annual biogenic particle fluxes to the interior of the north atlantic ocean studied at 34 n 21 w and 48 n 21 w abstract in order to clarify the annual quality quantity and export processes of biogenic particles from the euphotic zone to the deep ocean interior an array of automated time series sediment traps were deployed for 1 year from 4 april 1989 to 17 april 1990 at 34 n 21 w and 48 n 21 w as part of the joint global ocean flux program jgofs north atlantic bloom experiment nabe three sediment traps with 13 time series sediment collectors were placed at both stations approximately 1 and 2 km below the surface and 0 7 km above the bottom they collected settling particles during 26 14 day intervals for 376 days with an 20 day hiatus in september october 1989 for changeover of the trap moorings the collection periods of the six traps were synchronized forming a spatio temporal matrix of 156 samples the annual mass flux at about 2 km deep during this experiment was 22 and 27 g m 2 y 1 at the 34 and 48 n stations consisting of biogenic particles with traceable quantities of lithogenic particle flux the spring particle bloom characterized by the sedimentation of particles relatively enriched by norg began in january at the 34 n station and in march at the 48 n station the bloom continued for 4 5 and 3 months and provided 62 and 50 of the annual biogenic particle mass flux at 2 km at the 34 and 48 n stations the surface bloom penetrated to the ocean interior within a few weeks with apparently accelerated settling speed at deeper layers the order of susceptibility of biogenic elements to mineralization while settling in the 0 7 1 km a b water column was from least to most resistant p norg corg si and ca the c n p ratio at 0 7 km a b was 154 18 1 at the 34 n station and 148 18 1 at the 48 n station

neritic and pelagic carbonate sedimentation in the marine environment ignorance is not bliss synthesis of available data allows us to define general patterns of late quaternary carbonate production and sedimentation in the global ocean during high stands of sea level the neritic and pelagic environments appear to sequester approximately similar amounts of carbonate whereas during low stands of sea level the decreased neritic zone produces and accumulates approximately an order of magnitude less carbonate assuming that global accumulation of deep sea carbonates remains more or less constant during glacially induced changes in sea level the ocean becomes depleted with respect to calcium carbonate during high stands and recharges during low stands before we can achieve a better understanding of the global carbonate system however we need a better understanding of key environments and processes a production and accumulation on continental shelves both as potential sinks accumulation and as sources export to the deep sea b a better measure of pelagic carbonate production and c late quaternary late pleistocene and holocene mass accumulation rates in the deep sea

fluxes of planktic foraminifera in response to monsoonal upwelling on the somalia basin margin abstract sediment trap samples collected off somalia bi weekly from early june 1992 through mid february 1993 show large seasonal variations in the shell flux and species composition of planktic foraminifera these variations mirror the monsoon driven circulation which resulted in massive upwelling and offshore eddy transport in summer during the sw monsoon june september and accounted for nearly 90 of the estimated annual shell flux of planktic foraminifera g bulloides dominated the sw monsoon along with g glutinata t quinqueloba n dutertrei and t iota in a fauna rich species but with a low diversity and equitability during the autumn intermonsoon november december the ocean became stratified and the nutrient exhausted whereas during the ne monsoon january march a warm surface mixed layer developed with the nutrients entrained by deep wind mixing it was dominated by g ruber in association with g tenella g aequilateralis g trilobus g sacculifer and g menardii in a fauna poorer in species but with a higher diversity and equitability consequently the ratio in the abundance of g bulloides and g ruber follows the monsoonal cycle closely as both species persist throughout the year only the rare g theyeri seems to be restricted to the sw monsoon in october at the end of the sw monsoon the trap at 265 m above the bottom intercepted massive amounts of fine grained carbonate m resuspended from the shelf and upper slope it contained extreme fluxes of bioclastic fragments benthic foraminifera and benthic ostracods as well as enhanced fluxes of small sized planktic foraminifera m notably g rubescens integrated over the year the species composition intercepted by the trap shows a very good match with the shell fauna in a core top sediment from the same site apparently the year estimated monsoonal record of planktic foraminifera which is strongly dominated by the sw monsoon production survives burial intact consequently the species composition of planktic foraminifera and measures such as the bulloides ruber ratio can be applied as a proxy for past changes in the intensity of the sw monsoon off somalia compared to the modern condition

impacts of storms on recent planktic foraminiferal test production and caco3 flux in the north atlantic at 47 n 20 w jgofs planktic foraminiferal assemblages are well known to vary in accordance with seasonal fluctuations in ocean properties periodic reproduction cycles and variations between water masses here we report that storms also can significantly influence foraminiferal assemblages during the rv meteor cruise 21 to the northeast atlantic ocean biotrans area from march to may 1992 planktic foraminifera were sampled using a multiple opening closing net while sampling two storms with wind forces up to 12 beaufort caused intensified surface layer mixing with shifts in the depth of the upper ocean mixed layer from 20 40 m to 170 240 m subsequently planktic foraminiferal growth rates increased resulting in an elevated quantity of small 100 150 m tests phase 1 when the wind strength increased a second time the mixed layer deepened to a depth below the former position of the pycnocline and again the abundance of small tests increased phase 2 during phase 2 the weight of calcite in specimens of the productive zone reached its maximum in the export zone an associated increase in empty tests occurred with a lag time depending on the test sinking velocity in the upper export zone down to 700 m water depth caco3 flux increased from 9 3 to 49 8 mg caco3 m 2 d 1 after the first storm and from 8 9 to 19 9 mg caco3 m 2d 1 after the second storm in the 700 to 2500 m depth interval the flux increased from 5 1 mg caco3 m 2 d 1 to about 9 2 mg caco m 2 d 1 thus the standing stock of living foraminifera and export of empty tests from the productive zone increased after the storms leading to pulses of caco3 exported from the surface to deep water

planktic foraminiferal sedimentation and the marine calcite budget 1 the vertical flux and sedimentation rate of planktic foraminiferal tests are quantified and a global planktic foraminiferal cac0 3 budget is presented test and calcite flux rates are calculated according to the distribution of species obtained from multinet and sediment trap samples modern planktic foraminiferal population dynamics are discussed as a prerequisite for the quantification of the calcite budget highlighting the importance of ecological autecological e g reproduction and biogeochemical conditions that determine the presence or absence of species to complete the open marine particulate caco 3 inventory the contribution of coccolithophores pteropods and calcareous dinophytes is discussed based on the studied regions the global planktic foraminiferal calcite flux rate at 100 m depth amounts to 1 3 3 2 gt yr 1 equivalent to 23 56 of the total open marine caco 3 flux the preservation of tests varies on a regional and temporal scale and is affected by local hydrography and dissolution during most of the year off peak periods many tests dissolve above 700 m water depth while settling through the water column with on average only 1 3 of the initially exported caco 3 reaching the deep seafloor pulsed flux events mass dumps of fast settling particles yield a major contribution of tests to the formation of deep sea sediments on average 25 of the initially produced planktic foraminiferal test cac0 3 settles on the seafloor the total planktic foraminiferal contribution of caco 3 to global surface sediments amounts to 0 36 0 88 gt yr 1 32 80 of the total deep marine calcite budget

comparison of pelagic and nepheloid layer marine snow implications for carbon cycling abstract marine snow from upper and mid water i e pelagic depths on the california margin is texturally and compositionally different from that traveling in the nepheloid layer transmission electron microscopy shows that pelagic marine snow consists primarily of bioclasts e g diatom frustules foram tests organic matter and microbes these components are entrained as discrete particles or small aggregates 10 m in diameter in a loose network of exocellular muco polysaccharide material clays are infrequent but when present are constituents of comparatively compact organic rich microaggregates microbes are abundant and appear to decrease in number with increasing water depth in contrast marine snow aggregates collected from just above the sea floor in the nepheloid layer are assemblages of clay particles clay flocs and relatively dense clay organic rich microaggregates in an exocellular organic matrix bioclasts and microorganisms occur only rarely the prevalence of clay organic rich aggregates in the nepheloid layer suggests that prior to final deposition and burial marine snow from the pelagic zone is subject to disaggregation and recombination with terrigenous detrital material near or at the sea floor results have significant implications for the accumulation and burial rates of organic carbon on continental margins and the aging and bioavailability of sedimentary organic matter samples examined were collected offshore of northern and central california

seasonal changes in species composition numbers mass size and isotopic composition of planktonic foraminifera settling into the deep sargasso sea abstract planktonic foraminifera recovered from six successive sediment trap samples collected over fourteen months at a depth of 3200 m near bermuda show varied responses to seasonal changes in the near surface hydrography these include changes in species composition and within species changes in number of individuals collected per unit time average size and weight of individuals and in oxygen and carbon isotopic composition a total of seventeen species were identified and eleven were sufficiently abundant for isotopic analysis in at least some of the samples the accumulations on the sea floor of some of these species can be expected to contain a strong seasonal component in their oxygen isotopic composition which may be utilized in paleoclimatic and paleo oceanographic reconstructions best suited to this purpose are those species which form their tests in isotopic equilibrium with surface water and whose abundance is confined to a portion of the year only the best examples are pulleniatina o bliguiloculata globigerinoides conglobatus and neogloboquadrina dutertrei globigerinoides ruber is a good estimator of the average surface water temperature because it occurs in rather even abundance throughout the year hastigerina pelagica though very abundant in surface water is practically eliminated even before arrival on the sea floor by partial resorption and structural weakening of the tests during gametogenesis orbulina universa globorotalia truncaltulinoides and globorotalia hirsuta show morphological and istopic evidence for contibnued calcification in deeper water after initial test formation near ther surface globigerina bulloides represents the most striking example of oxygen isotopic disequilibrium with sea water

planktic foraminifers and hydrography of the eastern and northern caribbean sea the distribution of living planktic foraminifers and their relation to the hydrography of the caribbean sea was investigated in plankton net tows and surface sediment samples taken along the antilles island arc during april may 1996 the planktic foraminiferal community was strongly influenced by spatial variations in salinity that were largely due to the influx of orinoco river water into the southeastern caribbean sea and inflowing sargasso sea water in the north along the antilles island arc globigerinoides ruber was the dominant species in the surface waters throughout in the southeastern caribbean sea where orinoco river outflow influences the planktic community standing stocks of planktic foraminifers 100 m between 4 and 50 specimens m 3 were medium to low the southeastern faunas between tobago and guadeloupe were characterized by increased proportions of neogloboquadrina dutertrei highest standing stocks of 159 specimens m 3 in the upper 20 m of the water column were recorded in the northeastern caribbean sea and the assemblages were characterized by high proportions of globigerinita glutinata associated with cyclonic eddies in the anegada passage where sargasso sea water flows into the caribbean sea low standing stocks of 18 specimens m 3 indicate oligotrophic conditions together with the oligotrophic surface waters the subtropical underwater enters the caribbean sea through the anegada passage in water depths between 100 and 300 m these waters are characterized by higher concentrations of globorotalia truncatulinoides relative to the adjacent water masses comparison of the living planktic foraminiferal fauna with empty test assemblages from the water column and from surface sediments shows that differences in the faunal composition mostly correspond to the distribution of water masses and to the differential dissolution of species in the vicinity of islands globigerinoides ruber reaches higher relative frequencies than in the open ocean pointing towards a higher tolerance of this species towards neritic conditions than in other species

trapping efficiencies of sediment traps from the deep eastern north atlantic the 230th calibration bottom tethered sediment traps deployed in the deep eastern north atlantic between 54 n 20 w and 33 n 20 w l1 l2 l3 at the european continental margin at 49 n omex and off the canary islands estoc were investigated for the determination of 230th trapping efficiencies the ratios of 230th flux measured in the traps fa to the expected 230th flux from the production rate of 230th in the overlying water column fp ranged between 0 09 and 1 26 for the traps with deployment periods 300 days the interannual variation of fa fp ratios different years but same location and water depth were up to 10 suggesting that the average 230th flux to the sediment traps did not vary significantly the influence of lateral advection on the 230th flux was taken into account either by applying a mass balance of 230th and 231pa or by assuming a constant removal rate of 230th from the water column an assumption based on similar 230th concentration depth profiles observed at most locations investigated 230th trapping efficiencies were between 9 and 143 showing a trend of increasing efficiencies with increasing water depth no relation was found between current velocities and 230th trapping efficiencies our investigations suggest that the observation of constant or even increasing particle flux rates with increasing water depths in several sediment trap arrays investigated may be a result of sediment trap biases the correction for the trapping biases is important for the understanding of the regional differences in the particle flux in the eastern north atlantic

shell size variation of the planktonic foraminifer neogloboquadrina pachyderma sin in the norwegian greenland sea during the last 1 3 myrs implications for paleoceanographic reconstructions we present measurements of the maximum diameter of the planktonic foraminifer neogloboquadrina pachyderma sin from six sediment cores ocean drilling program sites 643 644 907 909 985 and 987 from the norwegian greenland sea our data show a distinct net increase in mean shell size of n pachyderma sin at all sites during the last 1 3 ma with largest shell sizes reached after 0 4 ma external factors such as glacial interglacial variability and carbonate dissolution alone cannot account for the observed variation in mean shell size of n pachyderma sin we consider the observed shell size increase to mirror an evolutionary trend towards better adaptation of n pachyderma sin to the cold water environment after 1 1 1 0 ma probably the mid pleistocene climate shift and the associated change of amplitude and frequency of glacial interglacial fluctuations have triggered the evolution of this planktonic foraminifer oxygen and carbon stable isotope analyses of diverent shell size classes indicate that the observed shell size increase could not be explained by the functional concept that larger shells promote increasing sinking velocities during gametogenesis for paleoceanographic reconstructions the evolutionary adaptation of neogloboquadrina pachyderma sin to the cold water habitat has significant implications carbonate sedimentation in highest latitudes is highly dependent on the presence of this species in the norwegian greenland sea carbonate poor intervals before 1 1 ma are therefore not necessarily related to severe glacial conditions they are probably attributed to the absence of this not yet polar adapted species further transfer function and modern analog techniques used for the reconstruction of surface water conditions in high latitudes could therefore contain a large range of errors if they were applied to samples older than 1 1 1 0 myrs 2000 elsevier science b v all rights reserved

coccolithophorids on the continental slope of the bay of biscay production transport and contribution to mass fluxes abstract coccoliths collected by sediment traps deployed on the slope of the bay of biscay northeastern atlantic from june 1990 to august 1991 were examined to determine their contribution to the transport of carbonate on a mid latitude continental margin they also were used as tracers of particle transfer processes on this slope two traps located at 1900 m respectively at 2300 mooring site 1 and 3000 m mooring site 2 water depths provided high resolution 4 7 days time series samples covering a 14 month period at ms2 and a 3 month period at ms1 coccoliths from 28 species were identified over the course of the experiment among which emiliania huxleyi was always dominant relative abundance range 59 93 total coccoliths number fluxes were high but variable ranging from 390 106 to 1610 106 coccoliths m 2 day 1 at ms1 and from 58 106 to 1500 106 coccoliths m 2 day 1 at ms2 the time weighted mean flux calculated for the whole experiment at ms2 was 499 106 coccoliths m 2 day 1 estimate of coccoliths minimal contribution to total carbonate flux at 1900 m depth averaged 12 which represented a weighted mean flux of 7 3 mg m 2 day 1 2 7 g m 2 yr 1 lateral transport of coccoliths resuspended from shelf and or upper slope sediments seems to be the dominant transfer process to depth on this northeastern atlantic slope nevertheless the clear seasonal succession observed in the species composition implies that the deposition resuspension transport sequence is rapid presumably less than a few months several short and unsmoothed signals directly issued from coccoliths bloom events also were recorded in our traps a result that indicates rapid settling rates the overall coccolith sedimentation processes appear as being quite diversified but quantitative and qualitative analyses of aggregates collected by the traps suggest that they are important carriers of coccoliths in this margin environment

large extensional structures developed during emplacement of a crystalline thrust sheet the mondonedo nappe nw spain the mondonedo nappe is a crystalline thrust sheet characterized by large recumbent folds regional intermediate pressure metamorphism synkinematic intrusion of granitoids during nappe emplacement and an extensional ductile shear zone developed within the nappe during thrusting a large tectonic window permits the study of the footwall unit revealing another extensional shear zone contemporaneous with thrusting and a low pressure metamorphic evolution in contrast to that of the hanging wall unit the two main extensional shear zones produced e w extension parallel to the direction of orogenic shortening and normal to the orogenic structural trend furthermore subordinate n s longitudinal extension was accommodated by normal faults in the footwall and some of these faults were used as lateral ramps in late stages of thrusting the role of the extensional shear zones and faults described is discussed in the context of an evolving orogenic wedge dominated by plate convergence but characterized by large scale rheological heterogeneities within it deep seated viscous flow triggered by heat accumulation seems to account for the horizontal stretching and probable tapering of the orogenic wedge which was induced by gravitational instabilities due to partial melting and underplating by buoyant continental crust

production and accumulation of calcium carbonate in the ocean budget of a nonsteady state present day production of caco3 in tne world ocean is calculated to be about 5 billion tons bt per year of which about 3 bt accumulate in sediments the other 40 is dissolved nearly half of the carbonate sediment accumulates on reefs banks and tropical shelves and consists largely of metastable aragonite and magnesian calcite deep sea carbonates predominantly calcitic coccoliths and planktonic foraminifera have orders of magnitude lower productivity and accumulation rates than shallow water carbonates but they cover orders of magnitude larger basin area twice as much calcium is removed from the oceans by present day carbonate accumulation as is estimated to be brought in by rivers and hydrothermal activity 1 6 bt suggesting that outputs have been overestimated or inputs underestimated that one or more other inputs have not been identified and or that the oceans are not presently in steady state one missing calcium source might be groundwater although its present day input is probably much smaller than that of rivers if as seems likely caco3 accumulation presently exceeds terrestial and hydrothermal input this imbalance presumably is offset by decreased accumulation and increased input during lowered sea level shallow water accumulation decreases by an order of magnitude with a 100 m drop in sea level while groundwater influx increases because of heightened piezometric head and the diagenesis of metastable aragonite and magnesian calcite from subaerially exposed shallow water carbonates

using pliocene palaeoclimatic data to postulate dispersal pathways of early hominins abstract a geographic information system gis simulation is used here to test postulated early hominin expansions into eurasia to understand fully plio pleistocene hominin dispersals that is the increasing geographic range of hominins a number of interacting factors must be examined the most important variables affecting the first hominin dispersals would have been environmental and it is thus imperative that appropriate climatic and geographical factors are included in analyses of possible plio pleistocene hominin expansions as the late pliocene climate was substantially different from that of today the present model utilises the pliocene research interpretation and synoptic mapping middle pliocene paleoenvironmental reconstruction prism2 dowsett h j barron j a poore r z thompson r s cronin t m ishman s e willard d a 1999 middle pliocene paleoenvironmental reconstruction prism2 u s geological survey http pubs usgs gov of 1999 of99 535 accessed 3 02 00 the combination of data from palaeoclimate reconstructions and existing archaeological and palaeontological sites sheds light on the out of africa 1 problem suggesting that expansion of hominins out of africa would have been most feasible from a palaeoclimatic perspective at and around the plio pleistocene boundary the model predicts that the most likely dispersal route out of africa would have been through the levantine region but a lack of research in the region of the bab el mandeb straits means that this alternative pathway should not be ignored there is also a small possibility of dispersals across the mediterranean sea but with expansion limited to the northern coastline further easterly expansion of the hominin range is likely to have occurred through the more northerly countries of central asia specifically georgia ukraine russia and kazakhstan a hominin dispersal route thus existed from africa through central asia to china during the plio pleistocene period following the grassland vegetation belt that was present in this region at the time

interannual variability of planktic foraminiferal populations and test flux in the eastern north atlantic ocean jgofs planktic foraminiferal assemblages vary in response to seasonal fluctuations of hydrographic properties between water masses and after periodical changes and episodic events e g reproduction storms distinct annual variability of the planktic foraminiferal flux is also known from sediment trap data in this paper we discuss the short term impacts on interannual flux rates based on data from opening closing net hauls obtained between the ocean surface and 500 m water depth data were recorded during april may june and august at around 47 n 20 w biotrans in 1988 1989 1990 1992 1993 and during may 1989 and 1992 at 57 n 20 22 w species assemblages closely resemble each other when comparing the mixed layer fauna with the fauna of the upper 100 m and the upper 500 m of the water column in addition species assemblages 100 m are almost indistinguishable from assemblages that are 125 m in test size the standing stock of planktic foraminifers at biotrans can vary by more than one order of magnitude over different years however species assemblages may be similar when comparing corresponding seasons early summer assemblages june are distinctly different from late summer assemblages august significant variations in the species composition during spring april may are independent of the mixed layer depth spring assemblages are characterized by high numbers of globigerinita glutinata in particular day to day variations of the number of specimens and in species composition may have the same order of magnitude as interannual variations this appears to be independent of the reproduction cycle species assemblages at 47 n and 57 n are similar during spring although surface water temperatures and salinities differ by up to 10 c and 0 7 psu we suggest that the main factors controlling the planktic foraminiferal fauna are the trophic properties in the upper ocean productive layer planktic foraminiferal carbonate flux as calculated from assemblages reveals large seasonal variations a quasi annual periodicity in flux levels and substantial differences in timing and magnitude of peak fluxes at the biotrans station the average annual planktic foraminiferal caco 3 fluxes at 100 and 500 m depth are estimated to be 22 4 and 10 0 g m 2 yr 1 respectively

an atlas of the distribution of calcium carbonate in sediments of the deep sea historical observations of the concentration of calcium carbonate in global deep sea sediments are compiled and compared with a new gridded field of seawater co3 concentration to reveal regional variations in the calcite lysocline the most obvious mode of variability of the calcite lysocline is the thickness of the lysocline defined here as the difference in overlying water carbonate saturation co3 between high and low calcite sediments with a thicker lysocline in the atlantic than in the pacific i attribute this variation to differences in the delivery rate of terriginous material a recent model for the lower glacial atmospheric pco2 proposed to change the relationship between the depth of the lysocline and the co3 of the water column by changing the rain rate ratio of organic carbon to calcite production the rain ratio model archer and maier reimer 1994 i search the data set for analogs to the proposed glacial world by looking for a link between the regional climate at the sea surface and the depth of the lysocline below the co3 at the carbonate compensation depth ccd in the tropics appears to be 10 20 mol kg 1 co3 more undersaturated than in high latitudes but this is smaller than the 40 mol kg 1 shift required by the model in addition the general resemblance of the glacial lysocline to the present day requires that the proposed shift in co3 at the ccd be globally uniform rather than locally variable as climate forcing would probably generate i conclude that the rain ratio model would probably require some globally uniform perturbation during glacial time such as a change in ocean si content if it is to explain the entire pco2 decrease observed in the glacial atmosphere finally i grid the sedimentary data to estimate that the inventory caco3 which is available to neutralize fossil fuel co2 is approximately 1600 gt carbon a quantity which may be exceeded by fossil fuel release in the next several centuries

a calcite dissolution pulse in the norwegian greenland sea during the last deglaciation carbonate preservation profiles in the atlantic ocean and the norwegian greenland sea seem to be out of phase during the last deglaciation a possibly time transgressive deglacial preservation spike in the n atlantic overlaps with a major calcite dissolution pulse in the norwegian sea contemporaneously major changes in surface and bottom water circulation of the norwegian greenland sea occurred isotopic and sedimentological evidence suggests that bottom water formation in the norwegian greenland sea was almost shut down during the last maximum glaciation and probably also during the first part of the last deglaciation during that time corrosive bottom waters might have filled the norwegian sea deep sea basins causing carbonate dissolution at the sea floor subsequently the reinitiation of deep water formation could have been coupled with increased resuspension of organic matter and or reworking of older organic matter rising the p co2 of bottom waters and contributing to carbonate dissolution at the sea floor additionally large volumes of atmospheric co2 stored before the younger dryas might have been pumped into the deep sea possibly by downwelling of surface waters and been neutralized against carbonate at the sea floor

quaternary paleo oceanography pteropod preservation and stable isotope record of the red sea abstract an abundance of pteropod species their mode of preservation and stable isotope ratios of carbon and oxygen in benthic and planktic foraminifera have been determined in core md76 140 from the northern red sea isotopic data and benthic foraminiferal species indicate changes in o2 concentration in the bottom sediments the glacial stages are characterized by an almost anoxic bottom while the interglacial bottom was generally well ventilated in general very well preserved pteropod shells occur in glacial oxygen depleted intervals and also on recent well ventilated bottoms in the interglacial stages the preservation deteriorates and in the most extreme cases there is a complete dissolution of the aragonitic shells in these cases most of the assemblage is lost and the only traces of the original shells are their internal molds these are cemented with high magnesium calcite hmc which is a more stable carbonate phase in bottom sediments of the interglacial dissolution of aragonite discriminates between large and small shells the latter tend to be removed completely while the larger ones may be preserved as internal hmc molds limacina inflata is more readily molded than l trochiformis and thus its relative abundance increases in poorly preserved assemblages the state of aragonitic shell preservation depends on the degree of organic matter combustion in the sediments well preserved shells occur in sediments where the supply of organic matter is limited and o2 levels are high preservation deteriorates with increasing supply of organic carbon o2 consumption and increasing production of co2 in this case the dissolved carbonate is reprecipitated as hmc cement with further increase in organic matter supply and reduced ventilation nearly all oxygen is consumed and no3 and subsequently so42 become the oxidizing agents this is followed by an increasing carbonate alkalinity and excellent aragonitic shell preservation

a high resolution record of late quaternary upwelling along the oman margin arabian sea based on planktonic foraminifera planktonic foraminiferal abundances fluxes test sizes and coiling properties are influenced in various ways by the southwest monsoon winds and associated upwelling in the western arabian sea to determine the short term changes in the southwest monsoon we have carried out a high resolution time series analysis of three upwelling indices total flux of planktonic foraminiferal tests and flux and relative abundance of the planktonic foraminiferal species globigerina bulloides from ocean drilling program odp site 723a oman margin western arabian sea spanning the last 19 kyr in addition we have determined the relationships between upwelling intensity and the relative abundance fluxes and shell concentrations of various planktonic foraminiferal species upwelling indices suggest that from 19 to 16 ka 22 to 18 2 cal kyr b p the sw monsoon was relatively strong compared to the period 15 8 to 12 5 ka 17 8 to 13 8 cal kyr b p the intensification of the sw monsoon took place at 12 ka 13 1 cal kyr b p and reached a peak between 10 and 5 ka 10 6 and 4 8 cal kyr b p the high resolution data further demonstrate that the sw monsoon has started weakening from 5 ka 4 8 cal kyr b p and the weakest phase was in place at 3 5 ka 3 cal kyr b p which coincides with evidence of an arid climate in western tibet fluxes and shell concentrations of many of the planktonic foraminiferal species increased between 12 and 5 ka in response to the intensification of the sw monsoon winds after the last glacial period globigerina bulloides shows a fivefold to tenfold increase in flux during this period of intense upwelling the other species whose fluxes are influenced by this upwelling change are in order from strongest to weakest response globigerinita glutinata globigerinoides ruber neogloboquadrina dutertrei globigerinella aequilateralis globigerina falconensis and globigerinoides sacculifer the relative abundances of g bulloides and g ruber increased during intense upwelling whereas the relative abundances of g glutinata n dutertrei g falconensis and g sacculifer did not increase during this period which might be due to differences in the productivity of various species in relation to upwelling change therefore the fluxes and shell concentrations provide better and more reliable information about the changes in the monsoon system in the arabian sea than relative abundance data

the fate of coccoliths at 48 n 21 w northeastern atlantic abstract as part of the joint global ocean flux study program jgofs north atlantic bloom experiment nabe coccolithophore fluxes were recorded over one year period from april 1989 to april 1990 three vertically moored sediment traps synchronized on a 14 day interval were deployed at 48 n 21 w at three water depths at 1 2 and 3 7 km sea floor at 4 4 km we present the results on the vertical variability of coccolith and coccosphere fluxes through the water column calcium carbonate was the largest component of the total mass flux followed by particulate organic matter and opal coccolith and coccosphere seasonal fluxes followed the seasonality of the biogeochemical particle fluxes at all three depths with maxima occurring during the spring particle bloom the decrease in correlation between coccosphere and coccolith fluxes with depth from r 0 98 at 1 km depth to 0 60 at 3 7 km depth documents that disaggregation of the coccospheres occurs during settling in the deeper part of the water column between 1 and 3 7 km water depth annual coccosphere and coccolith fluxes were 1 5 108 and 0 9 10 11 m 2 year 1 respectively at 1 km depth or 3 1 108 and 1 4 10 11 m 2 year 1 including the spring 1990 episode for the 3 7 km trap the corresponding values were 0 5 108 and 1 1 1011 0 6 108 and 1 1 1011 including the partially recorded spring 1990 episode the slight increase in coccolith numbers with water depth can be explained by coccosphere disintegration and lateral influx nevertheless selective dissolution of delicate coccolith species from 1 to 3 7 km water depth occurs e g relative abundances of oolithothus fragilis coccoliths decrease from 3 at 1 km to less than 0 5 at 3 7 km water depth emiliania huxleyi gephyrocapsa muellerae calcidiscus leptoporus and coccolithus pelagicus constitute from 65 at 1 km depth to 95 at 3 7 km depth of the total coccolith assemblages the same four coccolith taxa dominate the fossil assemblages in the underlying surface sediments selective dissolution continues in the sediments most notably the abundance of the solution resistant species c leptoporus increases from 10 to 15 in the trap samples to 15 40 in the sediments annual flux of calcium carbonate in sediment trap samples accounts for about 60 of the total annual mass flux with the coccolith fraction m being about 45 of the annual carbonate flux mean value of 41 at 1 km and 50 at 3 7 km water depth the correlation between the m caco 3 and the calculated coccolithophore and calcisphere caco3 fluxes decreases with depth r 0 92 at 1 km and r 0 86 at 3 7 km indicating that partial dissolution and fragmentation of the delicate less calcified coccolithophore and planktic foraminifera species occurs during settling this fine fragmentation of biogenic caco3 can contribute to the m fraction causing the decrease of correlation with the calculated caco3 fluxes being based only on the flux of intact coccoliths and calcispheres

planktonic foraminiferal flux to the deep ocean sediment trap results from the tropical atlantic and the central pacific abstract the flux of planktonic foraminifera between 100 m and 1 mm to the seafloor has been estimated for the central pacific abyssal plain east of hawaii and the tropical atlantic demerara abyssal plain based on sediment trap samples collected from various water depths the faunas in each region are quite similar with 4 to 5 species generally accounting for 70 80 of the total assemblage at both study sites the total foraminiferal flux and the carbonate flux tend to decrease with depth in addition the flux of individual species of planktonic foraminifera varies significantly with depth with the number of small solution susceptible species decreasing with increasing water depth these results suggest that there is significant dissolution of small m foraminifera as they settle through the water column material collected from the sediment water interface directly below the pacific sediment trap array contains no planktonic foraminifera suggesting that the residence time of an individual on the seafloor before it dissolves is extremely short

coccolithophore export production species composition and coccolith caco3 fluxes in the ne atlantic and abstract coccolithophores were examined in two sediment traps deployed for one year april 1989 to april 1990 at approximately 1 km water depth in the subtropical ne atlantic at 34 n 21 w station nabe 34 and in the temperate ne atlantic 48 n 21 w station nabe 48 as part of the jgofs north atlantic bloom experiment nabe at both stations maximum coccolith and coccosphere fluxes were recorded during local spring bloom periods the average coccolith and coccosphere fluxes at nabe 34 were 1 12 10 9 m 2 d 1 and 4 90 10 6 m 2 d 1 respectively at this station coccoliths from 51 taxa were identified a marked seasonal change was noticed in relative abundances of emiliania huxleyi and deep photic zone species e huxleyi dominated during spring while in late autumn the coccolith assemblage was dominated by florisphaera profunda and the coccosphere assemblage by g flabellatus and algirosphaera spp emiliania huxleyi was the most abundant species in the overall coccolith and coccosphere assemblages with 69 and 64 respectively at nabe 48 the average daily fluxes were significantly lower with 3 88 10 8 coccoliths m 2 d 1 and 8 45 10 5 coccospheres m 2 d 1 and only 36 taxa were recorded in the trap samples emiliania huxleyi was the dominant species in the coccolith assemblage throughout the year with 72 and showed maximum fluxes during spring 1990 coccolithus pelagicus exhibited a distinct seasonal pattern with maximum fluxes in early june 1989 probably related to the development of a mesoscale cyclonic eddy in the vicinity of the trap site holococcolithophore fluxes were highest during maximum sea surface temperatures in summer the coccosphere assemblage was dominated by gephyrocapsa spp syracosphaera spp and e huxleyi estimated caco3 fluxes of coccoliths coccospheres and calcareous dinophytes were compared with the caco3 content in the fine m fraction the annual mass estimated caco3 flux of coccoliths and coccospheres was 5 1 g m 2 yr at nabe 34 and 2 7 g m 2 yr at nabe 48 and that of calcareous dinophytes 0 9 and 0 1 g m 2 yr respectively these summed mass estimated values contributed on average only 55 at nabe 34 and 60 at nabe 48 to the fine fraction caco3

sedimentation in the western arabian sea the role of coastal and open ocean upwelling monsoon induced coastal and open ocean upwelling explain 84 of the variations of the organic carbon fluxes measured in the deep western arabian sea in this paper sea level measurements satellite derived wind speeds sea surface temperatures and nutrient profiles are used to discern the relative importance of these factors on fluxes measured during nine years of continuous sediment trap deployments this exercise shows i the increase in fluxes observed during the initial stages of the sw monsoons are caused by open ocean upwelling which develops faster than the coastal upwelling ii coastal upwelling triggers diatom blooms from nutrients from subsurface water and sediment resuspension and more importantly by injecting resting stages of diatoms back into the euphotic zone iii silica depletion resulting from diatom blooms in laterally advecting water masses leads to a replacement of diatoms by other nitrate limited organisms iv organic carbon fluxes to the deep arabian sea increase in response to an intensification of both coastal and open ocean upwelling weak coastal upwelling and strong open ocean upwelling also increase organic carbon fluxes the varying dominance of their influence is reflected in the timing and the composition of the peak fluxes v the link between organic carbon flux and monsoon strength is non linear probably due to changes in the surface currents and to vigorous turbulence in the surface water during strong sw monsoons these processes could reduce the organic carbon flux in the western arabian sea by about 65

export production of coccolithophores in an upwelling region results from san pedro basin southern california borderlands a seven month long time series sediment trap project was carried out in san pedro basin southern california borderlands in order to evaluate the response of calcareous nannoplankton to seasonal hydrographic changes this region is periodically influenced by upwelling particularly during the spring and early summer the highest fluxes of both whole coccospheres and individual coccoliths occurred during winter january february a period when the fluxes of diatoms and planktic foraminifera were low the highest coccolithophore fluxes were recorded in the mid february with 860 106 coccoliths m 2 day 1 8 106 whole coccospheres m 2 day 1 and 80 mg of coccolith carbonate m 2 day 1 coccolith carbonate fluxes in january and february account for most of the total carbonate fluxes measured during this period the season of maximum coccolithophore production in this region winter is correlated with weak stratification of the upper water column low total primary production low nutrient contents and low temperatures emiliania huxleyi and florisphaera profunda are the two most abundant species in this region while e huxleyi displays no distinct seasonal changes in flux f profunda shows a clear preference for cold low nutrient water conditions and low light levels helicosphaera spp flux is positively correlated to the total coccosphere fluxes and is indicative of high coccolithophore productivity

global deep sea burial rate of calcium carbonate during the last glacial maximum global databases concentrations rates in holocene and last glacial of calcium carbonate and mass accumulation maximum sediments were used to estimate the deep sea sedimentary calcium carbonate burial rate during these two time intervals sparse calcite mass accumulation rate data were extrapolated across regions of varying calcium carbonate concentration using a gridded map of calcium carbonate concentrations and the assumption that accumulation of noncarbonate material is uncorrelated with calcite concentration within some geographical region mean noncarbonate accumulation rates were estimated within each of nine regions determined by the distribution and nature of the accumulation rate data for core top sediments data coverage 67 of the high calcite the regions of reasonable encompass 75 sediments globally and within these regions we estimate an accumulation rate of 55 9 3 6x1011 mol yr 1 the same regions cover 48 of glacial high caco3 sediments the smaller fraction is due to a shift of calcite deposition to the poorly sampled south pacific and total 44 1 6 0x1011 mol yr 1 projecting to 100 coverage south and total 44 1 6 0x1011 both estimates yields accumulation estimates of 8 3x1012 mol yr 1 today and 9 2x1012 mol yr 1 during glacial time this is little better than a guess given the incomplete data coverage but it suggests that glacial deep sea calcite burial rate was probably not considerably faster than today in spite of a presumed decreasein shallow water burial during glacial time

seasonal sedimentation and stable isotope records of pteropods off cap blanc abstract sediment trap studies conducted off cap blanc mauretania during 1989 1990 showed large seasonal flux variations of pteropods limacina inflata clearly dominated pteropod flux in terms of numbers and mass veligers of this species had a prominent peak of about 10 600 ind m 2 day 1 in boreal summer during the period of highest organic carbon flux the flux of adult individuals 1 mm peaked about 3 weeks earlier 312 ind m 2 day 1 aragonite produced by this size fraction of l inflata then amounted to about 75 of total carbonate clio pyramidata the second most abundant species showed less tendency toward seasonal variation and peaked in spring 43 ind m 2 day 1 although many pteropods undergo vertical migrations of several hundred meters the oxygen isotope compositions of l inflata and c pyramidata shells appear to reflect water temperatures of the species shallower limits 50 120 m since most larger pteropods have life spans of approximately one year shells derived from sedimentary pteropod layers may record past mean annual surface water temperatures a few specimen of c pyramidata with shell lengths from 8 5 to 13 mm were collected in august 1989 and march 1990 and were cut into several segments and analyzed for stable isotope composition although growth is discontinous c pyramidata appears to record summer to winter temperature fluctuations in its upper migration range around 120 m thus larger specimens taken from pteropod rich sedimentary layers could provide past seasonal variations in surface or subsurface water temperatures there appears to be a substantial enrichment of 13c in precipitated aragonite in l inflata 250 m size relative to bicarbonate we suggest that the 13c values in pteropod shells are primarily controlled by the isotope ratio of total dissolved co2 and by seasonal temperature variations the stable carbon isotope values of l inflata covary with the oxygen isotope values and thus correlate negatively with water temperature 13c 3 41 0 10t our data confirm the negative temperature dependency reported in the literature a four year seasonal sedimentation record for the cap blanc site g wefer and g fischer unpubl data shows only one distinct pteropod sedimentation event in august 1989 this indicates strong interannual variability of pteropod production and sedimentation the distinct maximum in august 1989 is probably related to episodically occurring unusually high surface water temperatures similar conditions might have been present during the sedimentation of pteropod rich layers described from the deglaciation period and holocene maximum in the atlantic ocean

impact of the azores front on the distribution of planktic foraminifers shelled gastropods and coccolithophorids abstract the azores front current system south of the azores islands has been studied in order to reveal the direct impact of an open oceanic thermohaline front on the distribution of the calcareous plankton planktic foraminifers pteropods heteropods and coccolithophorids were sampled from the upper 2500 m of the water column during august 1997 and january 1999 in addition the hydrography was recorded across the frontal jet on combined ctd and xbt transects in august 1997 a strong seasonal thermocline capped the azores front af at about 60 90 m water depth below the thermocline a distinct hydrographic front was indicated by temperature and salinity gradients the central af was an area of low planktic foraminiferal gastropod and coccolithophorid production and was a faunal barrier for shallow and deep dwelling species highest numbers of planktic foraminifers gastropods and coccolithophores were recorded from above the thermocline north of the af the most frequent planktic foraminiferal and pteropod species were globigerinoides ruber white and limacina inflata respectively below 100 m planktic foraminifers were most frequent north of the af and gastropod shells were rare in particular the deep dwelling planktic foraminifer globorotalia scitula was frequent only at sampling sites north of the af in january 1999 the surface water temperature of the azores front current system was lower and the thermocline was deeper than in august 1997 the planktic foraminiferal standing stock was three times higher than in august 1997 and no water depth related faunal changes occurred the fauna was dominated by globorotalia truncatulinoides and turborotalita humilis was frequent during both january and august the fauna from south of the azores was different from that to the southeast recording the occurrence of two different water masses in this study we present a micropaleontological definition of the af in order to provide a paleoceanographic tool that may be used to decipher the late quaternary current system of the north atlantic ocean