Question: Who is the paternal grandmother of Despina Khatun?
Sentence (from "Despina Khatun"): Theodora Megale Komnene , also known as Despina Khatun (from the Greek title "Despoina" and Mongol title "Khatun", both meaning "Lady, princess, queen"), was the daughter of John IV of Trebizond, who married the Aq Qoyunlu ruler Uzun Hasan in 1458.
Relevant evidence: Yes