeff, 1902
                           * Genus Brindleiana Steinmann, 1975
                           * Genus Eulithinus Hincks, 1935
                     * Subfamily Ancistrogastrinae Verhoeff, 1902a
                           * Genus Ancistrogaster Stål, 1855
                           * Genus Litocosmia Hebard, 1917
                           * Genus Osteulcus Burr, 1907
                           * Genus Paracosmia Borelli, 1909
                           * Genus Praos Burr, 1907
                           * Genus Sarcinatrix Rehn, 1903
                     * Subfamily Anechurinae Verhoeff, 1902a
                           * Genus Anechura Scudder, 1876
                           * Genus Chelidura Latreille, 1825
                           * Genus Chelidurella Verhoeff, 1902
                           * Genus Eumegalura Bey-Bienko, 1934
                           * Genus Mesasiobia Semenov, 1908
                           * Genus Mesochelidura Verhoeff, 1902
                           * Genus Neopterygidea Srivastava, 1984
                           * Genus Oreasiobia Semenov, 1936
                           * Genus Perirrhytus Burr, 1911
                           * Genus Pseudochelidura Verhoeff, 1902
                           * Genus Pterygidea Verhoeff, 1902
                     * Subfamily Diaperasticinae Burr, 1907
                           * Genus Diaperasticus Burr, 1907
                     * Subfamily Forficulinae Latreille, 1810
                           * Genus Afroforficula Steinmann, 1990
                           * Genus Apterygidea Westwood, 1840
                           * Genus Champaipites Burr, 1907
                           * Genus Doru Burr, 1907
                           * Genus Elaunon Burr, 1907
                           * Genus Forficula Linnaeus, 1758
                           * Genus Guanchia Burr, 1911
                           * Genus Mesolabia Shiraki, 1905
                           * Genus Parlax Burr, 1911
                           * Genus Proforficula Steinmann, 1990
                           * Genus Skalistes Burr, 1907
                           * Genus †Succiniforcipula Engel, 2016
                           * Genus Tauropygia Brindle, 1970
                     * Subfamily Neolobophorinae Burr, 1907
                           * Genus Dubianus Estrada-Álvarez & Núñez-Bázan, 2022
                           * Genus Eudohrnia Burr, 1907
                           * Genus Metresura Rehn, 1922
                           * Genus Neolobophora Scudder, 1875
                           * Genus Setocordax Brindle, 1970
                           * Genus Tristanella Borelli, 1909
                     * Subfamily Opisthocosmiinae Verhoeff, 1902a
                           * Genus Acanthocordax Günther, 1929
                           * Genus Chaetocosmia Nishikawa, 1973
                           * Genus Cipex Burr, 1910
                           * Genus Cordax Burr, 1910
                           * Genus Eparchus Burr, 1907
                           * Genus Eutimomena Bey-Bienko, 1970
                           * Genus Hypurgus Burr, 1907
                           * Genus Neoopisthocosmia Steinmann, 1990
                           * Genus Opisthocosmia Dohrn, 1865
                           * Genus Parasondax Srivastava, 1978
                           * Genus Parasyntonus Steinmann, 1990
                           * Genus Paratimomenus Steinmann, 1974
                           * Genus Pareparchus Burr, 1911
                           * Genus Prosadiya Hebard, 1923
                           * Genus Sondax Burr, 1910
                           * Genus Spinosocordax Steinmann, 1988
                           * Genus Syntonus Burr, 1910
                           * Genus Timomenus Burr, 1907
                     * Subfamily Skendylinae Burr, 1907
                           * Genus Afrocosmia Hincks, 1960
                           * Genus Brachycosmiella Steinmann, 1990
                           * Genus Cosmiella Verhoeff, 1902
                           * Genus Cosmiola Bey-Bienko, 1959
                           * Genus Forcepsia Moreira, 1930
                           * Genus Kleter Burr, 1907
                           * Genus Liparura Burr, 1907
                           * Genus Lipodes Burr, 1907
                           * Genus Mixocosmia Borelli, 1909
                           * Genus Neocosmiella Hebard, 1919
                           * Genus Obelura Burr, 1907
                           * Genus Paracosmiella Steinmann, 1990
                     * Subfamily Incertae sedis in Forficulidae Latreille, 1810
                           * Genus †Rupiforficula Engel & Chatzimanolis, 2010
                  * Family Incertae sedis in Forficuloidea Latreille, 1810
                           * Genus †Litholabis Engel & Chatzimanolis, 2010
                           * Genus †Petrolabis Engel & Chatzimanolis, 2010
               * Superfamily Incertae sedis in Epidermaptera Engel, 2003
                           * Genus †Autrigonoforceps Engel & Peris, 2015
                           * Genus †Laasbium Scudder, 1900
                           * Genus †Labiduromma Scudder, 1890
                           * Genus †Rhadinolabis Engel, Ortega-Blanco, & Azar, 2011

## Nomina dubia

Family †Ocelliidae Ewing, 1942 is considered a nomen dubium.