# Obrimini

The Obrimini are the most species-rich tribe of the Phasmatodea family of the Heteropterygidae native to Southeast Asia.

## Description

The Obrimini differ from their sister tribe the Hoplocloniini by the structure of the secondary ovipositor at the abdomen of the females, which surrounds the actual ovipositor. It is formed in the Obrimini dorsally from the eleventh abdominal tergum called supraanal plate or epiproct, while in the Hoplocloniini it originated from the tenth tergum.

## Distribution area

The distribution area of the Obrimini extends from Borneo to the east and includes the Philippines, Sulawesi, most of the Moluccas and New Guinea. Farthest east lies with Viti Levu, the main island of the Fiji group, the distribution area of Pterobrimus depressus.

## Taxonomy

Obrimini
Miroceramia
Pterobrimus
Theramenes
Tisamenus
Stenobrimus
Obrimus
Brasidas Euobrimus
Mearnsiana
Aretaon
Eubulides
Obrimini sp. 'Negros' = Trachyaretaon negrosanon
Sungaya
Trachyaretaon :
Relationships of the examined Obrimini genera by Sarah Bank et al (2021) :
Brunner von Wattenwyl built in 1893 for the genera already described Obrimus, Hoploclonia, Tisamenus, Pylaemenes, Dares and Datames (today a synonym to Pylaemenes) the tribe Obrimini (abbreviated there as Obrimi.). He placed these together with the genus Heteropteryx and the South American tribe Cladomorphini (abbreviated there as Cladomorphi.) in the family Cladomorphidae (now synonymous with Cladomorphinae). In the following years, basesd on this work, mostly genera of today's Obriminae and Dataminae were listed in this tribe, for example by Josef Redtenbacher in 1906. Lawrence Bruner raised the Obrimini to the rank of a family in 1915. James Abram Garfield Rehn and his son John William Holman Rehn divided the Obriminae, which they only referred to as a subfamily, into the tribes Obrimini and Datamini in 1939. Both tribes were transferred in 1953 by Klaus Günther to the subfamily Heteropteryginae. In 2004 Oliver Zompro raised this subfamily to the rank of family and the tribes contained in the rank of subfamilies or in the rank of a separate family (Anisacanthidae). At the same time he built three tribes in the subfamily Obriminae. In addition to the Obrimini, these were the Miroceramiini and the Eubulidini. The latter was in 2016 by Frank H. Hennemann et al synonymized with the Obrimini. At the same time, the new tribe Tisamenini was established within the Obriminae. In a study based on genetic analysis, published in 2021, Sarah Bank et al. synonymized the tribe Miroceramiini and the tribe Tisamenini with the Obrimini and placed next to this the Hoplocloniini as the only other tribe in the subfamily Obriminae. In addition, a species previously referred to as Trachyaretaon sp. 'Negros' was identified as a representative of an as yet undescribed genus. Hennemann described this in 2023 alongside many other species of the tribe as Trachyaretaon negrosanon.

### Genera

| Genus | Species |
| Aretaon Rehn, J.A.G. & Rehn, J.W.H., 1939 | Aretaon asperrimus (Redtenbacher, 1906) Aretaon muscosus (Redtenbacher, 1906) |
| Armadolides Hennemann, 2023 | Armadolides manobo (Acola, Naredo & Eusebio, 2022) |
| Brasidas Rehn, J.A.G. & Rehn, J.W.H., 1939 | Brasidas bakeri (Rehn, J.A.G. & Rehn, J.W.H., 1939) Brasidas cavernosus (Stål, 1877) Brasidas foveolatus (Redtenbacher, 1906) Brasidas lacerta (Redtenbacher, 1906) Brasidas malaki Hennemann, 2023 Brasidas manobo Hennemann, 2023 Brasidas rehni Hennemann, 2023 Brasidas samarensis Rehn, J.A.G. & Rehn, J.W.H., 1939 Brasidas viscayanus Rehn, J.A.G. & Rehn, J.W.H., 1939 Brasidas waray Hennemann, 2023 |
| Eubulides Stål, 1877 | Eubulides alutaceus Stål, 1877 Eubulides blaan Hennemann, 2023 Eubulides constanti Hennemann, 2023 Eubulides igorrote Rehn, J.A.G. & Rehn, J.W.H., 1939 Eubulides lumawigi Hennemann, 2023 Eubulides taylori Rehn, J.A.G. & Rehn, J.W.H., 1939 Eubulides timog Hennemann, 2023 |
| Heterocopus Redtenbacher, 1906 | Heterocopus leprosus Redtenbacher, 1906 |
| Mearnsiana Rehn, J.A.G. & Rehn, J.W.H., 1939 | Mearnsiana bullosa Rehn, J.A.G. & Rehn, J.W.H., 1939 Mearnsiana maranao Hennemann, 2023 |
| Miroceramia Günther, 1934 | Miroceramia westwoodii (Bates, 1865) |
| Obrimus Stål, 1875 | Obrimus bicolanus Rehn, J.A.G. & Rehn, J.W.H., 1939 Obrimus bufo (Westwood, 1848) Obrimus mesoplatus (Westwood, 1848) Obrimus uichancoi Rehn, J.A.G. & Rehn, J.W.H., 1939 |
| Pterobrimus Redtenbacher, 1906 | Pterobrimus depressus Redtenbacher, 1906 |
| Stenobrimus Redtenbacher, 1906 | Stenobrimus bolivari Redtenbacher, 1906 Stenobrimus lumad Lit & Eusebio, 2010 Stenobrimus pilipinus Eusebio, Lit & Lucañas, 2023 Stenobrimus tagalog Rehn, J.A.G. & Rehn, J.W.H., 1939 |
| Sungaya Zompro, 1996 | Sungaya aeta Hennemann, 2023 Sungaya dumagat Hennemann, 2023 Sungaya ibaloi Hennemann, 2023 Sungaya inexpectata Zompro, 1996 |
| Theramenes Stål, 1875 | Theramenes dromedarius Stål, 1877 Theramenes exiguus Hennemann & Conle, 2003 Theramenes letiranti Hennemann, 2023 Theramenes mandirigma Zompro & Eusebio, 2001 Theramenes olivaceus (Westwood, 1859) |
| Tisamenus Stål, 1875 | Tisamenus alviolanus Lit & Eusebio, 2010 Tisamenus armadillo Redtenbacher, 1906 Tisamenus asper (Bolívar, 1890) Tisamenus atropos (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus cervicornis Ignacio Bolívar, 1890 Tisamenus clotho (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus deplanatus (Westwood, 1848) Tisamenus draconina (Westwood, 1848) Tisamenus fratercula (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus hystrix (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus hebardi (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus kalahani Lit & Eusebio, 2005 Tisamenus lachesis (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus polillo (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus ranarius (Westwood, 1859) Tisamenus serratorius Stål, 1875 Tisamenus spadix (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus summaleonilae Lit & Eusebio, 2005 Tisamenus tagalog (Rehn, J.A.G. & Rehn, J.W.H., 1939) |
| Trachyaretaon Rehn, J.A.G. & Rehn, J.W.H., 1939 | Trachyaretaon bresseeli Hennemann, 2023 Trachyaretaon carmelae Lit & Eusebio, 2005 Trachyaretaon echinatus (Stål, 1877) Trachyaretaon gatla Zompro, 2004 Trachyaretaon maliit Hennemann, 2023 Trachyaretaon mangyan Hennemann, 2023 Trachyaretaon nakatago Hennemann, 2023 Trachyaretaon negrosanon Hennemann, 2023 Trachyaretaon tumandok Hennemann, 2023 |