Question: The son of who was a German botanist, horticulturist, and microscopist, who joined the Schleswig-Holstein army to fight the first round of military conflict in southern Denmark and northern Germany rooted in the Schleswig-Holstein Question?
Sentence (from "Johannes Groenland"): Groenland was trained in pharmacology in his youth and served as a pharmacist in Altona, Hamburg, and Jena in his early 20s.
Relevant evidence: No