puppigerus is an extinct genus of sea turtle from the eocene .

it is known from finds in the united states , england , belgium , denmark , and uzbekistan .

puppigerus

the taxonomy and morphology of puppigerus camerpi [ gray ] , an eocene sea - turtle from northern europe .
palatal view of plastic cast of puppigerus camperi , irsnb ig8402 / r . 17 . exhibiting development of ' spatulate’ palatines
puppigerus nessovi : zin ph 1 / 46 , a mandible . its type locality is dzheroi 2 , which is in a bartonian marine sandstone in uzbekistan .
puppigerus longiceps : its type locality is london clay , isle of sheppey turtles , which is in a ypresian marine claystone in the london clay formation of the united kingdom .
puppigerus camperi : i . r . s . n . b . 1687 / r . 4 . its type locality is bruxelles sands ( sables d ' bruxelles ) - bruxelles formation , which is in a lutetian coastal sandstone in the bruxelles formation of belgium .
haiyan tong , ren hirayama , jérôme tabouelle ; puppigerus camperi ( testudines : cryptodira : cheloniidae ) from the ypresian ( early eocene ) of ouled abdoun basin , morocco . bulletin de la société géologique de france ; 183 ( 6 ) : 635–640 . doi : urltoken
a well preserved skull of puppigerus camperi ( testudines : cryptodira : cheloniidae ) is reported from the ypresian ( early eocene ) phosphate deposits of the ouled abdoun basin , morocco . this discovery adds an additional element to the rich turtle assemblages hitherto known in the ouled abdoun basin , suggesting further close relationships with the paleogene sea turtle fauna from northwestern europe .
ty - jour ti - the taxonomy and morphology of puppigerus - camperi an eocene sea turtle from northern europe t2 - bulletin of the british museum ( natural history ) . vl - 25 ur - urltoken pb - bm ( nh ) cy - london : py - 1974 sp - 155 ep - 186 sn - 0007 - 1471 au - moody , r t j er -
@ article { bhlpart83415 , title = { the taxonomy and morphology of puppigerus - camperi an eocene sea turtle from northern europe } , journal = { bulletin of the british museum ( natural history ) . } , volume = { 25 } , copyright = { copyright permission granted } , url = urltoken publisher = { london : bm ( nh ) } , author = { moody , r t j } , year = { 1974 } , pages = { 155 - - 186 } , }
typical reptiles of the eocene london clay of sheppey . ( a )  ( d ) the turtle puppigerus camperi ( gray , 1831 ) , skull in ( a ) dorsal and ( b ) ventral views , ( c ) carapace in dorsal view , ( d ) plastron in ventral view ; ( e ) chrysemys bicarinata ( bell , 1849 ) , partial carapace in dorsal view ; ( f ) platemys bullocki owen , 1841 , plastron in ventral view ; ( g ) and ( h ) the crocodile crocodilus spenceri buckland , 1837 , skull in ( g ) dorsal and ( h ) lateral views ; ( i ) the snake palaeophis toliapicus owen , 1841 , thirty dorsal vertebrae in side view . ( a )  ( f ) after owen and bell ( 1849 ) ; ( g ) and ( h ) after owen ( 1850b ) ; ( i ) after owen ( 1850c ) .
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full reference : e . d . cope . 1870 . synopsis of the extinct batrachia , reptilia and aves of north america . transactions of the american philosophical society , new series 14 ( 1 ) : 1 - 252
see also averianov 2005 , cope 1870 , lydekker 1889 and tong et al . 2012
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this is an abstract which has been accepted for the 5th turtle evolution symposium .
sandra chapman is employed at the natural history museum in london . christopher king was a consultant to oil industry .
richard t . j . moody analyzed the data , wrote the paper , prepared figures and / or tables , reviewed drafts of the paper , data comparison .
sandra chapman analyzed the data , wrote the paper , prepared figures and / or tables , reviewed drafts of the paper , data comparison .
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bulletin de la société géologique de france ( 2012 ) 183 ( 6 ) : 635 - 640 .
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british museum , 1974 . paperback . condition : very good . 1974 . british museum . softback . book - vg . 9 . 5x7 . 5 . pages 153 - 186 . 8 b / w photos , 15 b / w line illus . ex - college library .
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full reference : r . owen . 1841 . description of the remains of six species of marine turtles ( chelones ) from the london clay of sheppy and harwich . proceedings of the geological society of london 3 part 2 ( 83 ) : 565 - 578
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the cretaceous system in britain ( figures 8 . 1 , 8 . 2 ) is represented by two broad phases of deposition which relate to palaeogeography . earth movements during the late jurassic uplifted most of north - west europe to form land . in the british region , there were initially two main basins of deposition , in the east anglia - north sea area , and in the wessex - weald region and northern france .
these keywords were added by machine and not by the authors . this process is experimental and the keywords may be updated as the learning algorithm improves .
benton m . j . , spencer p . s . ( 1995 ) british cretaceous fossil reptile sites . in : fossil reptiles of great britain . springer , dordrecht
narrow belt e of the matawan formation from raritan bay - patuxent river . very fossiliferous
cardioceras is generally older than upper cretaceous . it was entered as dioceras which could be a typo . dioceras might be a misspelling of ' didymoceras ' which has been found in nj . arca originally cibota
can ' t identify exeter village , nj . most likely a reference to exeter twp , berks cnty , pa
in the cliffs along raritan bay between highland & cliffwood there are outcrops of cretaceous marine formations . reported to be closed off with seawalls to prevent erosion
there does not seem to be a town of highland . possibly atlantic highlands or highland park
original says ' martisburg ' . in thin dark shale beds below second watchung basalt layer
unable to identify a grist mills in nj . there are a large number of old grist mills in nj
marl pits mined for potash rich greensand @ birmingham , medford , marlton , ewansville in 1920s . fossils are reported from the marl @ birmingham latlong is the burlington pit
at alloway pit ( overgrown irrigation pond excavation ) e of newbold corner rd . close to nike missile towers
the original taxa list includes zygaena which is a moth . posssibly refers to the hammerhead shark sphyrna zygaena . website
in marl pits on shepard farm in shiloh . couldn ' t identify shepard farm on internet . lat lon is ~ 1200m n in excavated area that might be old marl pit w of junction of 49 & 635
in roadcuts along i280 into west orange @ exposures of the passaic formation under the orange mountain basalt . a few reptile _ tracks found during construction
rosedale . exit 4a i280w s on eisenhower pkwy . 1 . 7km . left on beaufort ave . 600m to park entrance . collecting permits sometimes available from cnty park service
jackson road adjacent to jackson road @ mantua - harrison twp lines in gloucester cnty . in
39 . 7489 , - 75 . 3796 , i295 - oldmans creek bridge . nj , ga
locality - - oldman creek is roughly 10km long . latlon is i295 bridge over oldmans creek
locality - - oldman creek is roughly 10km long . latlon is i295 bridge over oldmans creek4
locality 1 - - oldman creek is roughly 10km long . latlon is i295 bridge over oldmans creek
locality 2 - - oldman creek is roughly 10km long . latlon is i295 bridge over oldmans creek
39 . 7489 , - 75 . 3796 , i295 - oldmans creek bridge . nj , ga4
locality 5 - - oldman creek is roughly 10km long . latlon is i295 bridge over oldmans creek
junction of main / broadway . ceres / mantua park next to pitman golf course - chestnut branch
at remains of granton quarry ( a sometime road metal quarry ) between the lowes home building center & tonelle plaza . shale & remains of 11 von houten flooding cycles exposed on hillside ( ' 04 )
there are several outcrops in the area . one is @ the clinton , block & supply co quarry
google books a dictionary of the fossils of pennsylvania & neighboring states named in . . . by j . peter lesley
original says only ' clark quarry ' . assume this is clark quarry which did produce material about 800m e of the smith - clark quarry @ milford . latlon is downtown milford
on the campus of princeton u . , fish & crustacea have been found in basement excavations
in 1990 or so , well preserved cretaceous insect fossils were reported from amber in a then unspecified spot . reported to be @ the end of white oak road @ a depth of about a meter
in pit of crossman clay products w of garden state pkwy & perth amboy . redeveloped around 2000 for housing & commercial development . some exposures may remain after redevelopment , but the more interesting material in the south part of the quarry is now inaccessible . well preserved leaves - two dozen taxa ; insecta ; amber ; charcoal
per the january 97 scientific american , the amber site in nj was found by jerry case & is in the dry bed of a large river about 90 minutes ( i . e . 100 - 150 km ? ) from new york city . lignite is found in the bed of the river & also in the banks
south river , n . j . ( = washington , n . j . / south river n . j . , ' ash - colored clay near washington , middlesex cnty , n . j . ' )
in stream beds in monmouth cnty , nj in monmouth formation greensands . poricy brook , hop brook ( colts neck ) , holmdel creek
crosswicks creek @ mouth of shingles run . upstream of mt holly - freehold rd @ fishing access .
brachiopods - oleneothyris ; bivalves - ostrea , pycnodonte ; others brachiopods - oleneothyris , ostrea - pycnodonte . small inverts above cretaceous glauconite
on sandbars in crosswicks creek . downstream from mt holly - freehold rd @ fishing access
long overgrown walk from parking . get permission from the marina . about 400m to cliffs
sohl & koch ofr # 480 ( beers hill ; usgs # 31176 : 28868 ) . site seems to be something called the ' beers hill cut ' - - perhaps a rr cut ? . beers hill is apparentlya 343 ft hill 12 . 3 miles from deal , nj & 3 . 3km nnw of the old bell telephone labs facility ( in holmdel , nj ? ) used for communication testing . there is a group of hills in thatarea , but none seems to be 343 ft tall . latlon used is telegraph hill
on whale creek 2 . 5 km from cliffwood point in pit of cliffwood brick co in nodules
on big brook in area defined by 79 , 520 & 4 ( vanderburg road ) boundary rd ( nj4a ) transects area x - ing big brook n of vanderburg rd ( note - a collector was buried by a landslide in the cliff here . fortunately , he was with friends )
on ramanessin brook upstream from e . main / newman springs rd & middletown rd
fossils along the river between w . farms rd & preventorium rd . older outcrops to west
west of gsp on s side of river in a bank exposure reached via a sandy trail & access path beyond the first ravine . shark teeth , some large
west of nj turnpike . site used is arbitrary - - x - ing of gully rd over shark river
18n from gsp exit 105 . exit on wayside rd & proceed onto tinton ave / 537 . cross sycamore ave / water street & park on rt . near colt ' s neck line . invertebrates & crabs ( claws ) in indurated rock
on tributary of shark river just w of garden state pkwy turnpike interchange 100 . park @ shark river cnty park on schoolhouse rd
sohl & koch ofr # 472 ( = water witch station ; usgs # 16277 ) - - presumably this is an extinct train station in highlands , nj . latlon is @ jct of nj36 & waterwitch ave
this locality is cited in a number of old books & papers , but the name wordills is not recognized by mapping software . neither does it seem to be a simple mispelling .
many genera of fish as well as other vertebrates in area quarries . much or all of this locality was flooded by a reservoir
catopterusand / dictyopyge , both genera in redfieldiidae , reported , but nomenclature is ? . ransom , ' fossils in america '
2 km s of green pond mt . in shaley limestone along road s of newfoundland & in fields
original says ' arlington beach ' . there is a beach arlington bay beach park between 13th & 16th streets in ship bottom
at tilcon quarry - - plans in ' 04 were to terminate quarrying & develop as residential . google maps satellite images as of 2009 show only parts of the quarry to be developed access would be from valley road n of the us46 junction via four seasons road on the w . reptile _ tracks were fairly abundant here
at traprock quarry ( known variously as little ferry , ubc , r . h . hamilton , dell materials or tilcon quarry ) in two beds @ top of underlying mudstones
probably stegomus arcuatus jerseyensis 1 . 25 miles ( 2km ) southwest of neshanic station , somerset cnty ; j . higgins 1936
on left side of willow rd on path behind farm gate & up a dirt road in abandoned quarry . occasional trilobites in limestone
at a water treatment plant - - permission required . equally on the pa side
might be on pa side - within the national recreation area - no digging but surface collecting permitted . fish to 15cm
11km n of montague , w of clove rd . along ' trilobite ridge / trilobite mountain ' - an e - w trending structure south of the neversink river with devonian formations exposed
s to ne of long pond ( foerste ) . long pond ( greenwood lake ) is 20km ne of sparta
zaphrentis , monotrypa , pholidops , stropheodonta , orthothetes , dalmanella , chonetes , atrypa , reticularia , bivalves - loxonema , orthoceras , favosites , cyrtina , . . .
2km ' up ' from yards creek pumping station on old mine rd off walnut valley rd . - road cut
there does not seem to be a watertown in nj . oncoceras entered as onococeras
copyright 2006 - 2012 donald kenney ( donald . kenney @ gmail . com ) . unless otherwise stated , permission is hereby granted to use any materials on these pages under the
there may be errors in the list . some of the original material was incorrect or ambiguous .
many of the sites are closed to collecting or buried under shopping centers , highways , or housing developments . closed sites will stay on the list , as they are sometimes reopened , and sometimes the same rocks and fossils will turn up 100 yards or 100 miles down the road .
in general , permission is required from somebody to collect fossil vertebrate material in the us . an exception is often made in practice for shark teeth and occasionally for disarticulated marine vertebrate bones . always ask . rules for collecting invertebrate and plant material vary widely especially in ' wilderness areas ' . permission is always required to collect on private property . digging holes where you do not have permission to dig or are unfamiliar with the rules is almost always a bad idea .
some of the sites listed probably never existed in the first place , and other sites have been developed over and / or reclaimed out of existence . many of the localities when found will be closed to fossil collecting , and of those that are not , many will prove to be unproductive . a few of the sites are accessible to large groups without special arrangements .
in a narrow belt e of the matawan formation from raritan bay to patuxent river . very fossiliferous shellbeds
cardioceras is generally older than upper cretacsous . it was entered as dioceras which could be a typo . but dioceras might be a misspelling of ' didymoceras ' which has been reported from the new jersey cretaceous . arca originally cibota , aninaea originally axinea
new jersey - in the cliffs along raritan bay between highland and cliffwood there are outcrops of cretaceous marine formations . reported to be closed off with seawalls to prevent erosion
there does not seem to be a town of highland . possibly atlantic highlands , highlands , or highland park
barney ' s dock is a dock and commercial clam processing facility at 700 n rhode island ave . the beds are reported to be grey sand with abundant clam fragments . there is a small park area just south of the dock facility that might expose some beds
on ridge on w side of meadowlands 40d48 . 434mn , 074d06 . 506mw , a few reptile _ tracks from this and other area exposures ( 1960 - 2002 ) access from madison st and meadow rd on e side of new zion and hillside cemeteries or from parking lots of complex on nw corner of valley brook ave and polito ave
marl pits mined for potash rich greensand at birmingham , medford , marlton , ewansville in 1920s . fossils are reported from the marl at birmingham
flabellina is a shell - less nudibranch . how could it be recognized as a fossil ? . porosoma originally coptosoma . lingulina - - unclear if foram , or inarticulate brachiopod
probably should ref s branch of rancocas creek . n branch would be ewansville , etc .
the original taxa list includes zygaena however the genuszygaena is a moth . posssibly a reference to the hammerhead shark sphyrna zygaena
in marl pits on shepard farm in shiloh . couldn ' t identify shepard farm on internet . lat lon is about 1200m n in excavated area that might be old marl pit w of junction of nj49 and cr635
in roadcuts along i280 into west orange at exposures of the passaic formation under the orange mountain basalt . a few reptile _ tracks found during construction
in rosedale , nj . exit 4a i280w s on eisenhower parkway . 1 . 7km . left on beaufort ave . 600m to park entrance . collecting permits sometimes available from county park service
original says ' barnesboro ' which seems to be an old spelling . topo maps show an area about 500m east of barnsboro that might once have been a quarry or pit , but satellite photos show it to be built over with housing .
jackson road adjacent to jackson road at mantua - harrison township lines in gloucester county . in
39 . 7489 , - 75 . 3796 , location : i295 - oldmans creek bridge . nj , ajk73
locality - - oldman creek is roughly 10km long . latlon is the i295 bridge over oldmans creek
locality - - oldman creek is roughly 10km long . latlon is the i295 bridge over oldmans creek4
locality 1 - - oldman creek is roughly 10km long . latlon is the i295 bridge over oldmans creek
locality 2 - - oldman creek is roughly 10km long . latlon is the i295 bridge over oldmans creek
locality 5 - - oldman creek is roughly 10km long . latlon is the i295 bridge over oldmans creek
at junction of main and broadway . ceres / mantua park next of pitman golf course in banks of chestnut branch
probably sw of town . sat images show area that was probably a pit or quarry has been redeveloped as housing . same as barnsboro ?
at remains of granton quarry ( a sometime road metal quarry ) between the lowes home building center and tonelle plaza . shale and remains of 11 von houten flooding cycles exposed on hillside ( 2004 ) 40d48 . 431mn , 074d01 . 071mw
there are several outcrops in the area . one is at the clinton , block and supply co quarry
original says only ' clark quarry ' . assume this is the clark quarry which did produce material like this - - said to be about 800m e of the smith - clark quarry at milford . latlon is downtown milford
google books a dictionary of the fossils of pennsylvania and neighboring states named in . . . by j . peter lesley
per the january 97 scientific american , the amber site in new jersey was found by jerry case and is in the dry bed of a large river about 90 minutes ( i . e . 100 - 150 km ? ) from new york city . lignite is found in the bed of the river and also in the banks
in pit of crossman clay products w of garden state parkway and perth amboy . redeveloped around 2000 for housing and commercial development . some exposures may remain after redevelopment , but the more interesting material in the south part of the quarry is now inaccessible . well preserved leaves - two dozen taxa ; insecta ; amber ; charcoal
in 1990 or so , well preserved cretaceous insect fossils were reported from amber in a then unspecified location in new jersey . reported to be at the end of white oak road at a depth of a meter or more below the ground surface
south river , n . j . ( = washington , n . j . / south river n . j . , ' ash - colored clay near washington , middlesex county , n . j . ' )
in stream beds in monmouth county , new jersey in monmouth formation greensands . poricy brook , hop brook ( colts neck ) , holmdel creek
40 . 3203 , - 74 . 2142 , location : big brook . nj , ajj14 ?
on sandbars in crosswicks creek . downstream from mt holly - freehold rd at fishing access
on crosswicks creek at mouth of shingles run . upstream from mt holly - freehold rd at fishing access . brachiopods ( abundant ) - oleneothyris , ostrea - pycnodonte , and few other taxa . small invertebrates of various sorts in sands above the cretaceous glauconite
a fairly long walk from nearest parking and largely overgrown . accessible with permission from the marina . about 400m to cliffs
sohl and koch ofr # 480 ( = beers hill ; usgs # 31176 : 28868 ) . this locality seems to be something called the ' beers hill cut ' - - perhaps a rr cut ? . beers hill is apparently a 343 ft hill 12 . 3 miles from deal , nj and 3 . 3km nnw of the old bell telephone labs facility ( in holmdel , nj ? ) used for communication testing . there is a group of hills in that general area , but none seems to be 343 ft tall . latlon used is telegraph hill
verbal description ( which is kind of hazy ) would be several km w of the lat lon . either ( or neither ) could be correct .
on big brook in area defined by us79 , nj520 and nj4 ( vanderburg road ) boundary rd ( nj4a ) transects area crossing big brook n of vanderburg rd ( note - a collector was buried by a landslide in the cliff here . fortunately , he was with friends )
on ramanessin brook upstream from intersection of e . main / newman springs rd and middletown rd
the old marl pits ' near hornerstown ' do not seem to appear on topo or satellite maps .
howell is a township , not a town per se . there doesn ' t seem to be a town of howell to have a downtown .
possibly near squankum . no sign of the manesquan river pits ( there were several ) on modern maps .
miscellaneous fossils along the river between west farms rd and preventorium rd . older outcrops to the west
south river runs ns and seems to end near old bridge . loc is in spotswood near apparent end of the river .
there is no middletown mall and the state highway through middletown is nj35 . nj33 runs ew from trenton to bradley beach . could be a refernce to the monmouth mall on nj35 and nj36 at 40 . 287 , - 74 . 0571
west of garden state parkway on s side of river in a bank exposure reached via a sandy trail and access path beyond the first ravine . shark teeth , some large
latlon is the rr station in belmar . any marl pits in the area have presumably been long since built over or made into parks . ( could also be the rr station a km or two north in avon - by - the - sea . ) there is also a shark river station road near tinton falls that crosses the shark river at 40 . 2281 , - 74 . 1237
protocardia originally protocardium . aninaea originally axinea . location used is arbitary . actual site ( s ) could be further n and / or w .
nj18n from garden state parkway exit 105 . exit on wayside rd and proceed straight through lights onto tinton ave / nj537 . cross sycamore ave / water street and park on rt . near colt ' s neck line . invertebrates and crabs ( claws ) in indurated rock
on tributary of shark river just w of garden state parkway turnpike interchange 100 . park at shark river county park on schoolhouse rd
sohl and koch ofr # 472 ( = water witch station ; usgs # 16277 ) - - presumably this is an extinct train station in highlands , nj . latlon is at jct of nj36 and waterwitch ave
this locality is cited in a number of old books and papers , but the name wordills is not recognized by mapping software . neither does it seem to be a simple mispelling of some other named place
catopterusand dictyopyge both reported . both are genera within the redfieldiidae , but the nomenclature is confused
original says ' arlington beach ' . no such place , but there is a beach arlington bay beach park between 13th and 16th streets in ship bottom
at tilcon quarry - - plans in 2004 were to terminate quarrying and develop as residential property . google maps satellite images as of 2009 show only parts of the quarry to be developed 40d52 . 470mn , 074d11 . 382mw . access would be from valley road n of the us46 junction via four seasons road on the w . reptile _ tracks were fairly abundant here
at traprock quarry ( known variously as little ferry , ubc , r . h . hamilton , dell materials or tilcon quarry ) in two beds at top of underlying mudstones
probably stegomus arcuatus jerseyensis 1 . 25 miles ( 2km ) southwest of neshanic station , somerset county ; j . higgins 1936
on left side of willow rd on path behind farm gate and up a dirt road in abandoned quarry . trilobita ( occasional ) in limestone
at a water treatment plant - - permission required . might equally well be on the pa side
might equally well be on the pa side - - within the national recreation area - - no digging but surface collecting permitted . fish to 15cm
hoptacong is a fair sized town in the se part of sussex cty . ( 40 . 9337 , - 74 . 6617 ) whereas old mine rd runs along the ne boundary of the county ?
11km n of montague , w of clove rd . along ' trilobite ridge / trilobite mountain ' - - an e - w trending structure s of the neversink river with several devonian formations exposed
not really clear which direction ' below ' is . peters valley is at 41 . 1964 , - 74 . 8507
41 . 3322 , - 74 . 7135 , location : nearpass quarry . nj , akm73
2km ' up ' from yards creek pumping station on old mine rd off walnut valley rd . in road cut
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lower end of the thigh bone of megalosaurus , from cornwell , oxfordshire : one of the first fossil reptile bones to be illustrated from britain , and the oldest recorded figure of a dinosaur ( from plot , 1677 ) .
the skull patterns , in side view , of the major lineages of reptiles . the anapsid pattern ( a ) is plesiomorphic ( primitive ) , being present also in fishes and amphibians , while the diapsid ( b ) and synapsid ( c ) patterns define two major clades of amniotes , the diapsida ( thecodontians , dinosaurs , pterosaurs , crocodiles , birds ) and the synapsida ( mammal - like reptiles and mammals ) . the euryapsid pattern ( d ) may have arisen more than once , in different marine groups , and appears to be a derivative of the diapsid pattern ( after benton , 1990a ) .
cladogram of the major groups of reptiles , based on recent analyses ( after benton , 1990a ) .
evolutionary tree of the main groups of reptiles , with proposed relationships based on recent cladistic analyses , and the stratigraphical distributions based on global data . the width of the ‘spindles’ represents diversity of the groups ( after benton , 1990a ) .
table showing the classification of the major groups of reptiles , based on the cladograms summarized in figure 1 . 3 . symbols : â† extinct group ; * paraphyletic group ( after benton , 1990a ) .
( a ) map of great britain showing the distribution of the 50 gcr fossil reptile sites ; ( b ) the outcrop pattern of permian , triassic , jurassic , cretaceous and tertiary rocks in great britain . after benton ( 1988 ) .
generalized stratigraphic column showing the major british fossil reptile sites in sequence , and comparable sites elsewhere in the world .
map showing the distribution of permian rocks in great britain . gcr permian reptile sites : ( 1 ) middridge ; ( 2 ) cutties hillock ; ( 3 ) masonshaugh .
reptile specimens from the late permian marl slate of middridge quarry , county durham : ( a ) protorosaurus speneri meyer , 1830 , part of the backbone ( after hancock and howse , 1870b ) ; ( b ) adelosaurus huxleyi ( hancock and howse , 1870 ) , partial skeleton ( after evans , 1988b ) .
part of the worked face at the east end of the main quarry , cutties hillock , showing cross - bedding . the fossil reptile remains were recovered from the foot of the cross - bedded units . ( photo : m . j . benton . )
reptile specimens from the late permian cutties hillocks sandstone formation of cutties hillock quarry , morayshire . skulls of ( a ) gordonia ; ( b ) geikia ; and ( c ) elginia , all drawn to the same scale . after benton and walker ( 1985 ) .
reptile footprints from the late permian hopeman sandstone formation of masonshaugh quarry , morayshire : ( a ) small prints ; ( b ) medium prints , chelichnus megacheirus huxley , 1877 ; and ( c ) large prints . after benton and walker ( 1985 ) .
map showing the distribution of triassic rocks in great britain . gcr triassic reptile sites : ( 1 ) grinshill quarries ; ( 2 ) coten end quarry ; ( 3 ) guy’s cliffe ; ( 4 ) sidmouth coast section ; ( 5 ) otterton point ; ( 6 ) lossiemouth east quarry ; ( 7 ) spynie ; ( 8 ) findrassie ; ( 9 ) bendrick rock ; ( 10 ) aust cliff ; ( 11 ) slickstones ( cromhall ) quarry ; ( 12 ) durdham down ; ( 13 ) emborough quarry ; ( 14 ) tytherington quarry .
the stratigraphy of the british triassic reptile faunas . correlations of the standard triassic divisions and the german triassic sequence with the british triassic , as proposed by hull ( 1869 ) for the ‘classical’ british succession , and by warrington et al . ( 1980 ) for currently recognized lithostratigraphical units . skull symbols indicate the levels of the main tetrapod faunas , and asterisks denote palynological evidence of relative age . age dates ( ma â± 5 ) after forster and warrington ( 1985 ) . from benton et al . ( 1993 ) .
the grinshill localities . the map is based on published maps of the british geological survey ( bgs 1 : 63 , 360 scale geological sheet 138 , wem ) and on field observations by m . j . b . i . bridge quarries ; ii . working quarry ( ecc quarries ltd ) .
the operational quarry on grinshill : view of the north face , showing the massive cross - bedded helsby sandstone formation at the bottom , and the softer , more thin - bedded tarporley siltstone formation above . ( photo : m . j . benton . )
rhynchosaurus articeps , the only member of the grinshill skeletal fauna : typical fossil remains ( ac ) and restorations ( dh ) . ( a ) partial skeleton lacking the tail and the limbs of the left side , in ventral view ( bmnh r1237 , r1238 ) ; ( b ) dorsal vertebrae , ribs , and right forelimb in posteroventral view ( shrbm 6 ) ; ( c ) pelvic region , right leg with ankle bones , presacral vertebrae 2225 , sacral vertebrae 1 and 2 , and caudal vertebrae 18 ( batgm m20a / b ) ; ( d )  ( g ) restoration of the skull , based on shrbm g132 / 1982 and 3 and bmnh r1236 , in lateral ( d ) , dorsal ( e ) , ventral ( f ) , and occipital ( g ) views ; ( h ) restoration of the skeleton in lateral view in walking pose . all based on benton ( 1990a ) .
typical elements of the warwick fauna . ( a ) left postero - external angle of the skull of ‘ cyclotosaurus pachygnathus ’ ( cyclotosaurinae incertae sedis ) in lateral view ( warms gz13 ) ; ( b ) part of the snout of ‘ stenotosaurus leptognathus ’ ( stenotosaurinae incertae sedis ) in palatal view ( warms gz38 ) ; ( c ) posterior portion of a left lower jaw of ‘ stenotosaurus leptognathus ’ in lateral view ( warms gz35 ) ; ( d ) scattered bones of cf . macrocnemus ( rhombopholis scutulata ) ( warms gz10 ) ; ( e )  ( h ) , assorted remains of rhynchosaurus brodiei : ( e ) anterior part of the skull in lateral view ( warms gz6097 / bmnh r8495 ) , ( f ) anterior part of a dentary in medial view ( warms gz950 ) , ( g ) mid - dorsal vertebra in right lateral view ( warms gz17 ) , and ( h ) interclavicle in ventral view ( warms gz34 ) ; ( i ) right ilium of bromsgroveia walkeri in lateral view ( warms gz3 ) . after various sources ; from benton et al . ( 1993 ) .
map of the coastal outcrop of the otter sandstone formation between sidmouth and budleigh salterton , devon . the major triassic formations are indicated , together with mean fluvial palaeoflow directions , and principal tetrapod localities . from benton et al . ( 1993 ) .
the otter sandstone formation exposed in the cliffs west of sidmouth , view looking east . fish and reptile remains have been found at various horizons . ( photo : m . j . benton . )
smaller elements of the otter sandstone formation fauna of devon . right dentaries ( a , c ) and a left maxilla ( b ) of a procolophonid , all in lateral view ( exems 60 / 1985 . 311 , 3 , and 154 ) ; ( d ) dentary fragment of an unknown small pleurodont reptile , showing pits for teeth , in lingual view ; ( e ) tooth of ? tanystropheus , showing small accessory cusps ( exems 60 / 1985 . 143 ) ; ( f ) , ( g ) recurved teeth of two kinds of unknown archosaurs ( bristol univ . unnumb . ) ; ( h ) unidentified insect wing ( bristol univ . unnumb . ) ; ( i ) , ( j ) carapaces of the conchostracan euestheria ( bristol univ . unnumb . ) . after various sources ; from benton et al . ( 1993 ) .
imaginary scene during mid triassic times in devon , based on specimens recovered from the otter sandstone formation between sidmouth and budleigh salterton . a scorpion ( mid - foreground ) contemplates a pair of procolophonids on the rocks . opposite them , a hefty temnospondyl amphibian has spotted some palaeonisciform fishes , dipteronotus , in the water . two rhynchosaurus stand in the middle distance and , behind them , a pair of rauisuchians lurk . the plants include equisetites ( horsetails ) around the waterside and voltzia , a coniferous tree . drawn by pam baldaro , based on her colour painting .