꼭두서니아과

하위 = * 본문 참조 **꼭두서니아과**(----亞科, 학명: 루비오이데아이)는 꼭두서니과의 아과이다.Verdcourt, Bernard. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 280. 1958. 26개 족, 약 7600여 종으로 이루어져 있다.Juss., 1789. *Genera Plantarum* 196.B. Bremer and J.-F. Manen, *Phylogeny and classification of the subfamily Rubioideae (Rubiaceae)* Plant Syst. Evol. 225:43-72 (2000) PDF. 갈퀴덩굴, 계요등, 꼭두서니, 노니, 민둥갈퀴, 솔나물, 수정목, 호자나무 등을 포함하고 있다.
# 하위 분류
; 가이르트네라족(Gaertnereae Endl.)Endlicher, Stephan Friedrich Ladislaus. Genera Plantarum (Endlicher) 1: 577. 1838. * *Gaertnera* Lam. * *Pagamea* Aubl. ; 계요등족(Paederieae DC.)Candolle, Augustin Pyramus de. Prodromus Systematis Naturalis Regni Vegetabilis 4: 342, 470. 1830. * 계요등속(*Paederia* L.) * *Leptodermis* Wall. * *Pseudopyxis* Miq. * *Saprosma* Blume * *Serissa* Comm. ex A.Juss. * *Spermadictyon* Roxb. ; 꼭두서니족(Rubieae Baill.)Baillon, Henri Ernest. Histoire des Plantes 7: 365, 390. 1880 * 갈퀴덩굴속(*Galium* L.) * 갈퀴아재비속(*Asperula* L.) * 꼭두서니속(*Rubia* L.) * 꽃갈퀴덩굴속(*Sherardia* L.) * *Callipeltis* Steven * *Crucianella* L. * *Cruciata* Mill. * *Didymaea* Hook.f. * × *Galiasperula* Ronniger * *Kelloggia* Torr. ex Benth. & Hook.f. * *Mericarpaea* Boiss. * *Microphysa* Schrenk * *Phuopsis* (Griseb.) Hook.f. * *Valantia* L. ; 노니족(Morindeae Kostel.)Kosteletzky, Vincenz Franz. Allgemeine Medizinisch-Pharmazeutische Flora 2: 565. 1833. * 노니속(*Morinda* L.) * *Appunia* Hook.f. * *Coelopyrena* Valeton * *Coelospermum* Blume * *Gynochthodes* Blume ; 다나이스족(Danaideae B.Bremer & Manen)Bremer, Birgitta & Manen, Jean-François. Plant Systematics and Evolution 225: 60. 2000. * *Danais* Comm. ex Vent. * *Payera* Baill. * *Schismatoclada* Baker ; 둔니아족(Dunnieae Rydin & B.Bremer)Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 121. 2009. * *Dunnia* Tutcher ; 무주나무족(Lasiantheae B.Bremer & Manen)Bremer, Birgitta & Manen, Jean-François. Plant Systematics and Evolution 225: 57. 2000. * 무주나무속(*Lasianthus* Jack) * *Ronabea* Aubl. * *Saldinia* A.Rich. ex DC. * *Trichostachys* Hook.f. ; 스크라데라족(Schradereae Bremek.)Bremekamp, Cornelis Eliza Bertus. Recueil des Travaux Botaniques Néerlandais 31: 253. 1934. * *Lecananthus* Jack * *Leucocodon* Gardner * *Schradera* Vahl ; 스키조콜레아족(Schizocoleeae Rydin & B.Bremer)Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 120. 2009. * *Schizocolea* Bremek. ; 스페르마코케족(Spermacoceae Bercht. & J.Presl)Berchtold, Bedřich Všemír von & Presl, Jan Svatopluk. O Prirozenosti Rostlin 256. 1820. * 낚시돌풀속(*Leptopetalum* Hook. & Arn.) * 백령풀속(*Hexasepalum* Bartl. ex DC.) * 백운풀속(*Scleromitrion* (Wight & Arn.) Meisn.) * 제주백운풀속(*Oldenlandia* L.) * 큰백령풀속(*Diodia* L.) * 탐라풀속(*Neanotis* W.H.Lewis) * *Agathisanthemum* Klotzsch * *Amphiasma* Bremek. * *Amphistemon* Groeninckx * *Anthospermopsis* (K.Schum.) J.H.Kirkbr. * *Arcytophyllum* Willd. * *Astiella* Jovet * *Bouvardia* Salisb. * *Carajasia* R.M.Salas, E.L.Cabral & Dessein * *Carterella* Terrell * *Conostomium* (Stapf) Cufod. * *Cordylostigma* Groeninckx & Dessein * *Crusea* Schltdl. & Cham. * *Debia* Neupane & N.Wikstr. * *Denscantia* E.L.Cabral & Bacigalupo * *Dentella* J.R.Forst. & G.Forst. * *Diacrodon* Sprague * *Dibrachionostylus* Bremek. * *Dimetia* (Wight & Arn.) Meisn. * *Edrastima* Raf. * *Eleuthranthes* F.Muell. ex Benth. * *Emmeorhiza* Pohl ex Endl. * *Ernodea* Sw. * *Exallage* Bremek. * *Galianthe* Griseb. * *Gomphocalyx* Baker * *Hedyotis* L. * *Hedythyrsus* Bremek. * *Houstonia* L. * *Hydrophylax* L.f. * *Involucrella* (Benth. & Hook.f.) Neupane & N.Wikstr. * *Kadua* Cham. & Schltdl. * *Kohautia* Cham. & Schltdl. * *Lelya* Bremek. * *Leptomischus* Drake * *Manettia* Mutis ex L. * *Manostachya* Bremek. * *Mexotis* Terrell & H.Rob. * *Mitracarpus* Zucc. * *Mitrasacmopsis* Jovet * *Nesohedyotis* (Hook.f.) Bremek. * *Oldenlandiopsis* Terrell & W.H.Lewis * *Opercularia* Gaertn. * *Pentanopsis* Rendle * *Pentodon* Hochst. * *Phialiphora* Groeninckx * *Phyllocrater* Wernham * *Phylohydrax* Puff * *Planaltina* R.M.Salas & E.L.Cabral * *Pseudonesohedyotis* Tennant * *Psyllocarpus* Mart. & Zucc. * *Richardia* L. * *Sacosperma* G.Taylor * *Schwendenera* K.Schum. * *Spermacoce* L. * *Staelia* Cham. & Schltdl. * *Stenaria* (Raf.) Terrell * *Stenotis* Terrell * *Synaptantha* Hook.f. * *Tessiera* DC. * *Thamnoldenlandia* Groeninckx ; 아르고스템마족(Argostemmateae Bremek. ex Verdc.)Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958. * *Argostemma* Wall. * *Mouretia* Pit. * *Mycetia* Reinw. * *Neohymenopogon* Bennet ; 안토스페르뭄족(Anthospermeae Cham. & Schltdl.)Chamisso, Ludolf Karl Adelbert von & Schlechtendal, Diederich Franz Leonhard von. Linnaea 3: 309. 1828. * *Anthospermum* L. * *Carpacoce* Sond. * *Coprosma* J.R.Forst. & G.Forst. * *Durringtonia* R.J.F.Hend. & Guymer * *Galopina* Thunb. * *Leptostigma* Arn. * *Nenax* Gaertn. * *Nertera* Banks ex Gaertn. * *Normandia* Hook.f. * *Phyllis* L. * *Pomax* Sol. ex DC. ; 오피오리자족(Ophiorrhizeae Bremek. ex Verdc.)Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958. * *Coptophyllum* Korth. * *Keenania* Hook.f. * *Lerchea* L. * *Neurocalyx* Hook. * *Ophiorrhiza* L. * *Spiradiclis* Blume * *Xanthophytum* Reinw. ex Blume ; 우로필룸족(Urophylleae Bremek. ex Verdc.)Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958. * *Amphidasya* Standl. * *Antherostele* Bremek. * *Pauridiantha* Hook.f. * *Pentaloncha* Hook.f. * *Raritebe* Wernham * *Temnopteryx* Hook.f. * *Urophyllum* Jack ex Wall. ; 코우사레아족(Coussareeae Hook.f.)Hooker, Joseph Dalton. Genera Plantarum 2: 9. 1873. * *Bradea* Standl. * *Coccocypselum* P.Browne * *Coussarea* Aubl. * *Cruckshanksia* Hook. & Arn. * *Declieuxia* Kunth * *Faramea* Aubl. * *Heterophyllaea* Hook.f. * *Hindsia* Benth. ex Lindl. * *Oreopolus* Schltdl. ; 콜레토이케마족(Colletoecemateae Rydin & B.Bremer)Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 120. 2009. * *Colletoecema* E.M.A.Petit ; 크녹시아족(Knoxieae Benth. & Hook.f.)Bentham, George & Hooker, Joseph Dalton. Genera Plantarum 2: 9, 21. 1873. * *Batopedina* Verdc. * *Carphalea* Juss. * *Chamaepentas* Bremek. * *Dirichletia* Klotzsch * *Dolichopentas* Kårehed & B.Bremer * *Knoxia* L. * *Lathraeocarpa* Bremek. * *Otiophora* Zucc. * *Otomeria* Benth. * *Paracarphalea* Razafim., Ferm, B.Bremer & Kårehed * *Paraknoxia* Bremek. * *Parapentas* Bremek. * *Pentanisia* Harv. * *Pentas* Benth. * *Phyllopentas* (Verdc.) Kårehed & B.Bremer * *Rhodopentas* Kårehed & B.Bremer * *Triainolepis* Hook.f. ; 크라테리스페르뭄족(Craterispermeae Verdc.)Verdcourt, Bernard. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958. * *Craterispermum* Benth. ; 키아노네우론족(Cyanoneuroneae Razafim. & B.Bremer)Razafimandimbison, Sylvain Georges & Bremer, Birgitta. Taxon 64(2): 294. 2015. * *Cyanoneuron* Tange ; 텔리고눔족(Theligoneae Baill.)Baillon, Henri Ernest. Histoire des Plantes 4: 46, 55. 1872. * *Theligonum* L. ; 팔리코우레아족(Palicoureeae Robbr. & Manen)Robbrecht, Elmar & Manen, Jean-François. Systematics and Geography of Plants 76: 125, 136. 2006. * *Carapichea* Aubl. * *Chassalia* Comm. ex Poir. * *Ditrichanthus* Borhidi, E.Martínez & Ramos * *Eumachia* DC. * *Geophila* D.Don * *Hymenocoleus* Robbr. * *Notopleura* (Hook.f.) Bremek. * *Palicourea* Aubl. * *Puffia* Razafim. & B.Bremer * *Rudgea* Salisb. * *Tromlyca* Borhidi ; 페라마족(Perameae Bremek. ex S.P.Darwin)Bremekamp, Cornelis Eliza Bertus. Taxon 25(5-6): 605. 1976. * *Perama* Aubl. ; 퐁케비아족(Foonchewieae R.J.Wang)Wang, Rui-jiang. Journal of Systematics and Evolution 50(5): 472–473. 2012. * *Foonchewia* R.J.Wang ; 푸토리아족(Putorieae Sweet)Sweet, Robert. Hortus Britannicus, ed. 3 325. 1839. * *Plocama* Aiton ; 프리스마토메리스족(Prismatomerideae Y.Z.Ruan)Ruan, Yun Zhen. Acta Phytotaxonomica Sinica 26(6): 446. 1988. * *Gentingia* J.T.Johanss. & K.M.Wong * *Motleyia* J.T.Johanss. * *Prismatomeris* Thwaites * *Rennellia* Korth. ; 프시코트리아족(Psychotrieae Cham. & Schltdl.)Chamisso, Ludolf Karl Adelbert von & Schlechtendal, Diederich Franz Leonhard von. Linnaea 4(1): 4. 1829. * *Amaracarpus* Blume * *Anthorrhiza* C.R.Huxley & Jebb * *Calycosia* A.Gray * *Dolianthus* C.H.Wright * *Hedstromia* A.C.Sm. * *Hydnophytum* Jack * *Myrmecodia* Jack * *Myrmephytum* Becc. * *Psychotria* L. * *Squamellaria* Becc. * *Streblosa* Korth. ; 호자덩굴족(Mitchelleae Razafim. & B.Bremer)Razafimandimbison, Sylvain Georges & Bremer, Birgitta. Molecular Phylogenetics and Evolution 48(1): 221. 2008. * 호자나무속(*Damnacanthus* C.F.Gaertn.) * 호자덩굴속(*Mitchella* L.) ; 분류 불확실(*incertae sedis*) * *Aphanocarpus* Steyerm. * *Clarkella* Hook.f. * *Coryphothamnus* Steyerm. * *Pagameopsis* Steyerm.
# 계통 분류
2=루쿨리아족