Given the input sentence, please extract the subject and object containing a certain relation in the sentence according to the following relation types, in the format of "relation1: word1, word2; relation2: word3, word4". Relations include: product/material produced; manufacturer; distributed by; industry; position held; original broadcaster; owned by; founded by; distribution format; headquarters location; stock exchange; currency; parent organization; chief executive officer; director/manager; owner of; operator; member of; employer; chairperson; platform; subsidiary; legal form; publisher; developer; brand; business division; location of formation; creator.

Chris MacDonald, a business ethics expert at Ryerson University in Toronto who oversaw a case study about Google’s operations in China while at Duke University, says Chinese regulators will see Play as less threatening than Search and Gmail, reducing the frequency of government-led probes.

Answer: headquarters_location: Ryerson University, Toronto