the black carp ( mylopharyngodon piceus ) or black chinese roach is a species of cyprinid fish and the sole species of the genus mylopharyngodon .

it is native to lakes and rivers in east asia , ranging from the amur basin , through china , to vietnam .

it is widely cultivated for food and chinese medicine .

the black carp can reach up to 1.8 m ( 5.9 ft ) in length and 35 kg ( 77 lb ) in weight .

it generally feeds on snails and mussels .

the average length is 60 – 120 cm ( 23.5 – 47 in ) .

black carp , together with bighead , silver , and grass carps , make up the culturally important " four famous domestic fishes " used in polyculture in china for over a thousand years , and known as " asian carp " in the united states .

black carp are not as widely distributed worldwide as the other three .

in china , black carp are the most highly esteemed and expensive foodfish among the four domestic fishes , and partly because of its diet and limited food supply , is the most scarce and expensive in the marketplace .

black carp

more information on black carp and black carp distribution in the united states : u . s . geological survey animated map * u . s . geological survey black carp fact sheet identification of black carp and grass carp
urltoken - black carp university of georgia . center for invasive species and ecosystem health .
fws fact sheet on black carp draft environmental assesment draft economic analysis asian carp prevention and control act ( hr . 3049 and s . 1402 )
black carp ( jul 2002 ; pdf | 449 kb ) doi . fws . invasive species program .
black carp feed on mollusks and snails , consuming up to 20 % of their body weight per day .
black carp found at river mile 137 of illinois river in april of 2017 . photo courtesy of aaron roberts
while adult black carp have been found sporadically in the mississippi , the november discovery near cape girardeau of juvenile fish among the hundreds of fish caught showed the black carp population in the river is higher than scientists expected , missouri department of conservation resource scientist quinton phelps said , and that there’s a “high probability” that more black carp were caught .
what to do if you think you have found an asian carp ( 2010 ; pdf | 584 kb ) asian carp regional coordinating committee . see asian carp newsroom for updated news regarding asian carp response in the midwest .
resembles many of the carp species in the united states . including its asian cousins : grass carp ( ctenopharyngodon idella ) silver carp ( hypophthalmichthys molitrix ) largescale silver carp ( hypophthalmichthys harmandi ) bighead carp ( hypophthalmichthys nobilis ) common goldfish ( carassius auratus ) crucian carp ( carassius carassius ) mud carp ( cirrhinus molitorella ) also resembles the common carp ( cyprinus carpio ) ( common carp are european , not asian , and are sometimes considered
native
because they have been in the us since the 1800s )
a black carp captured this april in the illinois river by a commercial fisher highlights a unique partnership between fishers , the illinois department of natural resources , and southern illinois university . for surrendering this black carp , the commercial fisher received a $ 100 bounty , and in turn , helped resource agencies learn a little more about the range of black carp in the illinois river . the black carp was found south of peoria , illinois near copperas creek lock and extends the upstream detection of the species by 110 miles .
nonindigenous aquatic species database . fact sheet - black carp . usgs , gainesville , fl . [ accessed sep 16 , 2014 ] .
u . s . present : the black carp has been reported in arkansas , illinois , mississippi , and missouri . texas : while the black carp has been used in aquaculture in states as close as louisiana for decades , currently there are no reported texas invasions .
illustrations and detailed information useful for the positive identification of this species appear in nico et al . ( 2005 ) and schofield et al . ( 2005 ) . an identification key to introduced asian carps and other cyprinids , including black carp , is provided by schofield et al . ( 2005 ) . the black carp closely resembles the grass carp ctenopharyngodon idella . the two species are similar in overall body shape , size and placement of fins . both black carp and grass carp have very large scales . in contrast to grass carp , the black carp is slightly darker in coloration ( not black ) and its pharyngeal teeth ( throat teeth ) are large and similar in appearance to human molars , an adaptation for crushing the shells of mollusks ( nico et al . 2005 ) commercial fishers in louisiana have noted that black carp also have a somewhat pointed snout , a character they find useful in distinguishing it from grass carp . juveniles and larvae may be difficult to distinguish from those of grass carp and certain other cyprinids . illustrations and descriptions of juvenile and larval asian carps , including black carp , appear in nico et al . ( 2005 ) and chapman ( 2006 ) .
the black carp is one of four species of asian carp that threaten waterways in the central united states . as molluscivores , black carps consume native freshwater mussels and snails that live in our large rivers . 26 freshwater mussel species native to illinois are state - threatened or endangered , twelve of which are federally listed . the asian carp regional coordinating committee’s collaborative 2017 asian carp action plan recognizes the informational needs for black carp control and management , and is further addressing this need in the annual monitoring and response plan which focuses on the upper illinois waterway .
commercial fishers are valuable in the cooperative effort to better understand black carp because they are skilled in fishing large rivers throughout the region . beyond the illinois river , the bounty program also accounts for the majority of adult black carp reported in the rest of the country . in addition to commercial fishers , recreational anglers and bowfishers should be aware of these invasive fish and what to do if one is harvested . in appearance , black carp closely resemble the more common grass carp , another asian carp which is also found in large rivers of the central united states . proper identification of black carps is an essential component of the bounty program .
there is high potential that the black carp would negatively impact native aquatic communities by feeding on , and reducing , populations of native mussels and snails , many of which are considered endangered or threatened ( nico et al . 2005 ) . given their size and diet preferences , black carp have the potential to restructure benthic communities by direct predation and removal of algae - grazing snails . mussel beds consisting of smaller individuals and juvenile recruits are probably most vulnerable to being consumed by black carp ( nico et al . 2005 ) . furthermore , based on the fact that black carp attain a large size ( well over 1 meter long ) , both juvenile and adult mussels and snails of many species would be vulnerable to predation by this fish ( nico et al . 2005 ) . fish farmers report that black carp are very effective in reducing the numbers of snails in some ponds . recently , wui and engle ( 2007 ) argued that black carp can eliminate 100 % of the snails in a single pond . although their assumption that black carp are capable of eliminating all common pond snails in ponds is open to debate , the effectiveness of black carp in significantly reducing snail populations in aquaculture ponds indicates that any black carp occurring in the wild may cause significant declines in certain native mollusk populations in north american streams and lakes ( nico et al . 2005 ) . because the life span of black carp is reportedly over 15 years , sterile triploid black carp in the wild would be expected to persist many years and therefore have the potential to cause harm native mollusks by way of predation ( nico et al . 2005 ) .
black carp would compete with mollusk - eating native fish , including freshwater drum , redhorse species and the state threatened lake sturgeon , for limited food resources .
there is high potential that the black carp will negatively impact native aquatic communities by feeding on , and reducing , populations of native mussels and snails , many of which are considered endangered or threatened . given their size and diet preferences , black carp also have the potential to restructure benthic communities by direct predation and removal of algae - grazing snails and native mussels . furthermore , because the black carp can attain a large size ( well over 1 meter long ) , juvenile and adult mussels and many species of snails would be vulnerable to predation . since the life span of the black carp is reportedly over 15 years , sterile triploid black carp in the wild are expected to persist many years and therefore have the potential to cause harm native mollusks by way of predation . also , black carp juveniles feed on zooplankton and insect larvae while adults feed on benthic invertebrates such as snails and mussels , so many different resources maybe become exhausted .
chick et al . ( 2003 ) believed that their illinois capture was the first wild record of black carp in the united states , but nico et al . ( 2005 ) provided new information indicating that louisiana commercial fishers had been collecting black carp in louisiana since the early 1990s . however , until recently the louisiana commercial fishers thought that the black carp in their nets were just an unusual type of grass carp—somewhat darker and with a higher dorsal fin and more pointed head or snout ( nico et al . 2005 : xiii ) . the black carp that escaped in missouri may have been triploid and thus considered sterile ( anonymous 1994b ) . however , it also was rumored that these fish may have been brood stock . all wild black carp examined to date taken in louisiana waters have been found to be diploid ( nico et al . 2005 ) .
nonindigenous aquatic species database : point map - black carp doi . usgs . wetland and aquatic research center . provides detailed collection information as well as animated map .
because it’s a federal offense to carry a live black carp across state lines , anyone who finds one should kill it and try to bring it to authorities without freezing it , said chapman , who is expecting to get funding this fall to develop a bait that targets black carp while not endangering other species or the public .
black carp , like the other asian carps , are able to invade novel habitats and ecosystems readily due to a variety of factors . first , the carp are aggressive and prodigious feeders . individuals may consume as much as 20 % of their body mass in one day . further , black carp are explosive breeders . gaining sexual maturity after a few years , these carp may lay hundreds of thousands of eggs in one brood . unfortunately , as the carp mature and grow they produce even more eggs and increase their fecundity , allowing for further invasions .
nonindigenous aquatic species database : fact sheet - black carp doi . usgs . wetland and aquatic research center . provides distribution maps and collection information ( state and county ) .
black carp was first imported into the united states in the 1970s , and by the 1990s this species was being used in fish farms in several southern states to control pond snails . black carp were reported as having escaped into the osage river from a missouri fish farm during a major flood in april 1994 . due to its widespread use to control snails , escapees from aquaculture ponds have probably added to the wild population . during recent years there have been reports of black carp being captured in the wild . the first published report was that of a single black carp taken by a commercial fisher from horseshoe lake in southern illinois in march 2003 . other reports of captures have surfaced since .
voucher preserved specimens of the two wild - caught black carp taken in illinois are deposited in the ichthyological collection of southern illinois university - carbondale . several wild - caught black carp taken in louisiana waters were preserved and are in the possession of biologists at louisiana state university and at the u . s . geological survey - gainesville center in florida .
devaney et al . ( 2009 ) performed ecological niche modeling to examine the invasion potential for black carp and three other invasive cyprinids ( grass carp ctenopharyngodon idella , common carp cyprinus carpio , and tench tinca tinca ) . the majority of the u . s . between the mississippi river basin and the atlantic coast had a moderate to high predicted ecological suitability for this species , with the mississippi river drainage ( where individuals of black carp have been caught in the wild ) having the highest overall predicted suitability .
in aquaculture ponds ( nico et al . 2005 ) . the first known record of an introduction of black carp into open waters occurred in missouri in 1994 when thirty or more black carp along with several thousand bighead carp reportedly escaped into the osage river , missouri river drainage , when high water flooded hatchery ponds at an aquaculture facility near lake of the ozarks . recently , owners of the missouri facility where the escapes reportedly took place have denied that black carp ever escaped from their facility ( nico et al . 2005 ) . in any case , flooding of aquaculture facilities and associated numbers and types of escaped fishes are very poorly documented in the public record . there is evidence that large portions of the lower mississippi river basin where aquaculture farms are present have been subject to large - scale floods on a number of occasions over the past few decades . consequently , it is likely that the source of some or all of the black carp present in the lower mississippi river basin . nearly all fish farms with black carp are in lowland areas and flood events increase the probability that more black carp will eventually escape fish farms ( nico et al . 2005 : 245 ) . there is also risk that black carp may be spread by other means . according to one aquaculture farmer , hundreds of young black carp were accidentally included in shipments of live baitfish sent from arkansas to bait dealers in missouri as early as 1994 ( nico et al . 2004 : 5 ) . in addition , because of the continued widespread distribution of grass carp across the united states , there remains the possibility that shipments may inadvertently contain black carp ( nico et al . 2005 ) . juveniles , in particular , are difficult to distinguish from grass carp young . as such , nico et al . ( 2005 ) expressed concern over the increased risk that the species be misidentified and unintentionally introduced as
grass carp
to some areas .
black carp have been used in the aquaculture industry for decades so its removal in the near future seems unlikely . however , scientists , worried about an escape / colonization event , have urged fish farmers to use triploid black carp . these triploid individuals are sterile , so even if they do escape it will not lead to a breeding population . in fact , it is not illegal ( thanks to the lacey act ) to transport viable black carp across states . the lacey act has largely prevented the unwanted spread of black carp across the us . there is evidence that wild populations of black carp may have been present in the lower mississippi river basin , largely in and around the red river of louisiana , since the early 1990s . reproduction in the mississippi river has not been documented , but new information and recent collections suggest this species has likely established in the lower part of the mississippi basin .
the black carp is a blackish brown fish with blackish grey fins , an elongated and laterally compressed body . they average more than 3 feet in length and 33 pounds in weight , but can reach 5 feet in length and weigh up to 150 pounds . individuals of the species are known to live for at least 15 years . young black carp are difficult to distinguish from young grass carp ( ctenopharyngodon idella ) , another non - native species .
mississippi national river and recreation area - asian carp overview doi . national park service .
officials are asking the public to help stop the spread of black carp by not dumping their bait buckets indiscriminately and stocking ponds with fish from licensed vendors , phelps said . and in some states along the mississippi — including missouri , ohio , illinois and tennessee — any black carp caught in the wild can earn a $ 100 reward from southern illinois university , using money from the illinois department of resources .
the native range of black carp includes most major pacific ocean drainages of eastern asia from the amur river basin south to the west - pearl river basin , and possibly the red river of northern vietnam .
priority species : asian carp washington state recreation and conservation office . washington invasive species council .
ben - ami , f . , & heller , j . 2001 . biological control of aquatic pest snails by the black carp mylopharyngodon piceus . biological control , 22 ( 2 ) , 131 - 138 .
the black carp is a bottom - dwelling molluscivore that has been used by u . s . fish farmers to prey on and control disease - carrying snails in their farm ponds ; more recently , this species has been proposed as a biological control for the introduced zebra mussel dreissena polymorpha . although the subject has been debated , to date , there is no experimental evidence that indicates black carp would be effective in controlling zebra mussels . because black carp do not have jaw teeth and their mouths are relatively small , it is unlikely that these fish are capable of breaking apart zebra mussel rafts ( nico et al . 2005 ) .
aquatic invasive species : black carp ( apr 2009 ; pdf | 216 kb ) indiana department of natural resources . see also : invasive species for exotic animal and plant pests invading indiana , causing economic and visual damage
if they become established in the great lakes , black carp could pose a major threat to michigan’s native mussel populations , many of which are endangered , threatened , of special concern , or in need of conservation .
u . s . habitat : a freshwater fish , the black carp has found suitable habitat in the united states . the great lakes and mississippi river ( and others ) , offer the necessary habitat for a rapid growth in the population numbers of these fish . there are plenty of snail and mussel species for the fish to eat , however many of those species are already endangered and could be pushed to extinction by the black carp .
chick , j . h . , r . j . maher , b . m . burr , m . r . thomas . 2003 . first black carp captured in u . s . science . 300 : 1876 - 1877 .
wui , y . - s . & engle , c . r . 2007 . the economic impact of restricting use of black carp for snail control on hybrid striped bass farms . north american journal of aquaculture 69 : 127 - 138 .
this 2008 photo provided by the u . s . geological survey shows jeremy haley holding a 50 - pound black carp at the usgs laboratory , in columbia , mo . the discovery of two juvenile black carp in a ditch attached to the mississippi river in missouri is a troubling sign that the invasive species is reproducing in the wild , which could threaten already - endangered mollusks and native fish species in the river , research scientists said . ( duane chapman / u . s . geological survey via ap )
nico , l . g . , j . d . williams , and h . l . jelks . 2005 . black carp : biological synopsis and risk assessment of an introduced fish , american fisheries society special publication 32 , bethesda , md . 337 p .
kansas city • the discovery of two juvenile black carp in a ditch connected to the mississippi river in missouri is the first , troubling sign that the invasive species is reproducing in the wild and becoming more of a threat to already endangered mollusks and some native fish , scientists say .
black carp were brought to the u . s . in the 1970s to help fish farms fight snails , which carry parasites that are dangerous to several fish species . no native fish is as efficient and , at the time , farmers and the government didn’t want to use chemicals .
factsheet : asian carp pennsylvania state university . pennsylvania sea grant . see also : aquatic invasive species : resources for additional species information
“scientists really thought there were not enough adult black carp in the wild to find each other and reproduce , ” phelps said . “but what we found through this sampling is evidence there are enough to reproduce , and those young are surviving to a point where we are collecting them . ”
scientists from federal agencies including the u . s . fish and wildlife service and u . s . geological survey provide valuable analysis of the surrendered black carps . the most recent black carp reported near peoria , illinois was 28 inches in length and weighed eight pounds . analysis by the u . s . fish and wildlife service indicates that this fish was fertile , referred to as diploid , which is consistent with most fish recently captured in the bounty program . since the establishment of the bounty program in 2015 , 37 black carps have been collected . of these , 26 were collected in 2016 alone . four have been found in the illinois river .
this species was first brought into the united states in the early 1970s as a
contaminant
in imported grass carp stocks . these fish came from asia and were sent to a private fish farm in arkansas ( nico et al . 2005 ) . subsequent introductions of black carp into this country occurred in the early 1980s . during this period it was imported as a food fish and as a biological control agent to combat the spread of yellow grub
to report a black carp captured in the central united states from the mississippi , illinois , ohio or wabash rivers , please call a number below during regular business hours or report to your local natural resource agency . fish should be held cold , on ice , not alive , until passed to a natural resource agent .
nico , l . g . , j . d . williams , and h . l . jelks . 2005 . black carp : biological synopsis and risk assessment of an introduced fish , american fisheries society special publication 32 , bethesda , md . t . a . crowl . 1990 . life - history strategies of freshwater snail in response to stream permanence and predation : balancing conflicting demands . oecologia 84 : 238–243 . w . l . shelton , a . soliman and s . rothbard . 1995 . experimental observation on feeding biology of black carp ( mylopharyngodon piceus ) . bamidgeh 47 : pp . 59–67 . internet sources urltoken urltoken urltoken urltoken urltoken
“everyone is up in arms about damage from species like feral hogs , whitetail deer , and other native terrestrial species , ” he said . “the impact doesn’t seem to be as bad when the damage is under the water . but ( black carp ) is no different or less destructive than other invasive species’ interaction with native wildlife . ”
* updated on may 23 , 2017 to reflect the reported capture of a black carp at douglas lake near chillicothe , illinois , adjacent to the illinois river at approximately river mile 182 . the report was from a commercial fisher , although no specimen was provided . more information can be found on u . s . geological survey ' s nonindigenous aquatic species page .
black carp can grow to 150 pounds and are the most efficient prey of mollusks in freshwater streams , according to duane chapman , a research fish biologist with the u . s . geological survey . that’s a problem because the mississippi river basin has the most diverse mollusk population in the world , and three - fourths of the mussel species are threatened or endangered , he said .
schramm , h . l . , jr . & basler , m . c . 2005 . evaluation of capture methods and distribution of black carp in arkansas , louisiana , and mississippi : final report 1 june 2004 - 31 may 2005 submitted to region 4 , u . s . fish and wildlife service , fisheries , atlanta , georgia . mississippi state , mississippi : u . s . geological survey , mississippi cooperative fish and wildlife research unit .
aquatic invasive species : asian carp risk analysis for arizona ( oct 2011 ; pdf | 316 kb ) arizona game and fish department . see also : aquatic invasive species for additional risk analyses and related species information
freshwater aquatic invasive species in rhode island - asian carp ( sep 2010 ; pdf | 553 kb ) rhode island department of environmental management . office of water resources . see also : aquatic invasive animals for species of concern
devaney , s . c . , k . m . mcnyset , j . b . williams , a . t . peterson , and e . o . wiley . 2009 . a tale of four
carp
: invasion potential and ecological niche modeling . plos one 4 ( 5 ) : e5451 .
this species can be found in rivers , streams , or lakes ; however , it requires large rivers to reproduce ( nico et al . 2005 ) . reproduction takes place in late spring and summer when water temperatures and / or water levels rise ( nico et al . 2005 ) . both male and female black carp are broadcast spawners ; females are capable of releasing hundreds of thousands of eggs into flowing water , which then develop in the pelagic zone ( nico et al . 2005 ) . after fertilization , the eggs become semiboyant ( sukhanova , 1967 as cited in nico et al . 2005 ) . they hatch in 1 to 2 days , depending on water temperatures , and the yolk sac is absorbed in 6 to 8 days ( nico et al . 2005 ) . they become sexually mature at 4 to 6 years after which they migrate back to their spawning grounds ( nico et al . 2005 ) . successful reproduction is known only from riverine habitats ( nico et al . 2005 ) . their lifespan probably exceeds 15 years ( biro , 1999 ) .
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but it’s not just about losing the mollusk species — there’s a domino effect . lose the mollusks and there could be poorer water quality because some species clean impurities , as well as losing a food source for fish , muskrats , raccoons , otters and some birds , the conservation department said .
“a great number of species are in danger , species we’ve done a great deal of work to protect , ” chapman said . “now we have this new threat coming in . conceivably , one large fish in the wrong place could cause the extinction of an endangered species in one place in a year , before we even know it’s there . ”
“everyone was trying to help the environment , but it’s turned out to be a mistake , with potentially serious consequences , ” chapman said . it’s unclear how the species escaped from fish farms into the mississippi river , he said , though some blame flooding .
another important step is being mindful of rules involving invasive species and the potential dangers of bringing them into the country , phelps said .
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the nonindigenous occurrences section of the nas species profiles has a new structure . the section is now dynamically updated from the nas database to ensure that it contains the most current and accurate information . occurrences are summarized in table 1 , alphabetically by state , with years of earliest and most recent observations , and the tally and names of drainages where the species was observed . the table contains hyperlinks to collections tables of specimens based on the states , years , and drainages selected . references to specimens that were not obtained through sighting reports and personal communications are found through the hyperlink in the table 1 caption or through the individual specimens linked in the collections tables .
based on asian records , large adults may be more than 1 . 5 m total length and 70 kg or more in weight ; the largest specimen , unconfirmed , from the chang ( yangtze ) river basin reportedly measured 2 . 2 m .
most major pacific drainages of eastern asia from the pearl river ( zhu jiang ) basin in china north to the amur river ( heilong jiang ) basin of china and far eastern russia ; possibly native to the honghe or red rivers of northern vietnam ( nico et al . 2005 ) .
table 1 . states with nonindigenous occurrences , the earliest and latest observations in each state , and the tally and names of hucs with observations† . names and dates are hyperlinked to their relevant specimen records . the list of references for all nonindigenous occurrences of mylopharyngodon piceus are found here .
( richardson , 1846 ) . pages 345 - 365 in p . banarescu ( ed . ) . the freshwater fishes of europe : volume 5 / i , cyprinidae 2 / i . aula - verlag , wiebelsheim , germany .
chapman , d . w . ( editor ) 2006 . early development of four cyprinids native to the yangtze river , china . reston virginia : us geological survey data series 239 . ( available online as urltoken
schofield , p . j . , j . d . williams , l . g . nico , p . fuller , and m . r . thomas . 2005 . foreign nonindigenous carps and minnows ( cyprinidae ) in the united states—a guide to their identification , distribution , and biology . scientific investigations report 2005 - 5041 . u . s . geological survey , tallahassee , florida . 103 p . ( available online at urltoken
nico , l . g . , and m . e . neilson , 2018 , mylopharyngodon piceus ( richardson , 1846 ) : u . s . geological survey , nonindigenous aquatic species database , gainesville , fl , urltoken revision date : 6 / 27 / 2018 , peer review date : 4 / 1 / 2016 , access date : 7 / 9 / 2018
this information is preliminary or provisional and is subject to revision . it is being provided to meet the need for timely best science . the information has not received final approval by the u . s . geological survey ( usgs ) and is provided on the condition that neither the usgs nor the u . s . government shall be held liable for any damages resulting from the authorized or unauthorized use of the information .
the data represented on this site vary in accuracy , scale , completeness , extent of coverage and origin . it is the user ' s responsibility to use these data consistent with their intended purpose and within stated limitations . we highly recommend reviewing metadata files prior to interpreting these data .
citation information : u . s . geological survey . [ 2018 ] . nonindigenous aquatic species database . gainesville , florida . accessed [ 7 / 9 / 2018 ] .
contact us if you are using data from this site for a publication to make sure the data are being used appropriately and for potential co - authorship if warranted . for queries involving fish , please contact pam fuller . for queries involving invertebrates , contact amy benson .
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injurious wildlife doi . fws . fish and aquatic conservation . includes species listed as injurious wildlife under the federal lacey act , which makes it illegal in the u . s . to import , export , or transport between states without a permit . see also : injurious wildlife : a summary of the injurious provisions of the lacey act ( dec 2017 ; pdf | 401 kb )
texas commission on environmental quality , galveston bay estuary program ; houston advanced research center ( harc ) .
invaders factsheet : asian carps ontario ' s invading species awareness program ( canada ) .
pest risk assessment for asian carps in oregon ( dec 15 , 2009 ; pdf | 90 kb ) oregon state library . oregon documents repository . prepared by : portland state university , center for lakes and reservoirs
integrated taxonomic information system . mylopharyngodon piceus . [ accessed jan 16 , 2016 ] .
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if possible , please take one or more photos of the invasive species you are reporting . also make note of the location , date and time of the observation . this will aid in verification of your report . you may be asked to provide your name and contact information if follow - up is needed .
habitat : large rivers and lakes but require large rivers for reproduction ( water current keeps their eggs from sinking to the bottom ) .
diet : their diet consists primarily of mussels and snails , but also includes freshwater shrimp , crayfish , and insects .
u . s . distribution : reported in arkansas , illinois , louisiana , mississippi and missouri .
potential means of introduction : illinois river or flood connections with great lakes waters .
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thank you for taking the time to provide feedback on the iucn red list of threatened species website , we are grateful for your input .
has been assessed as data deficient , due to the lack of information regarding species population size , current population , impacts of threats and harvest trends .
has originally an east asian distribution , from in most pacific river drainages , from the amur river to the west river ( xi jiang ) ( kottelat and freyhof 2007 ) . introduced in many countries worldwide for control populations of molluscan vectors of fish and human parasites . furthermore , used to removed dreissena mussels that clog hydroelectric plants .
naturally reproducing populations established only in amu darya ( turkmenistan ) and possible in tone drainages ( japan ) ( freyhof and kottelat 2007 ) .
in the second half of the 20th century , a massive decline in the abundance of this species was observed in its native distribution range . its current population trend is unknown .
inhabits large lowland river and lakes , preferably with clear water and high oxygen concentration ( kottelat and freyhof 2007 ) .
spawns for the first time at 6 - 11 years , females later than males ( at about 1000 mm sl and 15 kg , males at 900 mm and 11 kg , fecundity is about 700 - 800 thousand eggs ) . migrates upriver and spawns in open water during flood phase . eggs are pelagic or semipelagic and hatch while drifting downstream . if the river flow is blocked or if available river stretches are too short , eggs cannot drift for long enough and fail to develop . larvae migrate into floodplain lakes and channels with little or no current . larvae feed on zooplankton , then on ostracods and aquatic insects . at about 120 mm sl , juveniles start to feed on small snails and clams . larger juveniles and adults feed almost entirely on molluscs ( source : kottelat and freyhof 2007 ) .
major threats to this species are overfishing , river modifications such as dam construction and the conversion of floodplains into agriculture land and water pollution .
it is not known if there are any conservation measures in place . more research is needed .
to make use of this information , please check the < terms of use > .
photographer : rob cosgriff affiliation : illinois natural history survey source : urltoken copyright : ( cc by - ny 3 . 0 )
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illinois dnr : 217 . 557 . 0719 or 618 . 462 . 0362 southern illinois university : 618 . 453 . 6089 u . s . geological survey : 573 . 876 . 1866


the large tree finch ( camarhynchus psittacula ) is a species of bird in the darwin 's finch group of the tanager family thraupidae .

it is endemic to the galapagos islands .

its natural habitats are subtropical or tropical dry forests and subtropical or tropical moist montane forests .

large tree finch

large tree finch ( camarhynchus psittacula ) is a species of bird in the thraupidae family .
protection / threats / status : the large tree - finch is common within its range , and its populations appear to be stable . this species is not currently threatened .
the large tree - finch is the largest and heaviest bodied of the three tree - finch species , imaginatively named the large , medium and small tree - finches . large tree - finches have a big and deep bill with a strongly arched culmen , being approximately as long as it is deep . the fine tips of the mandibles actually cross a feature that is difficult to see on live birds . males show a dark hood , greenish back and whitish underparts . this species is found in a number of the islands in the galapagos archipelago , and in many it is sympatric with the small tree - finch ( c . parvulus ) . it is never as common as the small tree - finch and it is found in areas with taller and larger trees .
habitat : the large tree - finch frequents mainly humid evergreen forest between 300 and 700 metres of elevation . however , during the dry season , it may move to lower areas with deciduous trees , shrubs and opuntia cacti .
the breeding season is associated with rains , involving abundant food resources . the large tree - finch is monogamous and the pair defends a small territory . they often have long - term pair - bonds . the male is territorial and guards the female during the egg - laying .
introduction : the large tree - finch is the largest of the genus camarhynchus . this is mainly an arboreal species and an insect - eater . both scientific and french names could illustrate the repetitive song of this species , but also its strongly arched bill a bit similar to that of a parrot .
calls and songs : sounds by xeno - canto the large tree - finch’s call is a nasal “tzeeuu” . the song is a series of repeated notes “chu - tzee chu - tzee chut - zee” . this song can be given by both mates . like in other darwin’s finches , the song includes various trills and buzzes .
conservation and research actions underway no actions are currently known . conservation and research actions proposed implement a full - scale monitoring programme across the galápagos islands . ensure that management activities to control invasive alien plant species do not have a negative impact on large tree - finch . investigate drivers behind observed declines and assess the impact of philornis downsi on the population . protect and enhance existing habitat .
behaviour in the wild : the large tree - finch feeds primarily on arthropods , but it also takes cactus fruits and other fruits , flowers and seeds . during the nesting season , the chicks are fed with a mixed diet including arthropods , fruits and seeds . outside the breeding season , it feeds mainly on seeds according to the size of its bill . it forages in trees , probing and biting into the bark of twigs , in order to extract insects and larvae , but also caterpillars .
identification : the largest of the tree finches with a large , rather parrot - like bill , the tips of the mandibles crossing . adult male : head , neck , breast and mantle black when fully mature , the remainder of the upperparts being olive - grey with some dark streaking . underparts pale , often with a yellow tinge . female / immature : upperparts olive - grown with faint streaking . underparts paler and virtually unstreaked .
13 cm , largest tree - finch . deep bill , approximately as long as it is deep . mandible tips cross slightly when bill closed . male has upperparts greyish - olive and whitish below , with blackish hood . female is dull greyish brown ( jaramillo and sharpe 2015 ) . voice song a repeated series of 4 - 6 notes given in pairs
chu - tzee chu - tzee chut - zee
. call includes a nasal
tzeeuu
.
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thank you for taking the time to provide feedback on the iucn red list of threatened species website , we are grateful for your input .
sacc . 2005 and updates . a classification of the bird species of south america . available at : urltoken .
ashpole , j , butchart , s . , ekstrom , j . , fisher , s . , harding , m . , sharpe , c . j .
justification : this species has been uplisted to vulnerable . the species has declined significantly on the island of santa cruz and is likely to also be declining on other islands within its range , owing to habitat loss and degradation .
this species is endemic to the galápagos islands , ( ecuador ) , with breeding populations on isabela , santa cruz , santa fé , fernandina , santiago , marchena , pinta and rábida ( castro and phillips 1996 , stotz et al . 1996 ) . it is extinct on pinzón ( castro and phillips 1996 ) and thought to be extinct on floreana ( kleindorfer et al . 2014 , jaramillo and sharpe 2015 ) .
the global population size has not been quantified , but this species is described as ' uncommon ' in at least parts of its range ( stotz et al . 1996 ) . on the island of santa cruz its population has been estimated at 9 , 000 singing males ( dvorak et al . 2012 ) . however no data exists for the islands of isabela and santiago . trend justification : the population is suspected to be decreasing rapidly . on the island of santa cruz , the species reportedly declined significantly in the dry zone between 1997 and 2010 , but not in the scalesia zone ( dvorak et al . 2012 ) . habitat alteration , introduced pathogens and parasites and changes in insect availability may have contributed to declines . on the islands of isabela and santiago , where native forest has also been degraded ( by introduced herbivores ) population declines are also suspected ( dvorak et al . 2012 ) .
this species inhabits lowland deciduous and montane evergreen forest , between 300 and 700 m altitude ( stotz et al . 1996 ) . on santa cruz in the dry zone it is restricted to areas with tall palo santo bursera graveolens trees , it is also found in the scalesia zone and locally in the agricultural zone ( dvorak et al . 2012 ) . it feeds on the fruits of native plant species , and on insects for which it forages under leaves and excavates dead branches ( castro and phillips 1996 ) . it may move to lower elevations during the dry season ( jaramillo and sharpe 2015 ) .
the species is likely to be affected by a number of threats . development , introduced herbivores , the spread of invasive alien plant species and the herbicides used to manage these invasions may all have contributed to unfavourable habitat conditions for the species on santa cruz ( dvorak et al . 2012 ) . the introduced parasitic fly philornis downsi is known to have a negative impact on nesting success in galápagos finches and the species may be susceptible to avian pox . severe weather and changes in rainfall patterns owing to climate change also pose a threat .
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) , in neotropical birds online ( t . s . schulenberg , editor ) . cornell lab of ornithology , ithaca , ny , usa . retrieved from neotropical birds online :
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the adult male of the nominate race has greyish - olive upperparts with blackish feather centres . the tail is short . the underparts are whitish with yellowish - buff wash . lower breast and flanks are streaked dark . the undertail - coverts are buffy - white and unstreaked . the head is dark , forming a blackish hood extending down to throat and breast . the deep , relatively long bill has strongly arched culmen . it is black when breeding , and dull orange with dark culmen outside the breeding season . the eyes are dark brown . legs and feet are black .
the female has duller greyish - brown upperparts . the upperwing is brownish with two narrow pale wingbars . the underparts are whitish with indistinct dark streaking on breast , variable according to each bird . from belly to undertail - coverts , the plumage is plain pale buff . the head is greyish - brown with pale supercilium . the bill is dull orange with darker culmen . the eyes are dark . legs and feet are blackish .
the immature male resembles female , with blackish forehead , face and lower throat .
subspecies and range : there are three subspecies . c . p . habeli occurs on pinta and marchena islands , in n galapagos islands . this race is smaller than nominate . the male is darker too . the bill is longer , with less arched culmen .
c . p . affinis is found on fernandina and isabela islands , in w galapagos islands . this one is smaller than nominate , with smaller bill too .
c . p . psittacula ( here described and displayed ) occurs on santiago , rábida , santa cruz , santa fe and floreana , in c and s galapagos islands .
this species is resident in its range , only performing some altitudinal movements during the dry season .
reproduction of this species : the breeding season occurs during the wet period . the male builds the nest , a spherical structure with lateral entrance towards the top . the nest is made with dry grasses , moss and lichen .
the female lays 4 whitish eggs with darker spots . she incubates alone during 12 days . the chicks are fed by both parents . they fledge about 13 - 15 days after hatching .
darwin finches , or galapagos finches , are small land birds with generally dull black , brown or olive , often streaky , plumage ; short tails ; and short , rounded wings . their bills vary greatly in size and shape ( a fact which was instrumental in inspiring charles darwin’s thinking in relation to the theory of evolution – and hence the name given to this fascinating group of species ) .
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camarhynchus psittacula psittacula : galapagos islands ( seymour , barrington , santa cruz , floreana , pinzón , rábida , santiago is . )
howard and moore 4th edition ( incl . corrigenda vol . 1 - 2 ) :
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gustavo jiménez - uzcátegui , david a . wiedenfeld , f . hernán vargas , howard l . snell .
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idiosoma nigrum , also called black rugose trapdoor spider , occurs only in south-western australia , in dry woodlands east of the darling scarp and north to moore river .

females can reach a length of about 30 mm , males about 18 mm .

i. nigrum digs burrows up to 32 cm deep .

idiosoma nigrum

ecologia environment ( 2013 ) . blue hills idiosoma nigrum targeted survey 2012 . sinosteel midwest corporation . urltoken
ecologia environment ( 2010a ) . weld range idiosoma nigrum population genetic study . sinosteel midwest corporation . available from : urltoken .
idiosoma nigrum main , 1952 : 133 , pl . i , f . 2 - 5 , f . 2c ( d f ) . idiosoma nigrum main , 1957a : 439 , f . 2g - h , 9d , 14b , 24b ( d m ) . idiosoma nigrum main , 1985a : 13 , f . 23 , 27 , 215 ( m f ) . idiosoma nigrum rix et al . , 2018b : 18 , f . 1 - 3 , 26 - 56 ( m f ) .
shield - backed trapdoor spider ( idiosoma nigrum ) conservation plan ( avon catchment council ( acc ) , 2007 ) [ state species management plan ] .
ecologia environment ( 2009a ) . shield - back spider idiosoma nigrum survey . weld range iron ore project . sinosteel midwest corporation . available from : urltoken .
gray , m . ( 2014 ) . idiosoma nigrum ( family idiopidae ) . species bank . australian biological resources study , canberra . available from : urltoken .
anonymous ( 2010 ) . idiosoma nigrum . form to nominate a western australian species for listing as threatened , change of category or delisting . available from : urltoken .
avon catchment council ( acc ) ( 2007 ) . shield - backed trapdoor spider ( idiosoma nigrum ) conservation plan . avon catchment council , western australia . available from : urltoken .
citation : department of the environment ( 2018 ) . idiosoma nigrum in species profile and threats database , department of the environment , canberra . available from : urltoken . accessed tue , 10 jul 2018 05 : 15 : 41 + 1000 .
main , b . y . ( 2003 ) . demography of the shield - back trapdoor spider idiosoma nigrum main in remnant vegetation of the western australian wheatbelt . records of the south australian museum , monograph series . 7 : 179 - 185 .
the closest relatives to the shield - backed trapdoor spider are idiosoma sigillatum and idiosoma hirsutum , both of which lack the hardened shield on the abdomen ( anonymous 2010 ) .
in synonymy : idiosoma hirsutum main , 1952 = idiosoma sigillatum ( o . pickard - cambridge , 1870 ) ( rix et al . , 2017a : 593 ) . idiosoma pulleinei ( hogg , 1902 ) = idiosoma subtriste ( o . pickard - cambridge , 1877 ) ( main , 1957a : 428 , sub aganippe ) . idiosoma schomburgki ( karsch , 1878 , t from idiommata ) = idiosoma subtriste ( o . pickard - cambridge , 1877 ) ( main , 1985a : 12 , sub aganippe ; placed by raven , 1985a : 161 in misgolas ) .
threatened species scientific committee ( tssc ) ( 2013cb ) . commonwealth listing advice on idiosoma nigrum ( shield - back trapdoor spider ) . canberra : department of sustainability , environment , water , population and communities . available from : urltoken . in effect under the epbc act from 14 - may - 2013 .
department of sustainability , environment , water , population and communities ( 2013g ) . approved conservation advice for idiosoma nigrum ( shield - back spider ) . canberra : department of sustainability , environment , water , population and communities . available from : urltoken . in effect under the epbc act from 14 - may - 2013 .
idiops sigillatus o . pickard - cambridge , 1870c : 105 , pl . 8 , f . 1 ( d m ) . idiosoma sigillatum ausserer , 1871a : 150 . acanthodon sigillatum simon , 1892a : 91 . idiosoma sigillatum pocock , 1897a : 109 ( d f ) . idiosoma sigillatum simon , 1903a : 901 , f . 1053 ( f ) . idiosoma hirsutum main , 1952 : 132 , f . 2b ( d f ) . idiosoma sigillatum main , 1952 : 131 , f . 1a - c , 2a ( m ) . idiosoma hirsutum main , 1957a : 440 , f . 15a - b ( considered a possible hybrid of i . sigillatum x aganippe rhaphiduca rainbow & pulleine , 1918 ) . idiosoma sigillatum main , 1964 : 32 , f . a - d , f ( m ) . idiosoma hirsutum main , 1985a : 54 ( considered a valid species ) . idiosoma sigillatum main , 1985a : 14 , f . 26 , 190 , 216 ( m f ) . idiosoma sigillatum rix et al . , 2017a : 593 , f . 83 - 84 , 87 , 91 , 93 , 97 ( m , s of i . hirsutum ) . idiosoma sigillatum rix et al . , 2018b : 64 , f . 4 - 6 , 351 - 372 ( m f ) .
in synonymy : aganippe o . pickard - cambridge , 1877 = idiosoma ausserer , 1871 ( rix et al . , 2017a : 590 ) anidiops pocock , 1897 = idiosoma ausserer , 1871 ( rix et al . , 2017a : 590 )
the shield - backed trapdoor spider ( idiosoma nigrum ) belongs to the suborder mygalomorphae , commonly known as “trapdoor” and “funnel - web” spiders . they are primarily terrestrial burrowing spiders which occasionally make tubular silk nests on tree trunks . mygalomorphs are able to persist in small isolated areas due to their low dispersion powers , long life cycle and sedentary life style ( main , 1987a ) .
idiosoma gardneri rix & harvey , in rix et al . , 2018b : 38 , f . 167 - 179 ( d m ) .
idiosoma gutharuka rix & harvey , in rix et al . , 2018b : 39 , f . 180 - 192 ( d m ) .
idiosoma incomptum rix & harvey , in rix et al . , 2018b : 41 , f . 193 - 205 ( d m ) .
idiosoma kwongan rix & harvey , in rix et al . , 2018b : 53 , f . 272 - 284 ( d m ) .
idiosoma clypeatum rix & harvey , in rix et al . , 2018b : 25 , f . 79 - 100 ( d m f ) .
idiosoma corrugatum rix & harvey , in rix et al . , 2018b : 30 , f . 101 - 122 ( d m f ) .
idiosoma dandaragan rix & harvey , in rix et al . , 2018b : 32 , f . 132 - 144 ( d m f ) .
idiosoma formosum rix & harvey , in rix et al . , 2018b : 35 , f . 145 - 166 ( d m f ) .
idiosoma intermedium rix & harvey , in rix et al . , 2018b : 43 , f . 206 - 227 ( d m f ) .
idiosoma mcnamarai rix & harvey , in rix et al . , 2018b : 57 , f . 307 - 328 ( d m f ) .
idiosoma jarrah rix & harvey , in rix et al . , 2018b : 46 , f . 7 , 228 - 249 ( d m f ) .
idiosoma kopejtkaorum rix & harvey , in rix et al . , 2018b : 50 , f . 9 , 250 - 271 ( d m f ) .
idiosoma mcclementsorum rix & harvey , in rix et al . , 2018b : 55 , f . 8 , 285 - 306 ( d m f ) .
idiosoma schoknechtorum rix & harvey , in rix et al . , 2018b : 60 , f . 10 , 329 - 350 ( d m f ) .
idiosoma arenaceum rix & harvey , in rix et al . , 2018b : 23 , f . 11 - 12 , 57 - 78 ( d m f ) .
idiosoma galeosomoides rix , main , raven & harvey , in rix et al . , 2017a : 599 , f . 99 , 116 - 128 ( d f ) .
aganippe winsori faulder , 1985 : 89 , f . 9a - c ( d m ) . idiosoma winsori rix et al . , 2017a : 601 ( t from aganippe ) .
aganippe cupulifex main , 1957a : 436 , f . 7f , 9b , 12a , 13c ( d m f ) . idiosoma cupulifex rix et al . , 2017a : 599 ( t from aganippe ) .
aganippe castellum main , 1986b : 101 , f . 2 , 4a - h ( d m f ) . idiosoma castellum rix et al . , 2017a : 594 , f . 89 , 92 , 95 ( m , t from aganippe ) .
aganippe montanus faulder , 1985 : 85 , f . 5a - c , 6a - c , 7a - c , 8a - b ( d m f ) . idiosoma montanum rix et al . , 2017a : 601 ( t from aganippe ) .
aganippe planites faulder , 1985 : 91 , f . 10a - c , 11a - c , 12a - c , 13a - b ( d m f ) . idiosoma planites rix et al . , 2017a : 571 ( t from aganippe ) .
aganippe occidentalis hogg , 1903b : 309 , f . 1 - 2 ( d m ) . aganippe occidentalis main , 1957a : 414 , f . 11b ( m , d f ) . idiosoma occidentale rix et al . , 2017a : 571 ( t from aganippe ) .
nomina dubia : idiosoma modestum ( rainbow & pulleine , 1918 : 98 , pl . 21 , f . 47 - 48 , f , australia ( south australia ) , originally in aganippe ) [ urn : lsid : nmbe . ch : spidersp : 000633 ] - - rix et al . , 2017a : 592 . idiosoma pelochroa ( rainbow & pulleine , 1918 : 100 , pl . 21 , f . 51 , f , originally in aganippe , australia ( south australia ) ) [ urn : lsid : nmbe . ch : spidersp : 000636 ] - - rix et al . , 2017a : 592 . idiosoma robustum ( rainbow & pulleine , 1918 : 97 , pl . 21 , f . 45 - 46 , f , originally in aganippe , australia ( south australia ) [ urn : lsid : nmbe . ch : spidersp : 000639 ] - - rix et al . , 2017a : 592 . idiosoma simpsoni ( hickman , 1944a : 22 , f . 7 - 10 , f , originally in aganippe , australia ( south australia ) ) [ urn : lsid : nmbe . ch : spidersp : 000640 ] - - rix et al . , 2017a : 592 . idiosoma whitei ( rainbow , 1915 : 774 , pl . 67 , f . 1 - 2 , f , originally in aganippe , australia [ south australia ] ) - - rix et al . , 2017a : 590 , contra main , 1957a : 426 , sub anidiops .
aganippe smeatoni hogg , 1902 : 126 , f . 23 , pl . 13 , f . 7 ( d m ) . aganippe smeatoni simon , 1903a : 901 , f . 1054 - 1055 ( m ) . aganippe smeatoni main , 1957a : 429 , f . 11a ( m , d f ) . idiosoma smeatoni rix et al . , 2017a : 601 ( t from aganippe ) .
aganippe berlandi rainbow , 1914a : 199 , f . 9 - 13 ( d m ) . aganippe berlandi faulder , 1985 : 83 , f . 1a - c , 2a - c , 3a - c , 4a - b ( m , d f ) . aganippe berlandi main , 1985a : 51 ( not a junior synonym of a . smeatoni hogg , 1902 ) . idiosoma berlandi rix et al . , 2017a : 594 , f . 96 ( m , t from aganippe ) .
aganippe rhaphiduca rainbow & pulleine , 1918 : 93 , pl . 21 , f . 38 - 42 ( d m f ) . aganippe raphiduca main , 1957a : 433 , f . 2e - f , 7d , 12b - c , 13d , 26a - g , 27a - c ( m ) [ 13a - b is eucanippe nemestrina per rix et al . , 2018 : 153 ] . aganippe raphiduca main , 1964 : 34 , f . e - h ( m ) . idiosoma rhaphiduca rix et al . , 2017a : 571 ( t from aganippe ) .
anidiops manstridgei pocock , 1897a : 114 ( d f ) . anidiops manstridgei simon , 1903a : 903 , f . 1052 . anidiops manstridgei rainbow & pulleine , 1918 : 101 , pl . 21 , f . 52 - 54 ( f ) . anidiops manstridgei main , 1957a : 426 , f . 2c - d , 3b , 5a , 10a - c ( m , s ) . anidiops manstridgei main , 1985a : 16 , f . 20 - 21 , 192 ( f ) . idiosoma manstridgei rix et al . , 2017a : 600 , f . 132 - 144 ( m , t from anidiops ) .
aganippe subtristis o . pickard - cambridge , 1877c : 28 , pl . 6 , f . 3 ( d f ) . idiommata schomburgki karsch , 1878c : 820 ( d m ) . aganippe subtristis simon , 1892a : 106 , f . 104 . aganippe pulleinei hogg , 1902 : 128 , f . 24 , pl . 13 , f . 3 - 4 ( d m f ) . aganippe subtristis simon , 1903a : 901 , f . 1050 . aganippe subtristis rainbow & pulleine , 1918 : 91 , pl . 21 , f . 32 , 35 - 37 ( d m ) . aganippe subtristis main , 1957a : 428 , f . 7c ( m , s ) . aganippe subtristis main , 1964 : 34 , f . a - d ( m ) . aganippe subtristis main , 1985a : 12 , f . 13 , 15 - 18 , 24 - 25 , 188 - 189 ( m f , s ) . idiosoma subtriste rix et al . , 2017a : 601 ( t from aganippe ) .
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for information to assist regulatory considerations , refer to policy statements and guidelines , the conservation advice , the listing advice and / or the recovery plan .
department of sustainability , environment , water , population and communities ( 2013 ) .
. canberra : department of sustainability , environment , water , population and communities . available from :
recovery plan not required , the approved conservation advice for the species provides sufficient direction to implement priority actions and mitigate against key threats ( 26 / 04 / 2013 ) .
the distribution shown is generalised from the departments species of national environmental significance dataset . this is an indicative distribution map of the present distribution of the species based on best available knowledge . some species information is withheld in line with sensitive species polices . see map caveat for more information .
there is currently some discussion about whether the species found in the rangelands of the midwest region is the same as that found in the wheatbelt and the coastal regions of the midwest . there appears to be some taxonomic differences in the male morphology ( framenau pers . comm . cited in anonymous 2010 ) and these are currently being investigated ( anonymous 2010 ) . there are only two significant populations in the wheatbelt , east yorkrakine and minnivale , and these would be particularly important if the species is polyphyletic ( i . e . if it has a northern and southern species ) ( anonymous 2010 ) .
the shield - backed trapdoor spider is dark brown to black , large ( females up to 30 mm in body length ) and with a distinctive thick and hard cuticle on the abdomen . the end of the abdomen is flattened into a shield and the sides are deeply corrugated . the burrows always have a lightweight , leaf litter and silk door , with leaf and twig trip - lines fanning out from the centre of the front of the burrow rim ( gray 2014 ) .
the shield - backed trapdoor spider is endemic to semi - arid south - west western australia ( wa ) . it occurs in a number of severely fragmented populations in the central and northern wheatbelt ( e . g . minnivale and east yorkrakine ) ( main et al . 2000 cited in tssc 2013ca ) . further north , the species occurs in more arid areas in the midwest ( e . g . large isolated ranges at jack hills ( tssc 2013ca ) , weld range ( ecologia environment 2009a ) and blue hills ( ecologia environment 2013 ) ) and coastal areas of the midwest ( e . g . zuytdorp station north of the murchison river and nanga station south of shark bay ) ( main et al . 2000 cited in anonymous 2010 ) . the arid midwest populations are naturally fragmented or isolated because they persist only on ranges , but the wheatbelt and coastal midwest populations are all severely fragmented as a result of land clearing ( anonymous 2010 ) .
the avon catchment council ( 2007 ) includes a number of eastern records ( e . g . at westonia in 2007 ) that are not reported elsewhere ( e . g . tssc 2013cb ) . post - 1980 , verified records only extend as far east into the wheatbelt as durokoppin and trayning ( ala 2014 ) .
in 2010 , there were around 6100 burrows databased , although some of these would have been inactive or burrows of juveniles . the overwhelming majority ( around 5400 ) of these are from midwest ranges and were recorded as part of the environmental impact assessment process . many wheatbelt records , outside of the main populations at minnivale and east yorkrakine , were recorded decades ago and many of these are likely to be extinct ( anonymous 2010 ) .
the species extent of occurrence is approximately 21 000 km² . it is highly likely that the species occurs throughout much of the midwest region in association with large ranges with deep gullies and possibly with breakaways ( anonymous 2010 ) .
the species occurs in three areas at weld range ( weld range north , weld range south and hampton hill ) over an area of 15 km . in one study , the mean number of burrows per hectare in the area ranged between 11 . 3 - 43 . 9 ( ecologia environment 2009a ) . genetic studies indicated a strong population subdivision between weld range north and weld range south ( ecologia environment 2010a ) . the collections at weld range extended the species distribution by approximately 200 km further north , into more arid areas ( ecologia environment 2009a ) .
at blue hills , one study showed the mean number of burrows per hectare of 4 . 08 in an area of 13 ha ( ecologia environment 2013 ) .
survey work has been carried out at minnivale and east yorkrakine nature reserves , karara , weld range and jack hills mining leases and , more recently , on likely conservation areas and department of environment and conservation managed lands through the northern wheatbelt and the southern / central midwest ( anonymous 2010 ) . these surveys did not detect the species in much of the northern wheatbelt and southern midwest . these surveys demonstrated that there are unlikely to be any populations in the interior of the wheatbelt and midwest until you reach the large banded iron formation ranges of the midwest to the north and the higher rainfall of the central wheatbelt to the south ( anonymous 2010 ) .
this work also showed that there is a likelihood that the species occurs throughout more of the midwest region than currently known , with up to 60 sites identified as possibilities ( anonymous 2010 ) . however more than half of these occur east of the current known eastern extent , and a number of mining tenements in these areas have not found the species despite significant survey work . at least half a dozen occur further north than the northern known extent . there are about 20 sites that have a high potential for harbouring significant populations of this species ( anonymous 2010 ) .
in 2010 , there were seven locations with populations larger than 30 shield - backed trapdoor spider individuals ( anonymous 2010 ) . ecologia environment ( 2009a ) estimated that there is over 1000 ha of suitable habitat in weld range that could support over 20 000 individuals . based on detected adults , the site is estimated to have an effective population size of approximately 4000 with wilgie mia and weld range north supporting the largest effective populations 2300 and 1000 , respectively ( ecologia environment 2009a ) . proposed mining activity in the area is estimated to impact 11 % of the population ( ecologia environment 2009a ) .
total population reduction has not been investigated , but data from a study in east yorkrakine reserve from 1989 to 1999 showed a 95 % reduction in abundance at the site ( main 2003 ) . future reductions are possible due to ongoing threats in the wheatbelt and mining in the vicinity of populations at karara , weld range and jack hills ( anonymous 2010 ) .
in the wheatbelt , the shield - backed trapdoor spider typically inhabits clay soils whereas the arid midwest populations are associated with rocky habitats , primarily in positions with increased moisture retention properties like gullies and drainage lines on southern facing slopes ( anonymous 2010 ; ecologia environment 2009a ) . the wheatbelt and coastal midwest populations are in areas with more consistent annual rainfall than those in the arid midwest , which is likely to be why the populations in these areas are primarily found in sheltered habitat ( anonymous 2010 ) . in the wheatbelt , populations are associated with eucalypt woodland and acacia shrubland , and in the arid midwest they are associated with acacia shrubland ( anonymous 2010 ) .
leaf litter and twigs are extremely important to the species as it provides material for the burrows , reduced soil moisture loss and increased prey availability ( anonymous 2010 ) . the species avoids areas of dense leaf litter as juveniles are unable to dig their initial hole in such areas ( main 1992 cited in ) . areas of lower grazing pressure may suport greater population abundance ( ecologia environment 2009a ) .
in the wheatbelt , habitat critical to the species is identified as open york gum ( eucalyptus loxophleba ) , salmon gum ( e . salmonophloia ) and wheatbelt wandoo ( e . capillosa ) woodland , where jam ( acacia acuminata ) forms a sparse understorey in heavy clay soils ( acc 2007 ) .
in a study at weld range , all burrows were found within boundaries of drainage lines underneath predominantly acacia vegetation ( mulga ( acacia aneura ) , a . sp . weld range , dead finish ( a . tetragonophylla ) , a . ramulosa , a . craspedocarpa , a . paraneura ) and also underneath hakea preissii and eremophila glutinosa ) . of 1708 burrows detected , 33 % were on slope - foot - plain , 30 % on the plain , 27 % on slopes and 10 % on hilltop - slope - foot ( ecologia environment 2009a ) . burrows were found in soil dominated by clay and rocks ( 54 % ) , or clay , rock and sand ( 38 % ) . the average leaf litter of sites ranged between 7 - 100 % with a mean of 59 % ( ecologia environment 2009a ) .
in a study at blue hills , 53 burrows were recorded under a . ramulosa or a . caesaneura , in microhabitat ranging from loamy plains to rocky hillslopes ( ecologia environment 2013 ) . the average leaf litter of sites ranged between 25 - 98 % with a mean of 63 % ( ecologia environment 2013 ) .
in a study at mummaloo , of a 52 quadrat survey the species was detected at 14 locations ( bennelongia 2012 ) . two records occurred in ecualyptus woodland and twelve occurred in mixed species shrubland . the eucalypt woodland consists of york gum , salmon gum and gimlet ( e . salubris ) with callitris columellaris usually present and occasionally casuarina obesa . the understorey contains 5 - 60 % cover of eremophila spp . , acacia spp . and allocasuarina spp . . the mixed species shrubland consists of melaleuca stereophloia , callitris columellaris and casuarina obesa ( of no more than 55 % overall cover ) , and 10 - 100 % cover of shrub species , especially kimberly ' s wattle ( acacia anthochaera ) and allocasuarina acutivalvis subsp . prinsepiana ( bennelongia 2012 ) .
the shield - backed trapdoor spider is protected from dessication partly through a deep burrow that extends into humid soil . additionally it has an extraordinary thickened abdominal integument ( tough outer protective layer ) . this is dorsally corrugated and posteriorly truncated with large button - like sigilla ( seal ) and in some parts of the geographic range has stout spines along the ridges . this morphology reduces evaporative water loss in contrast to most species in the related genus aganippe . the enlarged eyes and long legs together with the attached twiglines to the buttom rim are advantageous for foraging ( main 2010 ) .
spiderlings generally construct their burrows within a very short distance ( several centimetres ) of the matriarch female , forming a family cluster that is typical of mygalomorph spiders ( a group of large spiders that include tarantulas , trapdoor spiders and funnel - web spiders ) that do not have aerial dispersal . gene flow is maintained through the dispersal of mature males in search of females for mating ( travelling up to 500 m ) the only time males leave their burrows . females spend their entire life in the one burrow or within its proximity ( anonymous 2010 ) . the population at weld range north was shown to have uninterrupted gene flow over a range of at least 6 km ( ecologia environment 2010a ) .
both males and females reach maturity after approximately 5 - 6 years . males die shortly after reaching sexual maturity and mating , whereas females may live as long as 20 years , reproducing several times ( anonymous 2010 ) . it is believed that males mature and mate after the first significant rains of the year , dispersing up to 500 m ( main unpub . data cited in anonymous 2010 ) . there is some evidence that females may store sperm ( main unpub . data cited in anonymous 2010 ) but whether this means that males only mate with virgin females or whether adult females mate repeatedly during their life is unclear . it is also unknown whether males mate within their matriarchal unit and whether they mate with more than one female ( anonymous 2010 ) . the very low dispersal capabilities of the males mean that fragmentation and isolation are likely to be playing a major role in the declining populations in the wheatbelt ( anonymous 2010 ) .
the shield - backed trapdoor spider is an opportunistic feeder and feeds primarily on ants , but also includes beetles , cockroaches , millipedes and moths ( clark & spier - ashcroft 2003 ) . the species relies on the twigs and leaves they have attached to the rim of their burrow for the detection of prey within the vicinity of their burrow ( anonymous 2010 ) . this reliance on leaf litter means that leaf litter loss through inappropriate fire regimes and management may impact significantly on the species ability to feed ( anonymous 2010 ) .
searching for burrows during the wetter parts of the year is the most common and most effective way of surveying for the shield - backed trapdoor spider . the burrow is quite distinctive but can be easily mistaken for other twig - lining species . the burrow has two tufts of leaf litter radiating out from the centre of the burrow rim , similar to a moustache . the atrium of the burrow is cup shaped and narrows to the main shaft of the burrow . it is this characteristic that is often missed or misinterpreted by surveyors ( anonymous 2010 ) . searching for burrows within and on the edges of leaf litter in suitable habitats is the most effective approach for detection . pit traps placed around vegetation in suitable habitats during autumn can be used to catch wandering males ( anonymous 2010 ) .
secondary salinisation : the widespread clearing of the wheatbelt has resulted in the water table rising and an increase in salinisation close to the surface . this results in vegetation changes that directly affect the shield - backed trapdoor spider because of it’s reliance on the vegetation and associated leaf litter for habitat ( anonymous 2010 ) .
grazing : grazing by stock and feral animals affects both the wheatbelt and midwest populations largely through the disturbance of leaf litter , vegetation and soil . work in the midwest has shown that areas where grazing occurs have fewer emergents and juveniles ( ecologia environment unpub . data cited in anonymous 2010 ) .
fragmentation and clearing : the clearing of habitat has resulted in the severe fragmentation of populations in the wheatbelt . the populations at karara , weld range , jack hills and blue hills will all be negatively affected by land clearing because of the mining operations ( anonymous 2010 ; ecologia environment 2009a , 2013 ) , which will fragment these significant populations . dust pollution associated with mining could negatively impact the species
vibrations : vibrations associated with vehicles and exploration drilling have the potential to affect nearby populations . recent work at jack hills and weld range has shown a possible reduction in emergents and juveniles within 50 m of exploration drilling pads ( phoenix environmental unpub . data cited in anonymous 2010 ) . exploration restrictions have been put in place at weld range and jack hills based on this research .
inappropriate fire : in the wheatbelt , the combination of fragmentation and intense fire has a high potential to result in local extinctions , with little to no chance of recolonisation ( main 1995 ) . intense fires can not only remove burrow doors but also remove all the leaf litter , providing no material for reparation work and dramatically affecting the prey population ( anonymous 2010 ) .
lack of litter management in reserves : although the species requires leaf litter to survive , excessive , deep litter canrestrict the establishment of emergent burrows , forcing them further away from vegetation and exposing them to the elements that likely decrease their chances of surviving to adulthood . similarly deep litter reduces the chances of understorey vegetation growing , reducing the diversity of invertebrates and the health of the habitat and increases the chances of hot , intense fires going through a population ( anonymous 2010 ) .
recovery actions for this species are provided in the conservation advice for the shield - backed trapdoor spider ( tssc 2013ca ) and the avon catchment shield - backed trapdoor spider conservation plan ( acc 2007 ) at the start of the profile .
atlas of living australia ( ala ) ( 2014 ) . atlas of living australia . available from : urltoken .
bancroft , w . & m . bamford ( 2012 ) . karara iron ore project : annual monitoring survey of the shield - backed trapdoor spider 2010 to 2012 . prepared for : karara mining limited .
bennelongia ( 2012 ) . mummaloo hill project : short - range endemic invertebrates . prepared for top iron . available from : urltoken .
clarke , g . & f . spier - ashcroft ( 2003 ) . a review of the conservation status of selected australian non - marine invertebrates . environment australia , canberra . available from : urltoken .
main , b . y . ( 1992 ) . the role of life history patterns and demography of mygalomorph trapdoor spiders for assessing persistence in remnant habitats of the western australian wheatbelt . report for the world wide fund for nature , world wide fund for nature , sydney .
main , b . y . ( 1995 ) . survival of trapdoor spiders during and after fire . calm science supplement . 4 : 207 - 216 .
main , b . y . ( 2010 ) . interactions of water , plants and ground - dwelling fauna : water harvesting and tapping by trapdoor spiders . landscapes : the journal of the international centre for landscape and language . 4 ( 1 ) .
anonymous ( 2009 ) . australian faunal directory . australian biological resources study . available from : urltoken .
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urn : lsid : biodiversity . org . au : afd . taxon : 695e1e77 - a2b7 - 4786 - 9502 - 81be54ac63d9
urn : lsid : biodiversity . org . au : afd . taxon : a5fba25c - 9301 - 4111 - bd1d - f020b12a5b0a
urn : lsid : biodiversity . org . au : afd . taxon : e67764dc - eda3 - 4e87 - aec6 - 15c1bee8e0a9
urn : lsid : biodiversity . org . au : afd . taxon : b1892325 - 13b1 - 4247 - a2e0 - 986a69278e48
urn : lsid : biodiversity . org . au : afd . name : 301255
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these spiders are restricted to the northern wheatbelt region in western australia . they are currently ( 2017 ) the only species listed as threatened under federal legislation ( epbc act ) . they are believed to be threatened by habitat destruction and fragmentation . a number of spider species have a rugose abdomen like this and are subject to a taxonomic study by michael rix ( queensland museum ) and collaborators .
n . b . : considered a senior synonym of aganippe o . pickard - cambridge , 1877 ( type a . subtristis o . pickard - cambridge , 1877 ) [ urn : lsid : nmbe . ch : spidergen : 00072 ] and of anidiops ( type a . manstridgei pocock , 1897 ) [ urn : lsid : nmbe . ch : spidergen : 00073 ] by rix et al . , 2017 : 590 ; transferred from the ctenizidae to the idiopidae by raven , 1985a : 138 .
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050288 ]
| | australia ( new south wales ) [ urn : lsid : nmbe . ch : spidersp : 000630 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 000631 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050289 ]
| | australia ( south australia ) [ urn : lsid : nmbe . ch : spidersp : 050290 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 000632 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050291 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050292 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 049619 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050293 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050294 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050295 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050296 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050297 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050298 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050299 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 000644 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050300 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050301 ]
| | australia ( new south wales ) [ urn : lsid : nmbe . ch : spidersp : 000634 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 000806 ]
| | australia ( western australia , south australia ) [ urn : lsid : nmbe . ch : spidersp : 000635 ]
| | australia ( new south wales ) [ urn : lsid : nmbe . ch : spidersp : 000637 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 000638 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 050302 ]
| | australia ( western australia ) [ urn : lsid : nmbe . ch : spidersp : 000807 ]
| | australia ( south australia ) [ urn : lsid : nmbe . ch : spidersp : 000641 ]
| | australia ( south australia ) [ urn : lsid : nmbe . ch : spidersp : 000642 ]
| | australia ( victoria ) [ urn : lsid : nmbe . ch : spidersp : 000643 ]
these spiders , from the sub - order orthognatha ( mygalomorphae ) or primitive spiders , are famous for their falsely suggested
deadly
bites . more about the exaggerated poisonous of these and other spiders can be read here . the bird - eating spider and tarantula , which is a common name for these spiders , the goliath spider ( theraposa blondi ) which is the largest spider on the world with a leg span of 30 cm and a weight of 130 grams , sydney funnel web spider ( atrax robustus ) and the mouse spider ( missulena ) are names given to species belonging to the order . in europe only two members of this sub - order can be found . in australia 13 % ( > 240 ) of the spiders belong to the mygalomorphae . the ancestral lineage of these spiders goes back over 360 million years .
most of these spiders live fearful lives buried deep in holes . many species react on unexpected events by cowering in fear , unable to move , or by violently plunging their pickaxe fangs . ( more about the fangs ) the spider can often be spotted when the males start searching for females and leave their burrows . females are more difficult to find because they can live for several years in their burrow with out leaving it .
the two long spinnerets at the back end of their abdomen and their large fangs that move up and down instead of sideways , like the modern spiders , are characteristic for this order .
distribution the eastern mouse spider ( missulena bradleyi ) lives in eastern australia from queensland to victoria . the redheaded mouse spider ( missulena occatoria ) occurs across most of the mainland , except southern victoria and northern australia . the male of this species has a bright red cephalothorax . the northern mouse spider ( missulena pruinosa ) is found in northern australia around darwin . one species has been described outside australia in chile .
missulena bradleyi rainbow , 1914 ; new south wales missulena dipsaca faulder , 1995 ; australia missulena granulosa o . p . - cambridge , 1869 ; western australia missulena hoggi womersley , 1943 ; western australia missulena insignis o . p . - cambridge , 1877 ; australia missulena occatoria walckenaer , 1805 ; southern australia missulena pruinosa levitt - gregg , 1966 ; western australia , northern territory missulena reflexa rainbow & pulleine , 1918 ; south australia missulena rutraspina faulder , 1995 ; western australia , south australia , victoria missulena torbayensis main , 1996 ; western australia missulena tussulena goloboff , 1994 ; chile
there are over 150 species in the family barychelidae in australia . males are called silverbacks because they are silvery coloured on the head . females have dark to golden brown hairs on their heads .
idiomata are funnel web spiders that build their burrows with a door . not all genera in this family have a door to close their burrow . the flask - like chambers are just below the ground .
they are mostly terrestrial spiders , which build typical silk - lined tubular burrow retreats , with a collapsed
tunnel
or open
funnel
entrance from which irregular trip lines radiate out over the ground . exceptions , which lack trip lines but may have trapdoors , are those hadronyche from south australia , like
the silk entrance tube may be split into 2 openings , in a y or t form . in the case of
the burrow may be in the hollow of a tree trunk or limb , many meters above ground level .
adult male spiders leave the burrow permanently to seek a mate . such wandering male spiders may enter houses , sometimes even find their way into clothing , and thus account for many bites . most funnel - web spiders are ground or log dwellers but at least two are tree dwellers (
, the sydney funnelweb spider , has a distribution centering on sydney , extending north to the hunter river , south to shoalhaven river , and narrowing westwards as far as lithgow .
has a considerably wider distribution ; being the coastal areas and highland forest regions from tasmania to queensland .
the australian funnel - web spiders ( family hexathelidae , simon , 1892 ) are probably the most dangerous spiders we can encounter . the most famous spider is the sydney funnel web ( atrax robustus ) . chances to be bitten are small . there are only two cases of envenomation annually in the last 10 years . funnel - web spiders belong to the family hexathelidae and two ( atrax and hadronyche ) of the eleven genera are considered dangerous . of the 40 described species in this family , the six red printed species caused severe envenomation .
hadrochyne spiders are medium sized spiders with a size varying between 10 and 50 mm . the largest species , hadronyche formidabilis measures between 40 and 50 mm . the sparsely haired spiders are coloured between brown to black . not all species are dangerous but a bite from a funnel web spider should always be treated seriously .
the southern tree funnel - web spider , hadronyche cerberea , is common around sydney and in the central coast regions . they make their silk - lined retreats in rough - barked trees often covered with bark and other wood particles .
this family contains six subfamilies with 38 genera and 329 species , widely distributed worldwide . in australia 86 species in at least six genera are described .
these spiders live in silk - lined burrows up to 20 cm deep . some species close the burrow with a lid .
this is a large family of trap door spiders with about 270 species , mostly found in the southern hemisphere , worldwide . eight genera with about 70 species live in australia . all australian genera are endemic with the exception of misgolas that also occurs in new zealand . almost all members of this family live in arid areas where they live in burrows up to 60 cm deep .
misgolas spiders are 15 - 30 mm in length and commonly found in eastern australia . the spider can of course bite but the toxin is not dangerous . the penetration of the fangs throught the skin may hurt .
this is a large family with 111 genera and 883 species world wide . their body size varies from 13 to 90 mm . the largest spider theraphosa blondi also belongs to this family . spiders of this family are called tarantulas or bird - eating spiders . this is a very common name but it often refers to these spiders .
in australia inly six species occur ; selenocosmia crassipes , selenocosmia stirlingi , selenocosmia strenua , selenocosmia subvulpina , selenotholus foelschei , selenotypus plumipes .
a famous member is the barking spider or whistling spider selenocosmia crassipes . she makes a rasping sound with her mouthparts that can be heard from a one meter distance . spiders from this family make burrows that can reach a length of two meters when the spider is mature . young spiders can be found under rocks and roots . the spider lives in north eastern australia and papua new guinea . the spider is quite large with a body length of 70 to 90 mm . measured from the tip on her legs she can be 200 mm in length .
funnel web tarantulas occur almost worldwide in the tropics . the 175 world wide and 90 australian members of this family often build messy funnel webs . most species are uncommon and live in remote areas . most species are small hairy and dark brown to black . sometimes they can be found in holes in trees . their spinnerets are moderate long .
cethegus ischnotheloides has a body length of 15 mm . this spider lived in a dense web on the ground between shrub in leinster , western australia .
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