# خيتل

خَيْتَل أو بورتونوس هو جنس من السرطان ويتضمن العديد من الأنواع المهمة للمسمكة، كالسرطان الأزرق السابح و سرطان جازامي.
يحتوي جنس الخيتل على أكثر من 90 نوع، و40 منها معروف من الأحفوريات.
أحفوريات هذه السرطانات في هذا الجنس يمكن ان توجد رواسبًا في أوروبا، الولايات المتحدة، المكسيك، فينزويلا، البرازيل، وأستراليا (مدى العمر يتراوح بين 48.6 مليون إلى 0.0 مليون سنة ماضية)

## الأنواع

قائمة بالأنواع
| النوع | قسم | التاريخ | جنيس |
| Portunus aburatsubo | Balss !(Balss, 1922) | 1922 | Lupocycloporus |
| Portunus acerbiterminalis | Stephenson & Rees !Stephenson & Rees, 1967 | 1967 | Xiphonectes |
| Portunus acuminatus | Stimpson !(Stimpson, 1871) | 1871 | Portunus |
| Portunus affinis | Faxon !(Faxon, 1893) | 1893 | Portunus |
| Portunus alcocki | Nobili !(Nobili, 1905) | 1905 | Xiphonectes |
| Portunus anceps | Saussure !(Saussure, 1858) | 1858 | Portunus |
| Portunus andersoni | De Man !(De Man, 1887) | 1887 | Xiphonectes |
| Portunus angustus | Rathbun, 1898 !Rathbun, 1898 | 1898 | Achelous |
| Portunus arabicus | Nobili !(Nobili, 1905) | 1905 | Xiphonectes |
| Portunus argentatus | Milne-Edwards, A., 1861 !(A. Milne-Edwards, 1861) | 1861 | Monomia |
| Portunus armatus | Milne-Edwards, A., 1861 !(A. Milne-Edwards, 1861) | 1861 | Portunus |
| Portunus asper | Milne-Edwards, A., 1861 !(A. Milne-Edwards, 1861) | 1861 | Portunus |
| Portunus australiensis | Stephenson & Cook !Stephenson & Cook, 1973 | 1973 | Monomia |
| Portunus brevimanus | Faxon !(Faxon, 1895) | 1895 | Achelous |
| Portunus brockii | De Man !(De Man, 1887) | 1887 | Xiphonectes |
| Portunus convexus | De Haan !De Haan, 1835 | 1835 | Portunus |
| Portunus curvipenis | Stephenson !Stephenson, 1961 | 1961 | Monomia |
| Portunus dayawanensis | Chen !Chen, 1986 | 1986 | Xiphonectes |
| Portunus depressiformis | Stimpson !(Stimpson, 1859) | 1859 | Achelous |
| Portunus dubius | Laurie !(Laurie, 1906) | 1906 | Achelous |
| Portunus elongatus | Milne-Edwards, A., 1861 !A. Milne-Edwards, 1861 | 1861 | Achelous |
| Portunus euglyphus | Laurie !(Laurie, 1906) | 1906 | Monomia |
| Portunus floridanus | Rathbun, 1930 !Rathbun, 1930 | 1930 | Achelous |
| Portunus gibbesii | Stimpson !(Stimpson, 1859) | 1859 | Portunus |
| Portunus gladiator | Fabricius !Fabricius, 1798 | 1798 | Monomia |
| Portunus gracilimanus | Stimpson !(Stimpson, 1858) | 1858 | Lupocycloporus |
| Portunus gracillimus | Stimpson !(Stimpson, 1858) | 1858 | Xiphonectes |
| Portunus granulatus | Milne-Edwards, H. !(H. Milne-Edwards, 1834) | 1834 | Achelous |
| Portunus guaymasensis | Garth & Stephenson !Garth & Stephenson, 1966 | 1966 | Achelous |
| Portunus guinotae | Stephenson & Rees !Stephenson & Rees, 1961 | 1961 | Xiphonectes |
| Portunus hainanensis | Chen !Chen, 1986 | 1986 | Xiphonectes |
| Portunus hastatoides | Fabricius !Fabricius, 1798 | 1798 | Xiphonectes |
| Portunus hastatus | Linnaeus !(Linnaeus, 1767) | 1767 | Portunus |
| Portunus inaequalis | Miers !(Miers, 1881) | 1881 | Portunus |
| Portunus innominatus | Rathbun, 1909 !(Rathbun, 1909) | 1909 | Lupocycloporus |
| Portunus iranjae | Crosnier !Crosnier, 1962 | 1962 | Xiphonectes |
| Portunus iridescens | Rathbun, 1894 !(Rathbun, 1894) | 1894 | Achelous |
| Portunus isolamargaritensis | Türkay !Türkay, 1968 | 1968 | Achelous |
| Portunus laevis | Milne-Edwards, A., 1861 !(A. Milne-Edwards, 1861) | 1861 | Lupocycloporus |
| Portunus latibrachium | Rathbun, 1906 !(Rathbun, 1906) | 1906 | Xiphonectes |
| Portunus lecromi | Moosa !Moosa, 1996 | 1996 | Monomia |
| Portunus longispinosus | Dana !(Dana, 1852) | 1852 | Xiphonectes |
| Portunus macrophthalmus | Rathbun, 1906 !Rathbun, 1906 | 1906 | Xiphonectes |
| Portunus madagascariensis | Hoffman !(Hoffman, 1877) | 1877 | Portunus |
| Portunus mariei | Guinot !Guinot, 1957 | 1957 | Xiphonectes |
| Portunus mauricianus | Ward !Ward, 1942 | 1942 | Portunus |
| Portunus minimus | Rathbun, 1898 !Rathbun, 1898 | 1898 | Portunus |
| Portunus minutus | Shen !(Shen, 1937) | 1937 | Lupocycloporus |
| Portunus mokyevskii | Zarenkov !Zarenkov, 1970 | 1970 | Portunus |
| Portunus octodentatus | Gordon !(Gordon, 1938) | 1938 | Achelous |
| Portunus orbicularis | Richters !(Richters, 1880) | 1880 | Achelous |
| Portunus orbitosinus | Rathbun, 1911 !Rathbun, 1911 | 1911 | Achelous |
| Portunus ordwayi | Stimpson !(Stimpson, 1860) | 1860 | Achelous |
| Portunus paralatibrachium | Crosnier !Crosnier, 2002 | 2002 | Xiphonectes |
| Portunus pelagicus | Linnaeus !(Linnaeus, 1758) | 1758 | Portunus |
| Portunus petreus | Alcock !(Alcock, 1899) | 1899 | Monomia |
| Portunus ponticus | Fabricius !(Fabricius, 1798) | 1798 | Monomia |
| Portunus pseudoargentatus | Stephenson !Stephenson, 1961 | 1961 | Monomia |
| Portunus pseudohastatoides | Yang & Tang !Yang & Tang, 2006 | 2006 | Xiphonectes |
| Portunus pseudotenuipes | Spiridonov !Spiridonov, 1999 | 1999 | Xiphonectes |
| Portunus pubescens | Dana !(Dana, 1852) | 1852 | Portunus |
| Portunus pulchricristatus | Gordon !(Gordon, 1931) | 1931 | Xiphonectes |
| Portunus reticulatus | Herbst !(Herbst, 1799) | 1799 | Portunus |
| Portunus reufiarcus | Davie !Davie, 1987 | 1987 | Portunus |
| Portunus rubromarginatus | Lanchester !(Lanchester, 1900) | 1900 | Monomia |
| Portunus rufiremus | Holthuis !Holthuis, 1959 | 1959 | Portunus |
| Portunus rugosus | Milne-Edwards, A., 1861 !(A. Milne-Edwards, 1861) | 1861 | Xiphonectes |
| Portunus samoensis | Ward !(Ward, 1939) | 1939 | Monomia |
| Portunus sanguinolentus | Herbst !(Herbst, 1783) | 1783 | Portunus |
| Portunus sayi | Gibbes !(Gibbes, 1850) | 1850 | Portunus |
| Portunus sebae | Milne-Edwards, H. !(H. Milne-Edwards, 1834) | 1834 | Achelous |
| Portunus segnis | Forskål !(Forskål, 1775) | 1775 | Portunus |
| Portunus serratifrons | Montrouzier !(Montrouzier, 1865) | 1865 | Portunus |
| Portunus sinusodactylus | Stephenson !Stephenson, 1967 | 1967 | Lupocycloporus |
| Portunus spinicarpus | Stimpson !(Stimpson, 1871) | 1871 | Achelous |
| Portunus spiniferus | Stephenson & Rees !Stephenson & Rees, 1967 | 1967 | Xiphonectes |
| Portunus spinimanus | Latreille !Latreille, 1819 | 1819 | Achelous |
| Portunus spinipes | Miers !(Miers, 1886) | 1886 | Xiphonectes |
| Portunus stanfordi | Rathbun, 1902 !Rathbun, 1902 | 1902 | Achelous |
| Portunus stephensoni | Moosa !Moosa, 1981 | 1981 | Xiphonectes |
| Portunus suborbicularis | Stephenson !Stephenson, 1975 | 1975 | Achelous |
| Portunus tenuicaudatus | Stephenson !Stephenson, 1961 | 1961 | Xiphonectes |
| Portunus tenuipes | De Haan !(De Haan, 1835) | 1835 | Xiphonectes |
| Portunus tridentatus | Yang, Dai & Song !Yang, Dai & Song, 1979 | 1979 | Xiphonectes |
| Portunus trilobatus | Stephenson !Stephenson, 1972 | 1972 | Xiphonectes |
| Portunus trituberculatus | Miers !(Miers, 1876) | 1876 | Portunus |
| Portunus tuberculatus | Stimpson !(Stimpson, 1860) | 1860 | Achelous |
| Portunus tuberculosus | Milne-Edwards, A., 1861 !(A. Milne-Edwards, 1861) | 1861 | Xiphonectes |
| Portunus tweediei | Shen !(Shen, 1937) | 1937 | Xiphonectes |
| Portunus ventralis | Milne-Edwards, A., 1879 !(A. Milne-Edwards, 1879) | 1879 | Portunus |
| Portunus wilsoni | Moosa !Moosa, 1981 | 1981 | Lupocycloporus |
| Portunus xantusii | Stimpson !(Stimpson, 1860) | 1860 | Portunus |
| Portunus yoronensis | Sakai !Sakai, 1974 | 1974 | Achelous |