His principal contributions to science consisted in his anthropological inquiries, especially into the laws of heredity, where the distinguishing feature of his work was the application of statistical methods.
Punong-guro ang kanyang mga kontribusyon sa science consisted sa kanyang antropolohiko mga katanungan, lalo na sa mga batas ng pagmamana, kung saan ang tumutukoy na tampok ng kanyang trabaho ay ang application ng statistical methods.