He was the first biomathematician although he followed in the tradition of another great natural historian with mathematical skills, namely Buffon . His understanding of mathematics was of the modern subject but based on the firm foundations of an understanding of Greek mathematics.
Siya ay ang unang biomathematician bagama't siya sinundan sa tradisyon ng isa pang magaling na likas na istoryador sa mga kasanayan sa matematika, namely Buffon. Kanyang-unawa ng matematika ay ang modernong paksa ngunit batay sa matatag pundasyon ng isang unawa ng Greek matematika.