# Classificació dels coleòpters

Aquesta classificació està basada en Families and subfamilies of Coleoptera (with selectes genera, notes, refernces and data on family-group names), de Lawrence i Newton (1995), la més recent i exhaustiva revisió global de la classificació supragenèrica de l'ordre Coleoptera. Com tota classificació se sens dubte provisional; algunes subfamílies han estat considerades fins no fa gaire com a famílies independents (i alguns autors encara ho segueixen fent) o eventualment poden ser elevades a la categoria de família:

## Classificació

ORD. COLEOPTERA LINNAEUS, 1758
Subord. Archostemata Kolbe, 1908
             Fam. Jurodidae Ponomarenko, 1985
             Fam. Ommatidae Sharp & Muir, 1912
                Subfam. Tetraphalerinae Croeson, 1962
                Subfam. Ommatinae Sharp & Muir, 1912
             Fam. Crowsoniellidae Iablokoff-Khnzorian, 1983
             Fam. Micromalthidae Barber, 1913
             Fam. Cupedidae Laporte, 1836
Subord. Myxophaga Crowson, 1955
             Fam. Lepiceridae Hinton 1936 (= Cyathoceridae)
             Fam. Torridincolidae Steffan|, 1964
             Fam. Hydroscaphidae Leconte, 1874
             Fam. Microsporidae Crotch, 1873 (= Sphaeriidae)
Subord. Adephaga Schellenberg, 1806
             Fam. Aspidytidae Ribera,et al., 2002
             Fam. Meruidae Spangler & Steiner, 2005
             Fam. Gyrinidae Latreille, 1810
                Subfam. Spanglerogyrinae Folkerts, 1979
                Subfam. Gyrininae Latreille, 1810
             Fam. Haliplidae Aubé, 1836
             Fam. Trachypachidae C. G. Thomson, 1857
             Fam. Noteridae C. G. Thomson, 1860
                Subfam. Phreatodytinae Uéno, 1957
                Subfam. Noterinae C. G. Thomson, 1860
             Fam. Amphizoidae LeConte, 1853
             Fam. Hygrobiidae Régimbart, 1878 (1837)
             Fam. Dytiscidae Leach, 1815
                Subfam. Copelatinae Van den Branden, 1885
                Subfam. Laccophilinae Gistel, 1856
                Subfam. Hydroporinae Aubé, 1836
                Subfam. Colymbetinae Erichson, 1837
                Subfam. Dytiscinae Leach, 1815
                Subfam. Aubehydrinae Guingot, 1942
             Fam. Rhysodidae Laporte, 1840
             Fam. Carabidae Latreille, 1802
                Subfam. Paussinae Latreille, 1807
                Subfam. Gehringiinae Darlington, 1933
                Subfam. Omophroninae Bonelli, 1810
                Subfam. Carabinae Latreille, 1802
                Subfam. Cicindelinae Latreille, 1802
                Subfam. Hiletinae Schiödte, 1847
                Subfam. Loricerinae Bonelli, 1810
                Subfam. Elaphrinae Latreille, 1802
                Subfam. Migadopinae Chaudoir, 1861
                Subfam. Siagoninae Bonelli, 1813
                Subfam. Scaritinae Bonelli, 1810
                Subfam. Trechinae Bonelli, 1810
                Subfam. Harpalinae Bonelli, 1810
                Subfam. Pseydomorphinae Newman, 1842
                Subfam. Brachininae Bonelli, 1810
Subord. Polyphaga Emery, 1886
       Infraorden Staphyliniformia Lameere, 1900
          Superfamilia Hydrophiloidea Latreille, 1802
             Fam. Hydrophilidae Latreille, 1802
                Subfam. Helophirinae Leach, 1915
                Subfam. Epimetopinae Zaitzev, 1908
                Subfam. Georissinae Laporte, 1840
                Subfam. Hydrochinae C. G. Thomson, 1859
                Subfam. Spercheinae Erichson, 1837
                Subfam. Horelophinae Hansen, 1991
                Subfam. Hydrophilinae Latreille, 1802
                Subfam. Sphaeridiinae Latreille, 1802
             Fam. Sphaeritidae Shuckard, 1839
             Fam. Synteliidae Lewis, 1882
             Fam. Histeridae Gyllenhal, 1808
                Subfam. Niponiinae Fowle, 1912
                Subfam. Trypeticinae Bickhardt, 1914
                Subfam. Trypanaeinae Marseul, 1857
                Subfam. Saprininae Blanchard, 1845
                Subfam. Dendrophilinae Reitter, 1909
                Subfam. Onthophilinae MacLeay, 1819
                Subfam. Tribalinae Bickhardt, 1914
                Subfam. Histerinae Gyllenhal, 1808
                Subfam. Hetaeriinae Marseul, 1857
                Subfam. Chlamydopsinae Bickhardt, 1914
          Superfamilia Staphylinoidea Latreille, 1802
             Fam. Hydraenidae Mulsant, 1844
                Subfam. Prosthettopinae Perkins in Perkins & Balfour-Browne, 1994
                Subfam. Hydraeninae Mulsant, 1844
                Subfam. Ochthebiinae C. G. Thomson, 1859
             Fam. Ptiliidae Erichson, 1845/Motschulsky, 1845 (= Trichopterygidae)
                Subfam. Ptiliinae Erichson, 1845/Motschulsky, 1845
                Subfam. Nanosellinae Barber, 1924
                Subfam. Cephaloplectinae Sharp, 1883
                Subfam. Acrotrichinae Reitter, 1909 (1856)
             Fam. Agyrtidae C.G. Thomson, 1859
             Fam. Leiodidae Fleming, 1821 (= Anisotomidae, Liodidae)
                Subfam. Camiarinae Jeannel, 1911
                Subfam. Catopocerinae Hatch, 1927
                Subfam. Leiodinae Fleming, 1821
                Subfam. Coloninae Horn, 1880 (1859)
                Subfam. Cholevinae Kirby, 1837
                Subfam. Platypsyllinae Ritsema, 1869 (= Leptininae)
             Fam. Scydmaenidae Leach, 1815
                Subfam. Mastiginae Fleming, 1821
                Subfam. Scydmaeninae Leach, 1815
             Fam. Silphidae Latreille, 1807
                Subfam. Silphinae Latreille, 1807
                Subfam. Nicrophorinae Kirby, 1837
             Fam. Staphylinidae Latreille, 1802
                Subfam. Glypholomatinae Jeannel, 1962
                Subfam. Microsilphinae Crowson, 1950
                Subfam. Omaliinae MacLealy, 1825
                Subfam. Empelinae Newton & Thayer, 1992
                Subfam. Proteininae Erichson, 1839
                Subfam. Micropeplinae Leach, 1815
                Subfam. Neophoninae Fauvel, 1905
                Subfam. Dasycerinae Reitter, 1887
                Subfam. Protopselaphinae Newton & Thayer, 1995
                Subfam. Pselaphinae Latreille, 1802
                Subfam. Phloeocharinae Erichson, 1839
                Subfam. Olisthaerinae C.G. Thomson, 1858
                Subfam. Tachyporinae MacLealy, 1825
                Subfam. Trichophyinae C.G. Thomson, 1858
                Subfam. Habrocerinae Mulsant & Rey, 1877
                Subfam. Aleocharinae Fleming, 1821
                Subfam. Trigonurinae Reiche, 1865
                Subfam. Apateticinae Fauvel, 1895
                Subfam. Scaphidiinae Latreille, 1807
                Subfam. Piestinae Erichson, 1839
                Subfam. Osoriinae Erichson, 1839
                Subfam. Oxytelinae Fleming, 1821
                Subfam. Oxyporinae Fleming, 1821
                Subfam. Megalopsidiinae Leng, 1920 (=Stylopodinae)
                Subfam. Steninae MacLealy, 1825
                Subfam. Euaesthetinae C.G. Thomson, 1859
                Subfam. Solieriinae Newton & Thayer, 1992
                Subfam. Leptotyphlinae Fauvel, 1874
                Subfam. Pseudopsinae Ganglbauer, 1895
                Subfam. Paederinae Fleming, 1821
                Subfam. Staphylininae Latreille, 1802
       Infraorden Scarabaeiformia Crowson, 1960
          Superfamilia Scarabaeoidea Latreille, 1802
             Fam. Lucanidae Latreille, 1804
                Subfam. Aesalinae MacLeay, 1819
                Subfam. Nicaginae LeConte, 1861
                Subfam. Syndesinae MacLeay, 1819
                Subfam. Lampriminae MacLeay, 1819
                Subfam. Penichrolucaninae Arrow, 1950
                Subfam. Lucaninae Latreille, 1804
             Fam. Passalidae Leach, 1815
                Subfam. Aulococyclinae Kamp, 1868
                Subfam. Passalinae Leach, 1815
             Fam. Trogidae MacLeay, 1819
             Fam. Glaresidae Semenov Tian-Shanskij & Medvedev, 1932
             Fam. Pleocomidae LeConte, 1861
             Fam. Diphyllostomatidae Holloway, 1972
             Fam. Geotrupidae Latreille, 1802
                Subfam. Bolboceratinae Mulsant, 1842
                Subfam. Geotrupinae Latreille, 1802
                Subfam. Lethrinae Mulsant & Rey, 1871
             Fam. Belohinidae Paulian, 1959
             Fam. Ochodaeidae Mulsant & Rey, 1871
                Subfam. Ochodaeinae Mulsant & Rey, 1871
                Subfam. Chaetocanthinae Scholtz in Scholtz et. al., 1988
             Fam. Ceratocanthidae Cartwright & Gordon, 1971 (= Acanthiceridae)
             Fam. Hybosoridae Erichson, 1847
             Fam. Glaphyridae MacLeay, 1819
             Fam. Scarabaeidae Latreille, 1802
                Subfam. Aphodiinae Leach, 1815
                Subfam. Scarabaeinae Latreille, 1802
                Subfam. Pachypodinae Erichson, 1840
                Subfam. Orphninae Erichson, 1847
                Subfam. Allidiostomatinae Arrow, 1940
                Subfam. Dynamopodinae Arrow, 1911
                Subfam. Aclopinae Milne-Edwards, 1850
                Subfam. Euchirinae Hope, 1840
                Subfam. Phaenomeridinae Erichson, 1847
                Subfam. Melolonthinae Samouelle, 1819
                Subfam. Rutelinae Latreille, 1802
                Subfam. Dynastinae MacLeay, 1819
                Subfam. Cetoniinae Leach, 1815
       Infraorden Elateriformia Crowson, 1960 (= Dascilliformia, Eucinetiformia)
          Superfamilia Scirtoidea Fleming, 1821 (Eucinetoidea)
             Fam. Decliniidae Nikitsky i cols., 1994
             Fam. Eucinetidae Lacordaire, 1857
             Fam. Clambidae Fischer, 1821
                Subfam. Calyptomerinae Crowson, 1955
                Subfam. Acalyptomerinae Crowson, 1979
                Subfam. Clambinae Fischer, 1821
             Fam. Scirtidae Fleming, 1821 (= Helodidae)
          Superfamilia Dascilloidea Guérin-Méneville, 1843 (1834)
             Fam. Dascillidae Guérin-Méneville, 1843
                Subfam. Dascillinae Guérin-Méneville, 1843
                Subfam. Karumiinae Escalera, 1913
             Fam. Rhipiceridae Latreille, 1834
          Superfamilia Buprestoidea Leach, 1815
             Fam. Buprestidae Leach, 1815
                Subfam. Schizopodinae LeConte, 1861
                Subfam. Julodinae Lacordaire, 1857
                Subfam. Buprestinae Leach, 1815
                Subfam. Agrilinae Laporte, 1835
          Superfamilia Byrrhoidea Latreille, 1804 (=Dryopoidea)
             Fam. Byrrhidae Latreille, 1804
                Subfam. Byrrhinae Latreille, 1804
                Subfam. Syncalyptinae Mulsant & Rey, 1869
                Subfam. Amphicyrtinae LeConte, 1861
             Fam. Elmidae Curtis, 1830
                Subfam. Larainae LeConte, 1861
                Subfam. Elminae Curtis, 1830
             Fam. Dryopidae Billberg, 1820 (1817)
             Fam. Lutrochidae Kasap & Crowson, 1975
             Fam. Limnichidae Erichson, 1846
                Subfam. Hyphalinae Britton, 1971
                Subfam. Limnichinae Erichson, 1846
                Subfam. Cephalobyrrhinae Champion, 1925
                Subfam. Thaumastodinae Champion, 1924
             Fam. Heteroceridae MacLeay, 1825
                Subfam. Elythomerinae MacLeay, 1825
                Subfam. Heterocerinae MacLeay, 1825
             Fam. Psephenidae Lacordaire, 1854
                Subfam. Eubriinae Lacordaire, 1857
                Subfam. Psephenoidinae Hinton, 1939
                Subfam. Eubrianacinae Jacobson, 1913
                Subfam. Psepheninae Lacordaire, 1854
             Fam. Cneoglossidae Champion, 1897
             Fam. Anchytarsinae Champion, 1897
                Subfam. Cladotominae Pic, 1914
                Subfam. Apoglossinae Champion, 1897
                Subfam. Araeopidiinae Lawrence, 1991
                Subfam. Ptilodactylinae Laporte, 1836
             Fam. Chelonariidae Blanchard, 1845
             Fam. Eulichadidae Crowson, 1973 (= Lichadidae)
             Fam. Callirhipidae Emden, 1924
          Superfamilia Elateroidea Leach, 1815
             Fam. Artematopodidae Lacordaire, 1857
                Subfam. Electribiinae Crowson, 1957
                Subfam. Artematopodinae Lacordaire, 1857
             Fam. Brachypsectridae LeConte & Horn, 1883
             Fam. Cerophytidae Latreille, 1834
             Fam. Eucnemidae Eschscholtz, 1829 (= Melasidae)
                Subfam. Perothopinae Lacordaire, 1857
                Subfam. Phyllocerinae Reitter, 1905
                Subfam. Pseudomeninae Muona, 1993
                Subfam. Palaeoxeninae Muona, 1993
                Subfam. Phlegoninae Muona, 1993
                Subfam. Melasinae Fleming, 1821
                Subfam. Eucneminae Eschscholtz, 1829
                Subfam. Macraulacinae Fleutiaux, 1923
             Fam. Throscidae Laporte, 1840
             Fam. Elateridae Leach, 1815
                Subfam. Cebrioninae Latreille, 1802
                Subfam. Tetralobinae Laporte, 1840
                Subfam. Thylacosterninae Fleutiaux, 1920
                Subfam. Lissominae Laporte, 1835
                Subfam. Semiotinae Jacobson, 1913
                Subfam. Pityobiinae Hyslop, 1917
                Subfam. Oxynopterinae Candèze, 1857
                Subfam. Agrypninae Candèze, 1857
                Subfam. Denticollinae Stein & Weise, 1877
                Subfam. Negastriinae Nakane & Kishii, 1956
                Subfam. Diminae Candèze, 1863
                Subfam. Cardiophorinae Candèze, 1859
                Subfam. Hemiopinae Fleutiaux, 1941
                Subfam. Physodactylinae Lacordaire, 1857
                Subfam. Eudicronichinae Girard, 1971
                Subfam. Anischiinae Fleutiaux, 1936
                Subfam. Subprotelaterinae Fleutiaux, 1920
             Fam. Plastoceridae Crowson, 1972
             Fam. Drilidae Blanchard, 1845
             Fam. Omalisidae Lacordaire, 1857
             Fam. Lycidae Laporte, 1836
                Subfam. Lycinae Laporte, 1836
                Subfam. Leptolycinae Leng & Mutchler, 1922
                Subfam. Ateliinae Kleine, 1928
                Subfam. Metriorrhynchinae Kleine, 1926
                Subfam. Erotinae LeConte, 1881
                Subfam. Calochrominae Lacordaire, 1857
             Fam. Telegeusidae Leng, 1920
             Fam. Phengodidae LeConte, 1861
                Subfam. Rhagophthalminae Olivier, 1907
                Subfam. Phengodinae LeConte, 1861
             Fam. Lampyridae Latreille, 1817
                Subfam. Pterotinae LeConte, 1861
                Subfam. Ototretadrilinae Crowson, 1972
                Subfam. Cyphonocerinae Crowson, 1972
                Subfam. Ototretinae McDermott, 1964
                Subfam. Amydetinae Olivier, 1907
                Subfam. Lampyrinae Latreille, 1817
                Subfam. Luciolinae Lacordaire, 1857
                Subfam. Photurinae Lacordaire, 1857
             Fam. Omethidae LeConte, 1861
                Subfam. Driloniinae Crowson, 1972
                Subfam. Matheteinae LeConte, 1881
                Subfam. Omethinae LeConte, 1861
             Fam. Cantharidae Imhoff, 1856 (1815)
                Subfam. Cantharinae Imhoff, 1856 (1815)
                Subfam. Silinae Mulsant, 1862
                Subfam. Dysmorphocerinae Brancucci, 1980
                Subfam. Malthininae Kiesenwetter, 1852
                Subfam. Chauliognathinae LeConte, 1861
             Fam. Podabrocephalidae Pic, 1930
             Fam. Rhinorhipidae Lawrence, 1988
       Infraorden Bostrichiformia Forbes, 1926
          Superfamilia Derodontoidea LeConte, 1861
             Fam. Derodontidae LeConte, 1861
                Subfam. Peltasticinae LeConte, 1861
                Subfam. Derodontinae LeConte, 1861
                Subfam. Laricobiinae Mulsant & Rey, 1863-64
          Superfamilia Bostrichoidea Latreille, 1802
             Fam. Dermestidae Latreille, 1804
                Subfam. Dermestinae Latreille, 1804
                Subfam. Marioutinae Jacobson, 1913
                Subfam. Thorictinae Agassiz, 1846
                Subfam. Orphilinae LeConte, 1861
                Subfam. Trinodinae Casey, 1900
                Subfam. Thylodriinae Semenov-Tian-Shanskij, 1913
                Subfam. Attegeninae Laporte, 1840
                Subfam. Megatominae Leach, 1815
             Fam. Endecatomidae LeConte, 1861
             Fam. Bostrichidae Latreille, 1802
                Subfam. Dysidinae Lesne, 1921
                Subfam. Polycaoninae Lesne, 1896
                Subfam. Bostrichinae Latreille, 1802
                Subfam. Psoinae Blanchard, 1851
                Subfam. Dinoderinae C. G. Thomson, 1863
                Subfam. Lyctinae Billberg, 1820
                Subfam. Euderiinae Lesne, 1934
             Fam. Anobiidae Fleming, 1821
                Subfam. Eucradinae LeConte, 1861 (=Hedobiinae)
                Subfam. Ptininae Latreille, 1802
                Subfam. Dryophilinae LeConte, 1861
                Subfam. Ernobiinae Pic, 1912
                Subfam. Anobiinae Fleming, 1821
                Subfam. Ptilininae Schuckard, 1840
                Subfam. Alvarenganiellinae Viana & Martínez, 1971
                Subfam. Xyletininae Gistel, 1856
                Subfam. Dorcatominae C. G. Thomson, 1859
                Subfam. Mesocoelopodinae Mulsant & Rey, 1864
             Fam. Jacobsonidae Heller, 1926
       Infraorden Cucujiformia Lameere, 1938
          Superfamilia Lymexyloidea Fleming, 1821
             Fam. Lymexylidae Fleming, 1821
                Subfam. Hylecoetinae Gistel, 1856
                Subfam. Lymexylinae Fleming, 1821
                Subfam. Melittommatinae Wheller, 1986
          Superfamilia Cleroidea Latreille, 1802
             Fam. Phloiophilidae Kiesenwetter, 1863
             Fam. Trogossitidae Latreille, 1802
                Subfam. Protopeltinae Crowson, 1966
                Subfam. Larinotinae Slipinski, 1992
                Subfam. Peltinae Kirby, 1837 (=Ostominae)
                Subfam. Rentoniinae Crowson, 1966
                Subfam. Decamerinae Crowson, 1964
                Subfam. Lophocaterinae Crowson, 1964
                Subfam. Calitinae Reitter, 1922
                Subfam. Eugoliinae Lacordaire, 1854
                Subfam. Trogossitinae Latreille, 1802 (=Temnochilinae)
             Fam. Chaetosomatidae Crowson, 1952
             Fam. Cleridae Latreille, 1802
                Subfam. Thaneroclerinae Chapin, 1924
                Subfam. Tillinae Leach, 1815
                Subfam. Hydnocerinae Spinola, 1844
                Subfam. Clerinae Latreille, 1802
                Subfam. Epiphloeinae Kuwert, 1893
                Subfam. Enopliinae Gistel, 1856
                Subfam. Tarsosteninae Jacquelin du Val, 1860
                Subfam. Korynetinae Laporte, 1836
             Fam. Acanthocnemidae Crowson, 1964
             Fam. Phycosecidae Crowson, 1952
             Fam. Prionoceridae Lacordaire, 1857
             Fam. Melyridae Leach, 1815
                Subfam. Melyrinae Leach, 1815
                Subfam. Rhadalinae LeConte, 1861
                Subfam. Gietellinae Constantin & Menier, 1987
                Subfam. Dasytinae Laporte, 1840
                Subfam. Malachiinae Fleming, 1821
          Superfamilia Cucujoidea Latreille, 1802 (= Clavicornia)
             Fam. Protocucujidae Crowson, 1954
             Fam. Sphindidae Jacquelin du Val, 1860 (= Aspidiphoridae)
                Subfam. Protosphindidae Sen Gupta & Crowson, 1979
                Subfam. Odontosphindidae Sen Gupta & Crowson, 1979
                Subfam. Sphindiphorinae Sen Gupta & Crowson, 1979
                Subfam. Sphindinae Jacquelin du Val, 1860
             Fam. Brachypteridae Erichson, 1845 (= Cateretidae, Kateretidae)
             Fam. Nitidulidae Latreille, 1802
                Subfam. Calonecrinae Kirejtshuk, 1982
                Subfam. Carpophilinae Erichson, 1842
                Subfam. Meligethinae C. G. Thomson, 1859
                Subfam. Nitidulinae Latreille, 1802
                Subfam. Cillaeinae Kirejtshuk & Audisio in Kirejtshuk, 1986
                Subfam. Cryptarchinae C. G. Thomson, 1859
                Subfam. Cybocephalinae Jacquelin du Val, 1858
             Fam. Smicripidae Horn, 1879
             Fam. Monotomidae Laporte, 1840
                Subfam. Rhizophaginae Redtenbacher, 1845
                Subfam. Monotominae Laporte, 1840
             Fam. Boganiidae Sen Gupta & Crowson, 1966
                Subfam. Paracucujinae Endrödy-Younga & Crowson, 1986
                Subfam. Boganiinae Sen Gupta & Crowson, 1966
             Fam. Helotidae Reitter, 1876/Chapuis, 1876
             Fam. Phloeostichidae Reitter, 1911
                Subfam. Phloeostichinae Reitter, 1911
                Subfam. Hymaeinae Sen Gupta & Crowson, 1966
                Subfam. Agapythinae Sen Gupta & Crowson, 1969
                Subfam. Myraboliinae Lawrence & Britton, 1991
                Subfam. Priasilphinae Crowson, 1973
                Subfam. Tasmosalpinginae Lawrence & Britton, 1991
             Fam. Silvanidae Kirby, 1837
                Subfam. Brontinae Erichson, 1845/Blanchard, 1845
                Subfam. Silvaninae Kirby, 1837
             Fam. Cucujidae Latreille, 1802
             Fam. Laemophloeidae Ganglbauer, 1899
             Fam. Propalticidae Crowson, 1952
             Fam. Phalacridae Leach, 1815
                Subfam. Phaenocephalinae MAtthews, 1899
                Subfam. Phalacrinae Leach, 1815
             Fam. Hobartiidae Sen Gupta & Crowson, 1966
             Fam. Cavognathidae Sen Gupta & Crowson, 1966
             Fam. Cryptophagidae Kirby, 1837
                Subfam. Hypocoprinae Reitter, 1879
                Subfam. Cryptophaginae Kirby, 1837
                Subfam. Atomariinae LeConte, 1861
             Fam. Lamingtonoiidae Sen Gupta & Crowson, 1969
             Fam. Languriidae Crotch, 1873
                Subfam. Xenoscelinae Ganglbauer, 1899
                Subfam. Setariolinae Crowson, 1952
                Subfam. Languriinae Crotch, 1873
                Subfam. Cryptophilinae Casey, 1900
                Subfam. Toraminae Sen Gupta, 1967
             Fam. Erotylidae Latreille, 1802
                Subfam. Dacninae Gistel, 1856
                Subfam. Megalodacninae Sen Gupa, 1969
                Subfam. Encaustinae Crotch, 1876
                Subfam. Tritominae Curtis, 1834
                Subfam. Erotylinae Latreille, 1802
             Fam. Byturidae Jacquelin du Val, 1858
                Subfam. Platydascillinae Pic, 1914
                Subfam. Byturinae Jacquelin du Val, 1858
             Fam. Biphyllidae LeConte, 1861
             Fam. Bothrideridae Erichson, 1845
                Subfam. Teredinae Seidlitz, 1888
                Subfam. Xylariderinae Erichson, 1845
                Subfam. Anommatinae Ganglbauer, 1899
                Subfam. Bothriderinae Erichson, 1845
             Fam. Cerylonidae Bilberg, 1820
                Subfam. Euxestinae Grouvelle, 1908
                Subfam. Loeblioryloninae Slipinski, 1990
                Subfam. Ostomopsonae Sen Gupa & Crowson, 1973
                Subfam. Murmidiinae Jacquelin du Val, 1858
                Subfam. Ceryloninae Bilberg, 1820
             Fam. Alexiidae Imhoff, 1856
             Fam. Discolomatidae Horn, 1878
                Subfam. Notiophyginae Jacobson, 1915
                Subfam. Discolomatinae Horn, 1878
                Subfam. Aphanocephalinae Grouvelle, 1912
                Subfam. Cephalophaninae John, 1954
                Subfam. Pondonatinae John, 1954
             Fam. Endomychidae Leach, 1815
                Subfam. Merophysiinae Seidlitz, 1872
                Subfam. Holoparamecinae Seidlitz, 1888
                Subfam. Leiestinae C. G. Thomson, 1963
                Subfam. Eupsilobiinae Casy, 1895
                Subfam. Endomychinae Leach, 1815
                Subfam. Epipocinae Gorham, 1873
                Subfam. Lycoperdininae Redtenbacher, 1844
                Subfam. Mycetaeinae Jacquelin du Val, 1857
                Subfam. Anamorphinae Strohecker, 1953
                Subfam. Pleganophorinae Jacquelin du Val, 1858
                Subfam. Xenomycetinae Strohecker in Arnett, 1962
             Fam. Coccinellidae Latreille, 1807
                Subfam. Sticholotidinae Weise, 1901
                Subfam. Coccidulinae Mulsant, 1846
                Subfam. Scymninae Mulsant, 1846
                Subfam. Chilocorinae Mulsant, 1846
                Subfam. Coccinellinae Latreille, 1807
                Subfam. Epilachninae Mulsant, 1846
             Fam. Corylophidae LeConte, 1852
                Subfam. Peltinodinae Paulian, 1950
                Subfam. Corylophinae LeConte, 1852
                Subfam. Sericoderinae Matthews, 1888
                Subfam. Parmulinae Poey, 1854
             Fam. Latridiidae Erichson, 1842 (= Lathridiidae)
                Subfam. Latridiinae Erichson, 1842
                Subfam. Corticariinae Curtis, 1829
          Superfamilia Tenebrionoidea Latreille, 1802
             Fam. Mycetophagidae Leach, 1815
                Subfam. Esarcinae Reitter, 1882
                Subfam. Mycetophaginae Leach, 1815
                Subfam. Bergininae Leng, 1920
             Fam. Archeocrypticidae Kaszab, 1964
             Fam. Pterogeniidae Crowson, 1953
             Fam. Ciidae Leach in Samouelle, 1819 (= Cisidae)
                Subfam. Sphindociinae Lawrence, 1974
                Subfam. Ciinae Leach in Samouelle, 1819
             Fam. Tetratomidae Billberg, 1820
                Subfam. Piseninae Miyatake, 1960
                Subfam. Tetratominae Billberg, 1820
                Subfam. Penthinae Lacordaire, 1859
             Fam. Melandryidae Leach, 1915 (= Serropalpidae)
                Subfam. Hallomeninae Mulsant, 1856/Gistel, 1856
                Subfam. Eustrophinae Gistel, 1856
                Subfam. Melandryinae Leach, 1815
                Subfam. Osphyinae Mulsant, 1856 (1840)
             Fam. Mordellidae Latreille, 1802
                Subfam. Ctenididae Franciscolo, 1951
                Subfam. Mordellinae Latreille, 1802
             Fam. Rhipiphoridae Gemminger & Harold, 1870 (1853)
                Subfam. Pelecotominae Seidlitz, 1875
                Subfam. Micholaeminae Vianna, 1971
                Subfam. Ptilophorinae Gerstaecker, 1855
                Subfam. Hemirhipidiinae Heller, 1921
                Subfam. Rhipidiinae Gerstaecker, 1855
                Subfam. Rhipiphorinae Gemminger & Harold, 1870 (1853)
             Fam. Colydiidae Erichson, 1842
                Subfam. Pycnomerinae Erichson, 1845
                Subfam. Colydiinae Erichson, 1842
             Fam. Monommatidae Blanchard, 1845
             Fam. Zopheridae Solier, 1834
                Subfam. Zopherinae Solier, 1834
                Subfam. Usechinae Horn, 1867
             Fam. Ulodidae Pascoe, 1869
             Fam. Perimylopidae St. George, 1939
             Fam. Chalcodryidae Watt, 1974
             Fam. Trachelostenidae Lacordaire, 1859
             Fam. Tenebrionidae Latreille, 1802
                Subfam. Lagriinae Latreille, 1825 (1820)
                Subfam. Phrenapatinae Solier, 1834
                Subfam. Zolodininae Watt, 1974
                Subfam. Pimeliinae Latreille, 1802
                Subfam. Tenebrioninae Latreille, 1802
                Subfam. Alleculinae Laporte, 1840
                Subfam. Diaperinae Latreille, 1802
                Subfam. Coelometopinae Lacordaire, 1859/Schaum, 1859
                Subfam. Cossyphodinae Wasmann, 1899
             Fam. Prostomidae C. G. Thomson, 1859
             Fam. Synchroidae Lacordaire, 1859
             Fam. Oedemeridae Latreille, 1810
                Subfam. Nacerdinae Mulsant 1858
                Subfam. Calopodinae Costa 1852
                Subfam. Oedemerinae Latreille, 1810
             Fam. Stenotrachelidae C. G. Thomson, 1859 (= Cephaloidae)
                Subfam. Stenotrachelinae C. G. Thomson, 1859
                Subfam. Nematoplinae LeConte, 1862
                Subfam. Cephaloinae LeConte, 1862
                Subfam. Stoliinae Nikitsky, 1985
             Fam. Meloidae Gyllenhal, 1810
                Subfam. Eleticinae Wellman, 1910
                Subfam. Meloinae Gyllenhal, 1810
                Subfam. Nemognathinae Laporte, 1840
             Fam. Mycteridae Blanchard, 1845
                Subfam. Mycterinae Blanchard, 1845
                Subfam. Lacconotinae LeConte, 1862
                Subfam. Hemipeplinae Lacordaire, 1854
             Fam. Boridae C. G. Thomson, 1859
                Subfam. Borinae C. G. Thomson, 1859
                Subfam. Synercticinae Lawrence & Pollock, 1994
             Fam. Trictenotomidae Blanchard, 1845
             Fam. Pythidae Solier, 1834
             Fam. Pyrochroidae Latreille, 1807
                Subfam. Tydessinae Nikitsky, 1986
                Subfam. Pilipalpinae Abdullah, 1964
                Subfam. Pedilinae Lacordaire, 1859
                Subfam. Pyrochroinae Latreille, 1807
                Subfam. Agnathinae Lacordaire, 1859 (= Cononotinae)
             Fam. Salpingidae Leach, 1815
                Subfam. Othniinae LeConte, 1862
                Subfam. Prostominiinae Grouvelle, 1914
                Subfam. Agleninae Horn, 1878
                Subfam. Inopeplinae Grouvelle, 1908
                Subfam. Salpinginae Leach, 1815
                Subfam. Aegialitinae LeConte, 1862
                Subfam. Dacoderinae LeConte, 1862
             Fam. Anthicidae Latreille, 1819
                Subfam. Eurygeniinae LeConte, 1862
                Subfam. Lagrioidinae Abdullah & Abdullah, 1968
                Subfam. Afreminae Levey, 1985
                Subfam. Macratriinae LeConte, 1862
                Subfam. Stenoporinae Jacquelin du Val, 1863
                Subfam. Ischaliinae Blair, 1920
                Subfam. Copobaeninae Abdullah, 1969
                Subfam. Lemodinae, Lawrence & Britton, 1991
                Subfam. Tomoderinae Bonadona, 1961
                Subfam. Anthicinae Latreille, 1819
             Fam. Aderidae Winkler, 1927
             Fam. Scraptiidae Mulsant, 1856/Gistel, 1856
                Subfam. Scraptiinae Mulsant, 1856/Gistel, 1856
                Subfam. Anaspidinae Mulsant, 1856
          Superfamilia Chrysomeloidea Latreille, 1802
             Fam. Cerambycidae Latreille, 1802
                Subfam. Vesperinae Mulsant, 1839
                Subfam. Oxypeltinae Lacordaire, 1869
                Subfam. Disteniinae J. Thomson, 1860
                Subfam. Anoplodermatinae Guérin-Méneville, 1840
                Subfam. Philinae J. Thomson, 1860
                Subfam. Parandrinae Blanchard, 1845
                Subfam. Prioninae Latreille, 1802
                Subfam. Spondylidinae Audinet-Serville, 1832
                Subfam. Apatophyseinae Lacordaire, 1869
                Subfam. Necydalinae Latreille, 1825
                Subfam. Lepturinae Latreille, 1802
                Subfam. Cerembycinae Latreille, 1802
                Subfam. Lamiinae Latreille, 1825
             Fam. Megalopodidae Latreille, 1802
                Subfam. Palophaginae Kuschel & May, 1990
                Subfam. Zeugophorinae Böving & Craighead, 1931
                Subfam. Megalopodinae Latreille, 1802
             Fam. Orsodacnidae C. G. Thomson, 1859
                Subfam. Orsodacninae C. G. Thomson, 1859
                Subfam. Aulacoscelidinae Chapuis, 1874
             Fam. Chrysomelidae Latreille, 1802
                Subfam. Sagrinae Leach, 1815
                Subfam. Bruchinae Latreille, 1802
                Subfam. Donaciinae Kirby, 1837
                Subfam. Criocerinae Latreille, 1804
                Subfam. Hispinae Gyllenhal, 1813
                Subfam. Chrysomelinae Latreille, 1802
                Subfam. Galerucinae Latreille, 1802
                Subfam. Lamprosomatinae Lacordaire, 1848
                Subfam. Cryptocephalinae Gyllenhal, 1813
                Subfam. Eumolpinae Hope, 1840
          Superfamilia Curculionoidea Latreille, 1802
             Fam. Nemonychidae Bedel, 1882
                Subfam. Nemonychinae Bedel, 1882
                Subfam. Rhinorhynchinae Voss, 1922
                Subfam. Cimberidinae Gozis, 1882
             Fam. Anthribidae Billberg, 1820
                Subfam. Anthribinae Billberg, 1820
                Subfam. Choraginae W. Kirby, 1819
                Subfam. Urodontinae C. G. Thomson, 1859
             Fam. Belidae Schoenherr, 1826
                Subfam. Belinae Schoenherr, 1826
                Subfam. Oxycoryninae Schoenherr, 1840
                Subfam. Aglycyderinae Wollaston, 1864
             Fam. Rhynchitidae Gistel, 1856
                Subfam. Rhynchitinae Gistel, 1856
             Fam. Attelabidae Billberg, 1820
                Subfam. Attelabinae Billberg, 1820
                Subfam. Apoderinae Jekel, 1860
             Fam. Brentidae Billberg, 1820
                Subfam. Brentinae Billberg, 1820
             Fam. Apionidae Schoenherr, 1823
                Subfam. Apioninae Schoenherr, 1823
             Fam. Nanophyidae Gistel, 1856
                Subfam. Nanophyinae Gistel, 1856
             Fam. Brachyceridae Billberg, 1820
                Subfam. Brachycerinae Billberg, 1820
                Subfam. Ocladiinae Lacordaire, 1866
             Fam. Dryophthoridae Schoenherr, 1825
                Subfam. Dryophthorinae Schoenherr, 1825
                Subfam. Rhynchophorinae Schoenherr, 1833
             Fam. Platypodidae Shuckard, 1840
                Subfam. Platypodinae Shuckard, 1840
             Fam. Erirhinidae Schoenherr, 1825
                Subfam. Erirhininae Schoenherr, 1825
             Fam. Raymondionymidae Reitter, 1913
                Subfam. Raymondionyminae Reitter, 1913
             Fam. Curculionidae Latreille, 1802
                Subfam. Curculioninae Latreille, 1802
                Subfam. Entiminae Schoenherr, 1823
                Subfam. Cyclominae Schoenherr, 1826
                Subfam. Hyperinae Lacordaire, 1863
                Subfam. Molytinae Schoenherr, 1823
                Subfam. Lixinae Schoenherr, 1823
                Subfam. Cryptorhynchinae Schoenherr, 1825
                Subfam. Orobitidinae C. G. Thomson, 1859
                Subfam. Baridinae Schoenherr, 1836
                Subfam. Magdalininae Gistel, 1856
                Subfam. Ceutorhynchinae Gistel, 1856
                Subfam. Campyloscelinae Schoenherr, 1845
                Subfam. Cossoninae Schoenherr, 1825
                Subfam. Scolytinae Latreille, 1807

## Referències

* ↑ Lawrence, J. F. & Newton, A. F., Jr. 1995. Families and subfamilies of Coleoptera (with selectes genera, notes, refernces and data on family-group names). In: Pakaluk i Slipinski (Eds.). Biology, phylogeny and classification of Coleoptera: Papers celebrating the 80th birthday of Roy A. Crowson. Muzeum i Instytut Zoologii PAN, Warszawa. Pp. 779-1006. ISBN 83-85192-34-4
* ↑ Ribera, I, Beutel, R. G., Balke, M. & Vogler, A. P., 2002. Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proc. R. Soc. Lond. B, 269: 2351–2356. Artículo en pdf Arxivat 2007-02-07 a Wayback Machine.
* ↑ Spangler, P. J. & Steiner, W. E. 2005. A new aquatic beetle family, Meruidae, from Venezuela (Coleoptera: Adephaga). Systematic Entomology, 30(3): 339-357