# 2013 in paleobotany

This list of 2013 in paleobotany records new fossil plant taxa that were described during 2013, as well as other significant discoveries and events related to paleobotany that occurred in the year.

## Chlorophyta

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Cymatiosphaera scitula | Sp. nov. | Valid | Wicander & Playford | Devonian (Famennian) | Saverton Shale | United States | A member of Chlorophyta belonging to the group Prasinophyceae. | |

## Bryophyta

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Brachytheciites sachalinensis | Sp. nov | Valid | Ignatov & Perkovsky | Middle Eocene | Sakhalinian amber | Russia | A moss belonging to the family Brachytheciaceae. |
| Bryiidites | Gen. et sp. nov. | Valid | Barclay et al. | Late Cretaceous (late Cenomanian) | Middle Dakota Formation | United States | A member of Bryidae of uncertain phylogenetic placement. The type specimen is Bryiidites utahensis. |
| Bryokhutuliinia crassimarginata | Sp. nov | Valid | Ignatov et al. | Late Jurassic | | Russia | A moss. |
| Calymperites | Gen. et sp. nov | Valid | Ignatov & Perkovsky | Late Eocene | Rovno amber | Ukraine | A moss belonging to the group Dicranales (sensu lato). Genus includes new species C. ucrainicus. |
| Pseudoleskeellites | Gen. et sp. nov | Valid | Ignatov & Perkovsky | Middle Eocene | Sakhalinian amber | Russia | A moss belonging to the order Hypnales. Genus includes new species P. obscurus. |

## Ferns and fern allies

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Ashicaulis beipiaoensis | Sp. nov | Valid | Tian et al. | Middle Jurassic | Tiaojishan Formation | China | A member of Osmundaceae. Originally described as a species of Ashicaulis; transferred to the genus Millerocaulis by Bomfleur, Grimm & McLoughlin (2017). |
| Mygdykitia | Gen. et sp. nov | Valid | Golovneva & Shczepetov | Late Cretaceous (Campanian) | Ola Formation | Russia | A fern belonging to the family Dipteridaceae. Genus includes new species M. ochotica. |
| Palaeosorum | Gen. et sp. nov | Valid | Jacques & Zhou in Jacques, Su & Zhou | Miocene | Dajie Formation | China | A member of Polypodiaceae. The type specimen is Palaeosorum ellipticum. |
| Todea amissa | Sp. nov. | Valid | Carvalho in Carvalho et al. | Early Eocene | | Argentina | A species of Todea. |
| Yavanna | Gen. et sp. nov | Valid | Vera | Early Cretaceous (Aptian) | Cerro Negro Formation | Antarctica (Livingston Island) | A cyathealean of uncertain placement. Type species is Y. chimaerica. |

## Ginkgophytes

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Allicospermum angrenicum | Sp. nov. | Valid | Nosova | Middle Jurassic | Angren Formation | Uzbekistan | A Ginkgo-like plant described on the basis of seeds. | |
| Nagrenia | Gen. et sp. nov. | Valid | Nosova | Middle Jurassic | Angren Formation | Uzbekistan | A member of Ginkgoales described on the basis of peduncles furcating into several pedicels. Genus includes new species N. samylinae. | |

## Gnetophyta

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Chengia | Gen. et sp. nov. | Valid | Yang, Lin & Wang | Early Cretaceous (Aptian) | Yixian Formation | China | A member of Ephedraceae. The type specimen is Chengia laxispicata. |
| Friedsellowia gracilifolia | Gen. nov. et sp. nov. | Valid | Löwe et al. | Early Cretaceous | Crato Formation | Brazil | A gnetophyte. The type of a new genus. |
| Itajuba | Gen. et sp. nov | Valid | Ricardi-Branco et al. | Early Cretaceous (Aptian or Albian) | Crato Member of the Santana Formation | Brazil | A member of Gnetales. The type specimen is Itajuba yansanae. |

## Pachytestopsida

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Stephanospermum braidwoodensis | Sp. nov. | Valid | Spencer, Hilton & Sutton | Carboniferous | Mazon Creek fossil beds | United States | A member of Medullosales, a species of Stephanospermum. | |
| Stephanospermum shuichengensis | Sp. nov. | Valid | Spencer et al. | Permian (Lopingian) | Wangjiazhai Formation | China | A member of Medullosales, a species of Stephanospermum. | |

## Pinophyta

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Bancroftiastrobus | Gen. et sp. nov. | Valid | Rothwell et al. | Late Jurassic (early Kimmeridgian) | Kimmeridge Clay Formation | United Kingdom | A pinophyte of uncertain phylogenetic placement. Type species is B. digitata. |
| Brachyoxylon currumilii | Sp. nov. | Valid | Bodnar et al. | Early–Middle Jurassic | Cañadón Asfalto Formation | Argentina | A member of Pinophyta described on the basis of fossil wood. |
| Cunninghamia taylorii | Sp. nov. | Valid | Serbet, Bomfleur & Rothwell | Late Cretaceous (Campanian) | Horseshoe Canyon Formation | Canada | A species of Cunninghamia. |
| Elatocladus anorovii | Sp. nov | Valid | Golovneva & Shczepetov | Late Cretaceous (Campanian) | Ola Formation | Russia | A conifer of uncertain phylogenetic placement. |
| Pagiophyllum heshanense | Sp. nov. | Valid | Liu & Yao | Permian | | China | A species of Pagiophyllum. |
| Pararaucaria delfueyoi | Sp. nov. | Valid | Escapa et al. | Late Jurassic | Cañadón Calcáreo Formation | Argentina | A member of Pinophyta belonging to the family Cheirolepidiaceae, a species of Pararaucaria. |
| Picea yanensis | Sp. nov | Valid | Golovneva & Shczepetov | Late Cretaceous (Campanian) | Ola Formation | Russia | A spruce. |
| Prumnopitys anglica | Sp. nov. | Valid | Greenwood, Hill & Conran | Eocene | Branksome Formation | United Kingdom | A species of Prumnopitys. |
| Schizolepidopsis canicularis | Sp. nov. | Valid | Leslie et al. | Early Cretaceous (probably Aptian-Albian) | Tevshiin Govi Formation | Mongolia | A member of Pinophyta, possibly a member of the stem group of Pinaceae; a species of Schizolepidopsis. |
| Xuanweioxylon | Gen. et sp. nov. | Valid | He et al. | Permian (late Wuchiapingian-Changhsingian) | Xuanwei Formation | China | A member of Pinophyta. Genus includes new species X. scalariforme. |

## Angiosperms

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Anonaspermum warmanensis | Sp. nov. | Valid | Wang, Blanchard & Dilcher | Eocene | Cockfield Formation | United States | A member of Annonaceae. |
| Atli | Gen. et sp. nov. | Valid | Smith et al. | Late Cretaceous (Campanian) | | Canada | A ranunculalean known from a stem with lianoid anatomy. The type specimen is Atli morinii. |
| Bambusiculmus | Gen. et 2 sp. nov. | Valid | Wang & Zhou in Wang et al. | Miocene | Dajie Formation | China | A bamboo. The type specimen is Bambusiculmus angustus; genus also includes Bambusiculmus latus. |
| Bambusium angustifolia | Sp. nov. | Valid | Wang & Zhou in Wang et al. | Miocene | Dajie Formation | China | A bamboo, a species of Bambusium. |
| Bambusium latifolia | Sp. nov. | Valid | Wang & Zhou in Wang et al. | Miocene | Dajie Formation | China | A bamboo, a species of Bambusium. |
| Bombacacioxylon | Gen. et sp. nov. | Valid | Mehrotra et al. | Late Miocene to early Pliocene | Tipam Group | India | A possible relative of Bombax. The type specimen is Bombacacioxylon tertiarum. |
| Carpolithus warmanensis | Sp. nov. | Valid | Wang, Blanchard & Dilcher | Eocene | Cockfield Formation | United States | A fossil fruit. |
| Cornerocarpon | Gen. et sp. nov. | Valid | Grote in Wang, Blanchard & Dilcher | Eocene | Cockfield Formation | United States | A member of Rosales belonging or related to the family Moraceae. Genus includes new species C. copiosum. |
| Corylopsis grotei | Sp. nov. | Valid | Wang, Blanchard & Dilcher | Eocene | Cockfield Formation | United States | A species of Corylopsis. |
| Diospyros neyveliensis | Sp. nov. | Valid | Mukherjee & Prasad | Miocene | Neyveli Formation | India | A member of the family Ebenaceae described on the basis of fossil wood. Originally described as a species of Diospyros; Srivastava et al. (2018) transferred this species to the genus Ebenoxylon. |
| Firmiana oligocenica | Sp. nov. | Valid | Srivastava & Mehrotra | Oligocene (Chattian) | Tikak Parbat Formation | India | A species of Firmiana. |
| Grotea | Gen. et sp. nov. | Valid | Wang, Blanchard & Dilcher | Eocene | Cockfield Formation | United States | A member of the family Bignoniaceae. Genus includes new species G. warmanensis. |
| Guadua morronei | Sp. nov. | Valid | Brea, Zucol & Franco | Pliocene–Pleistocene | Ituzaingó Formation | Argentina | A bamboo, a species of Guadua. |
| Hamamelis warmanensis | Sp. nov. | Valid | Wang, Blanchard & Dilcher | Eocene | Cockfield Formation | United States | A witch-hazel. |
| Indovitis | Gen. et sp. nov. | Valid | Manchester, Kapgate & Wen | Late Cretaceous (Maastrichtian) | Deccan Intertrappean Beds | India | A member of the family Vitaceae. Genus includes new species I. chitaleyae. |
| Iterophyllum | Gen. et sp. nov. | Valid | Barral et al. | Early Cretaceous (late Barremian) | | Spain | A basal eudicot. The type specimen is Iterophyllum lobatum. |
| Jaguariba wiersemana | Gen. nov. et sp. nov. | Valid | Coiffard, Mohr & Bernardes-de-Oliveira | Early Cretaceous | Crato Formation | Brazil | Fossil Nymphaeales. |
| Lagerstroemia corvinusii | Sp. nov. | Valid | Prasad | Mio-Pliocene | Churia Formation | Nepal | A species of Lagerstroemia. |
| Laurelia otagoensis | Sp. nov | Valid | Conran, Bannister & Lee | Early Miocene | | New Zealand | A species of Laurelia. |
| Lauroflorum | Gen. et sp. nov. | Valid | Wang, Blanchard & Dilcher | Eocene | Cockfield Formation | United States | A member of Lauraceae. Genus includes new species L. warmanensis. |
| Lindera acuminatissima | Sp. nov | Valid | Dao & Sun in Dao et al. | Late Pliocene | Mangbang Formation | China | A species of Lindera. |
| "Magnolia" szakmanycsabae | Sp. nov. | Valid | Hably | Late Miocene | | Hungary | A plant of uncertain phylogenetic placement, possibly a relative of Magnoliaceae–Lauraceae. |
| Micropetasos | Gen. et sp. nov | Valid | Poinar, Chambers & Wunderlich | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | An early flowering plant of uncertain phylogenetic placement. Genus includes new species M. burmensis. |
| Myrtineoxylon | Gen. et sp. nov. | Valid | Oskolski, Feng & Jin | Late Eocene | Youganwo Formation | China | A member of Myrtaceae described on the basis of fossil wood. Genus includes new species M. maomingensis. |
| Palmoxylon vastanensis | Sp. nov. | Valid | Prasad et al. | Eocene | Cambay Shale Formation | India | A palm tree, a species of Palmoxylon. |
| Paranephelium makumensis | Sp. nov. | Valid | Srivastava & Mehrotra | Oligocene (Chattian) | Tikak Parbat Formation | India | A species of Paranephelium. |
| Poeciloneuron preindicum | Sp. nov. | Valid | Srivastava & Mehrotra | Late Oligocene | | India | A species of Poeciloneuron. |
| Pongamia arjunkholaensis | Sp. nov. | Valid | Prasad | Mio-Pliocene | Churia Formation | Nepal | A species of Pongamia. |
| Potomacapnos | Gen. et sp. nov. | Valid | Jud & Hickey | Early Cretaceous (Aptian) | Potomac Group | United States | A close relative of ranunculalean eudicots. The type specimen is Potomacapnos apeleutheron. |
| Pterygota palaeoalata | Sp. nov. | Valid | Srivastava & Mehrotra | Oligocene (Chattian) | Tikak Parbat Formation | India | A species of Pterygota. |
| Sapindus palaeoemarginatus | Sp. nov. | Valid | Srivastava & Mehrotra | Oligocene (Chattian) | Tikak Parbat Formation | India | A species of Sapindus. |
| Spixiarum kipea | Gen. nov. et sp. nov. | Valid | Coiffard, Mohr & Bernardes-de-Oliveira | Early Cretaceous | Crato Formation | Brazil | Fossil Aroid. |
| Trapa pannonica | Sp. nov. | Valid | Hably | Late Miocene | | Hungary | A water caltrop. |
| Trapa rozsaszentmartoni | Sp. nov. | Valid | Hably | Late Miocene | | Hungary | A water caltrop. |
| Virola dominicana | Sp. nov. | Valid | Poinar & Steeves | Cenozoic (Eocene or Miocene) | | Dominican Republic | A member of Myristicaceae found in Dominican amber, a species of Virola. |
| Vitis szakmanygyorgyi | Sp. nov. | Valid | Hably | Late Miocene | | Hungary | A species of Vitis. |

## Other seed plants

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Donponoxylon | Gen. et 2 sp. nov. | Valid | Tidwell, Britt & Wright | Middle to Late Jurassic | | Australia  New Zealand | A seed plant of uncertain phylogenetic placement. The type specimen is Donponoxylon bennettii; genus also contains Donponoxylon jacksonii. | |
| Dordrechtites arcanus | Sp. nov. | Valid | Bergene, Taylor & Taylor | Middle Triassic (Anisian) | Fremouw Formation | Antarctica | A gymnosperm of uncertain phylogenetic placement. | |

## Other plants

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Acinosporites tristratus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jubah Formation | Saudi Arabia | A trilete spore. |
| Alatisporites? trisacculus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Biornatispora elegantula | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Belgium  Germany  Saudi Arabia | A trilete spore. |
| Biornatispora microclavata | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Brochotriletes crameri | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation Jubah Formation | Saudi Arabia | A trilete spore. |
| Brochotriletes tenellus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Brochotriletes tripapillatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin II Formation | Libya | A trilete spore. |
| Camarozonotriletes asperulus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Awaynat Wanin II Formation Awaynat Wanin III Formation | Brazil  Tunisia | A trilete spore. |
| Chelinospora carnosa | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Chelinospora condensata | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Chelinospora densa | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Chelinospora laxa | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Chelinospora vulgata | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Coronaspora inornata | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Cristatisporites streelii | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin II Formation | Brazil  Tunisia | A trilete spore. |
| Cymbosporites variabilis | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Cymbosporites variegatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin II Formation | Brazil  Libya  Saudi Arabia  Tunisia  United Kingdom | A trilete spore. |
| Cymbosporites wellmanii | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Cyrtospora tumida | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Awaynat Wanin II Formation Awaynat Wanin III Formation | Saudi Arabia  Tunisia | A trilete spore. |
| Diaphanospora milleri | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation Ouan Kasa Formation | Saudi Arabia | A trilete spore. |
| Dibolisporites tuberculatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Awaynat Wanin II Formation Jauf Formation Jubah Formation | Brazil  Libya  Saudi Arabia  Spain  Tunisia  United States | A trilete spore. |
| Dibolisporites verecundus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Dictyotriletes hemeri | Sp. nov | Valid | Breuer & Steemans | Devonian | Jubah Formation | Saudi Arabia  Spain | A trilete spore. |
| Dictyotriletes marshallii | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation Ouan Kasa Formation | Saudi Arabia | A trilete spore. |
| Elenisporis gondwanensis | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Awaynat Wanin II Formation Jubah Formation | Saudi Arabia  Tunisia | A trilete spore. |
| Emphanisporites laticostatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin III Formation Jubah Formation | Brazil  Saudi Arabia  Tunisia | A trilete spore. |
| Emphanisporites plicatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation Jubah Formation | Saudi Arabia | A trilete spore. |
| Franhueberia | Gen. et sp. nov. | Valid | Hoffman & Tomescu | Devonian (Emsian) | Battery Point Formation | Canada | A basal member of Euphyllophytina. The type specimen is F. gerriennei. |
| Geminospora convoluta | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin II Formation Jubah Formation | Saudi Arabia | A trilete spore. |
| Grandispora maura | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Awaynat Wanin II Formation Awaynat Wanin III Formation | Saudi Arabia  Tunisia | A trilete spore. |
| Granulatisporites concavus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Awaynat Wanin II Formation Jauf Formation Jubah Formation | Libya  Saudi Arabia  Spain | A trilete spore. |
| Hystricosporites brevispinus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin II Formation | Tunisia | A trilete spore. |
| Raistrickia commutata | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin II Formation Awaynat Wanin III Formation | Saudi Arabia  Tunisia | A trilete spore. |
| Raistrickia jaufensis | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Retusotriletes atratus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation Jubah Formation | Australia  Saudi Arabia  Spain | A trilete spore. |
| Retusotriletes celatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Samarisporites tunisiensis | Sp. nov | Valid | Breuer & Steemans | Devonian | Ouan Kasa Formation | Tunisia | A trilete spore. |
| Verrucosisporites nafudensis | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Verrucosisporites onustus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation | Saudi Arabia | A trilete spore. |
| Verrucosisporites stictus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation Ouan Kasa Formation | Saudi Arabia | A trilete spore. |
| Zonotriletes brevivelatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Jauf Formation Jubah Formation Ouan Kasa Formation | Algeria  Libya  Saudi Arabia  Tunisia | A trilete spore. |
| Zonotriletes rotundus | Sp. nov | Valid | Breuer & Steemans | Devonian | Awaynat Wanin I Formation Awaynat Wanin II Formation Ouan Kasa Formation | Brazil  Tunisia | A trilete spore. |
| Zonotriletes venatus | Sp. nov | Valid | Breuer & Steemans | Devonian | Jauf Formation Ouan Kasa Formation | Algeria  Brazil  Saudi Arabia  Tunisia | A trilete spore. |