kurixalus idiootocus is a small species of frog in the rhacophoridae family .

it is endemic to taiwan and is commonly known as the temple tree frog .

its natural habitats are subtropical or tropical moist shrubland , seasonally wet or flooded lowland grassland , freshwater marshes , intermittent freshwater marshes and irrigated land .

it is listed as being of " least concern " by the iucn although there may be some destruction of its habitat .

kurixalus idiootocus

advertisement calls of four kurixalus species from taiwan . a . kurixalus berylliniris sp . n . “slow call” b . kurixalus berylliniris sp . n . “rapid call” c . kurixalus wangi sp . n . “slow call” d kurixalus wangi sp . n . “rapid call” e kurixalus eiffingeri f kurixalus idiootocus .
four kurixalus species of taiwan . a kurixalus berylliniris sp . n . ( holotype , adult , dark morph ) b kurixalus wangi sp . n . ( holotype ) c kurixalus berylliniris sp . n . ( sub - adult ) d kurixalus berylliniris sp . n . ( adult , light morph ) e kurixalus eiffingeri f kurixalus idiootocus .
kurixalus berylliniris sp . n . , kurixalus wangi sp . n . , oophagous tadpoles
lateral view of tadpoles of three oophagus kurixalus species . a kurixalus berylliniris sp . n . b kurixalus wangi sp . n . c kurixalus eiffingeri . scale bars 1 mm .
comparisons of the characteristics of slow mating calls among kurixalus eiffingeri , kurixalus berylliniris sp . n . and kurixalus wangi sp . n .
dorsal view of tadpole head region of three oophagus kurixalus species . a kurixalus berylliniris sp . n . b kurixalus wangi sp . n . c kurixalus eiffingeri . scale bars 1 mm .
temple tree frog ( kurixalus idiootocus ) temple tree frog stock photo , picture and royalty free image . image 56473078 .
chirixalus idiootocus — bossuyt and dubois , 2001 , zeylanica , 6 : 57 .
the guts of tadpoles of the two new species contained a yellow ‘yolky’ substance . when the same characteristic was observed in kurixalus eiffingeri it was confirmed as tadpole oophagy ( ueda 1986 , liang et al . 2002 ) . therefore , it is likely that the tadpole oophagy is a synapomorphy for the two new species and kurixalus eiffingeri . interestingly , kurixalus idiootocus , as well as all known kurixalus species from mainland china and southern asia lack this particular reproductive behavior . therefore , the oophagy reproductive behavior could also support the phylogenetic positions of kurixalus eiffingeri , kurixalus berylliniris sp . n . , and kurixalus wangi sp . n . within the kurixalus genus .
kurixalus eiffingeri , a native species in the island of taiwan , is the only rhacophorid within the genus kurixalus that has a tree - hole breeding reproductive mode and oophagous tadpoles ( ueda 1986 , lehtinen and nussbaum 2003 , wells 2007 ) . kurixalus idiootocus , a species endemic to taiwan , has a lentic feeding tadpole type , which is similar to most species in the genus kurixalus ( inger 1966 , inger et al . 1999 , kuramoto and wang 1987 ) . in previous molecular phylogenetic studies , kurixalus eiffingeri and kurixalus idiootocus have been recovered as sister taxa ( abraham et al . 2013 , yu et al 2013 , nguyen et al . 2014a , b ) . since kurixalus eiffingeri and kurixalus idiootocus are the only two species that have been described from the island of taiwan , rhacophorid frogs with similar life history to kurixalus idiootocus ( but see abraham et al 2013 ) , specifically any rhacophorid frogs with tree - hole breeding reproductive mode or lentic feeding tadpole type would be would be assigned to either of the two species .
nesting sites of three tree - hole breeding kurixalus species ( a nest is made by the animal ) . a eggs of kurixalus berylliniris sp . n . b eggs of kurixalus wangi sp . n . ; note that the parents were present with eggs c eggs of kurixalus eiffingeri .
molecular evidence for taxonomy of rhacophorus appendicularis and kurixalus species from northern vietnam , with comments on systematics of kurixalus and gracixalus ( anura : rhacophoridae ) .
analysis of covariance of morphometric characteristics of males and females among four kurixalus species .
aquixalus idiootocus — fei , hu , ye , and huang , 2009 , fauna sinica , amph . 2 : 671 .
effects of intermittent feeding on the growth of oophagous ( chirixalus eiffingeri ) and herbivorous ( chirixalus idiootocus ) tadpoles from taiwan .
measurements ( in mm ) of type series and other specimens of kurixalus berylliniris sp . n . and kurixalus wangi sp . n . abbreviations as in materials and methods .
genetic variation of three kurixalus species from taiwan according to mtdna co1 gene partial sequence .
a new tree frog of the genus kurixalus ( anura : rhacophoridae ) from vietnam .
molecular evidence for taxonomy of rhacophorus appendiculatus and kurixalus species from northern vi . . .
we follow wilkinson et al . ( 2002 ) in transferring this species from chirixalus to kurixalus .
fst ( above diagonal ) and nm ( below diagonal ) between three kurixalus species from taiwan .
a new cryptic tree frog species allied to kurixalus banaensis ( anura : rhacophoridae ) from vietnam .
annandale ' s high altitude frog ( kurixalus naso ) is a rhacophorid frog that . . .
in the latest study , a team led by zoologist yu - teh kirk lin at the national taiwan university in taipei studied kurixalus idiootocus tree frogs in a wooded suburb of taipei during the mating season , which lasts from february to september .
sampling localities of this study . localities 1 through 22 are around taiwan island , locality 23 from iriomote isle , locality 24 from ishigaki isle . the two isles belong to the southern end of ryukyu archipelago . color refers to the geographical distribution of the three kurixalus species . red : kurixalus eiffingeri ; green : kurixalus berylliniris sp . n . ( taxon 1 ) ; b : kurixalus wangi sp . n . ( taxon 2 ) . loc . 20 : ligia , type locality of kurixalus berylliniris sp . n . ; loc . 21 : shouka , type locality of kurixalus wangi sp . n .
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kurixalus naso ( annandale , 1912 )
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kurixalus naso ( annandale , 1912 )
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a species boundary within the chinese kurixalus odontotarsus species group ( anura : rhacophoridae ) : n . . .
aquixalus ( aquixalus ) idiootocus — delorme , dubois , grosjean , and ohler , 2005 , bull . mens . soc . linn . lyon , 74 : 166 .
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kurixalus naso
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the epithet is named and dedicated to mr . ching - shong wang for his pioneering work and contributions to the herpetology of taiwan ( wang 1962 ) . mr . wang discovered two rhacophorid frogs ( rhacophorus taipeianus and kurixalus idiootocus ) ( liang and wang 1963 , kuramoto and wang 1987 ) in taiwan and suggested , in the early 1980s , that some kurixalus specimens collected near the type locality of this new species might be different from kurixalus eiffingeri ( personal communication ) . the name is used in the genitive case .
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kurixalus naso ( annandale , 1912 )
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max : maximum frequency ; min : minimum frequency ; wid : width of frequency ( max - min ) ; dur : single note duration ; int : time interval between notes ; df : dominant frequency ; all data shown are mean and standard deviation ( in parentheses ) based on 30 advertisement calls recorded in the field under natural conditions . environmental parameters : 1 ) kurixalus berylliniris sp . n . : 18 °c , 93 % rh . 2 ) kurixalus wangi sp . n . : 25 °c , 84 % rh . 3 ) kurixalus eiffingeri : 19 °c , 91 % rh . 4 ) kurixalus idiootocus : 24 °c , 80 % rh .
phylogenetic relationships : k . idiootocus is closely related to k . eiffingeri . and has gone through several taxonomic changes . kuramoto and wang ( 1987 ) suggested these species might be related to the genus buergeria .
pca morphometric comparisons of four kurixalus species from taiwan . scatterplots of ( a ) principal components 1 and 2 , and ( b ) principal components 2 and 3 of size - adjusted morphometric data for male frogs of the four kurixalus species . the 95 % confidence ellipses for each population ( elm ) are shown .
two new species of rhacophorid tree frog were identified in taiwan . in both new taxa , derived reproductive characteristics of laying eggs in tree holes and oophagous tadpoles are shared with kurixalus eiffingeri , but they are divergent from each other in molecular genetics , mating calls , and tadpole and adult morphology . the morphological characteristics and the molecular phylogenetic evidence support the hypothesis that the two new species , kurixalus berylliniris sp . n . and kurixalus wangi sp . n . , are both monophyletic lineages .
abbreviations : k : kurixalus ; f : feihyla ; r : rhacophorus ; jp : ryukyu islands of japan ; ntw : northern taiwan ; ctw : central taiwan .
dana campbell marked the classification from
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. . . in this study , we measured the body weight , body length , jump height , jump length , and climbing abilities of three species of frogs indigenous to taiwan . the results were used to design an amphibian corridor suitable for amphibian mobility . twenty specimens for each of the three species , rana adenopleura , rana latouchii , and kurixalus idiootocus , were collected for testing . their climbing ability . . .
kurixalus idiootocus — wilkinson , drewes , and tatum , 2002 , mol . phylogenet . evol . , 24 : 272 ; frost , grant , faivovich , bain , haas , haddad , de sá , channing , wilkinson , donnellan , raxworthy , campbell , blotto , moler , drewes , nussbaum , lynch , green , and wheeler , 2006 , bull . am . mus . nat . hist . , 297 : 245 .
body size variation ( svl ) in female and male kurixalus species from taiwan ( mean ± standard deviation ) . ( * * : significant level < 0 . 01 ) .
factor loadings of the first three principal components of 18 size - adjusted morphometric characteristics of males of four kurixalus species . absolute values of loadings greater than 0 . 50 in boldface .
holotype of kurixalus berylliniris sp . n . dorsal ( a ) , ventral ( b ) , and ventral view of hand ( c ) and foot ( d ) . scale bars : 10 mm .
phylogeny of rhacophoridae is constructed using two mitochondrial ( 12s rrna and 16s rrna ) and two nuclear ( tyrosinase and rag - 1 ) genes in an attempt to test for the taxonomic status of rhacophorus appendiculatus and kurixalus species from tam dao . all phylogenetic analyses demonstrate that specimens from tam dao are nested in kurixalus bisacculus , indicating that they belong to k . bisacculus . . . . [ show full abstract ]
the notion of “frog song” needs more careful consideration , we have found a similar vocalizing behavior of the human and male frogs . many people sing in the bathroom because the acoustics make them feel like the voice is stronger and richer . male mientien tree frogs kurixalus idiootocus frequently perch and call in roadside concrete drainages – miniature urban canyons . in an indoor experiment using a replica of a concrete drain , males preferred one particular type of call perch that facilitated call transmission ( tan , tsai , lin , & lin , 2014 ) .
chirixalus idiootocus kuramoto and wang , 1987 , copeia , 1987 : 932 . holotype : ntuma 1010 , by original designation . type locality :
in a rain pool near sanshengkong ( a temple , altitude ca . 500 m ) in the southern slope of mount mientien - shan , taipei , taiwan
, china .
comparisons of measurements and ratios of tadpoles of three oophagus kurixalus species from taiwan ( anova significant level , * : 0 . 01 < p < 0 . 05 ; * * : p < 0 . 01 ) .
fei , 1999 , atlas amph . china : 252 - 253 , provided a brief account ( as philautus idootocus ) , figure , and map . bossuyt and dubois , 2001 , zeylanica , 6 : 57 - 58 , discussed with this species cannot be in philautus and , although they transferred it back to chirixalus , noted that the definition of chirixalus was so poorly resolved that this species might ultimately have to be moved to a distinct genus , a position formalized by wilkinson , drewes , and tatum , 2002 , mol . phylogenet . evol . , 24 : 272 . lue , tu , and hsiang , 1999 , atlas taiwan amph . rept . : 40 - 41 , provided a brief account for taiwan ( as chirixalus idiootocus ) . fei , hu , ye , and huang , 2009 , fauna sinica , amph . 2 : 671 - 676 , provided an account ( as aquixalus idiootocus ) and spot map . fei , ye , and jiang , 2010 , colored atlas of chinese amph . : 428 - 429 , provided a brief account ( as aquixalus idiootocus ) for china including photographs of specimens and habitat . fei , ye , and jiang , 2012 , colored atlas chinese amph . distr . : 498–499 , provided an account ( as aquixalus idiootocus ) , photographs , and a range map for china .
genetic distances among kurixalus species and two outgroup taxa in co 1 gene ( above diagonal ) and 16s rrna gene ( below diagonal ) . numbers on top row refer to species shown on the left column . genetic distances are shown as percentage .
buergeria japonica : nchuzool : 4021 , 4825 , 4826 ( taiwan ) . buergeria robusta : nchuzool 4025 , 4027 , 4831 ( taiwan ) . kurixalus eiffingeri : ntu 927–28 , 931 - 32 , 939 , 1052 , 1054–56 , 1649 ( shitou , nantou ) ; 1058 - 67 , 1645–46 , 1769 ( wulai , taipei ) ; nchuzool : 2502–03 , 2508 , 2509 ( c / s ) , 2514–15 , 2517–19 , 2523–24 , 2525 ( c / s ) , 2526–27 , 2535 , 2536 ( c / s ) , 2537 , 2538 ( c / s ) , 2539–40 , 2545–47 , 2550–51 , 2829 , 2831 , 4018 ( c / s ) , 11320 , 11436–40 ( shindian , taipei ) ; 11333 ( shitou , nantou ) . kurixalus idiootocus : ntu 929 , 1005–09 , 1033 , 1035–37 , 1038–42 ( shiding , taipei ) ; 1045 ( suao , yilan ) ; 1010 ( holotype of chirixalus idiootocus ) , 1011 , 1013–29 , 1657 , 1695–96 , ( yanminshan , taipei ) ; 1658 ( shindien , taipei ) ; 1708–09 ( juchi , chiayi ) ; 1770 ( wulai , taipei ) ; nchuzool 1010 , 2780 , 2782 , 2784 , 2785 ( c / s ) , 2787–88 , 2792 , 2795 , 2796 ( c / s ) , 2805 , 2845 , 2847 , 2954 , 2956 ( c / s ) , 2959–63 , 3709 , 3711 ( c / s ) , 4304 ( c / s ) , 4990 , 4993 , 4995 ( c / s ) , 7402 . kurixalus wangi sp . n . : ntu 1043–1044 , 1046 , 1647 - 48 ( manjo , pingtung county ) .
the population of kurixalus naso population is thought to be small , and susceptible to deforestation . little is known about this species , and the taxonomic identities and relationships within this genus are also poorly known and not distinguishable on the basis of morphology ( tao et al . 2014 ; nguyen et al . 2012 ) . kurixalus naso occurs in several preserves including dihang - dibang biosphere reserve and mouling national park in arunachal pradesh ( dutta et al . 2004 ) . more sampling and fieldwork is necessary to understand this species , its ecology , and its conservation status .
holotype of kurixalus wangi sp . n . dorsal ( a ) , ventral ( b ) , and ventral view of hand ( c ) and foot ( d ) . scale bars 10 mm in ( a ) and ( b ) ; 5 mm in ( c ) and ( d ) .
annandale ' s high altitude frog ( kurixalus naso ) is a rhacophorid frog that inhabits montane forests , shrublands and grasslands of northeastern india , the eastern xizang province of china , and myanmar . in india it is recorded from altitudes between 1 , 100 and 1 , 500m asl . kurixalus naso is a small , arboreal species , that breeds in temporary water bodies ( dutta et al . 2004 ) . annandale ( 1912 ) reported the type specimen as 43 mm ( 1 . 7 in ) snout to vent length ( svl ) , with a stout body , short stout limbs , and a broad head . it is purplish - brown on its dorsal surface , with grey mottling , and has an off - white belly ( annandale 1912 ) . a cone - shaped snout distinguishes this species from others in genus kurixalus . ( annandale 1912 ; tao et al . 2014 ) . very little literature describes this species .
phylogenetic relationship of all kurixalus species from taiwan . a phylogram showing the phylogenetic relationships of the four kurixalus species , obtained by a maximum likelihood search based on 1207 nucleotides from mtdna co1 and 16s rrna genes . feihyla palpebralis and rhacophorus moltrechti were used as outgroups . the three values on each branch are maximum likelihood ( ml ) , maximum parsimony ( mp ) , and neighbor - joining ( nj ) analyses with bootstrapping support based on 2000 replicates . bootstrapping values below 50 % are not shown . ( jp : ryukyu islands of japan ; n . tw : northern taiwan ; c . tw : central taiwan ) .
we would like to show our gratitude to hm huang for collecting the first specimen of kurixalus berylliniris sp . n . ; to jh chu , cc hwang , hj su and sm lin for advice on molecular analysis ; to h masaki for providing specimens of kurixalus eiffingeri from the ryukyu islands , to cy hsiao for photographs ; to yt chung for his dedicated fieldwork ; to cc wu for performing bench work ; and to cf lin for providing kurixalus eiffingeri tadpoles . thanks are extended to cj huang , ch chang , cw lin and yc liu for assistance with the fieldwork . we appreciate the help of sf chan with vocal analysis and yh chen for partial sample contribution . the map of taiwan was kindly prepared by pf lee . this study was partially supported by the council of agriculture grant ( 94–admin–4 . 1–conserv–03 ( 2 ) ) , by shei - pa national park , ministry of interior grant ( 094 - 301020500g1 - 005 ) , and a support from mr . cc chen . re brown greatly improved an early draft of the manuscript .
. . . phylogenetic data support the monophyletic origin of kurixalus despite its broad distri - bution ( li et al . , 2013 ; biju et al . , 2016 ; jiang et al . , 2016 ) . however , since these frogs are similar in morphology and overlap in body size , taxonomy is difficult and mainly based on molecular analyses ( wilkinson et al . , 2002 ; li et al . , 2008 ; li et al . , 2009 ; hertwig et al . , 2013 ; yu et al . , 2013 ; nguyen et al . , 2014a ; nguyen et al . , 2014b ; wu et al . , 2016 ) . currently , 14 species are recognized in kurixalus ( frost , 2017 ) . . . .
wu s - p , huang c - c , tsai c - l , lin t - e , jhang j - j , wu s - h ( 2016 ) systematic revision of the taiwanese genus kurixalus members with a description of two new endemic species ( anura , rhacophoridae ) . zookeys 557 : 121–153 . doi : 10 . 3897 / zookeys . 557 . 6131
we construct the phylogeny of the kurixalus odontotarsus species group using two mitochondrial ( 12s rrna and 16s rrna ) genes in an attempt to delimit species boundaries within the chinese k . odontotarsus group . with strong support values , three major clades are obtained , and all phylogenetic analyses reject monophyly of k . odontotarsus . the tibetan lineage of k . odontotarsus was clustered with . . . [ show full abstract ]
kurixalus eiffingeri is not an endangered or protected species and thus no specific permission is needed for field studies . some bamboo forests in chitou are privately owned , and we had verbal permission from land owners ( mr . pan yeong - sung and ms chen mei - chi ) to conduct observational and field studies ( 23 o 41’05 . 4” n 130 o 47’45 . 4” e and 23 o 41’22 . 8” n 130 o 47’22 . 5” e , respectively ) .
in this study , we used a taiwanese frog ( kurixalus eiffingeri ( anura : rhacophoridae ) ) that breeds in water - filled bamboo stumps as a model animal to study the parentage between overlapping offspring and its ecological consequence on reproductive strategy and nest site selection . specifically , we used ( 1 ) five microsatellite dna markers to analyze the parentage of adults and tadpoles and ( 2 ) paired bamboo cups with and without tadpoles to study the nest choice of frogs and to reveal the possible causes of nest reuse .
a small to moderate - sized kurixalus . females snout - vent length averaging about 34 mm ( range : 30 . 8–37 . 1 mm ) ; males averaging 30 mm ( range : 28 . 6–31 . 6 mm ) . iris golden - yellow . two anterior horns of the x - shaped marking on back extending to eyelid . webbing extensive on toes , extending to the toe disc on the inner margin of toe v . belly and throat whitish . anterior margin of tadpole dorsal fin extending to posterior body . tadpole with almost no pigment on region of tail muscle . upper lip of tadpole with shallow transverse furrow .
description diagnosis : kurixalus idiooticus is a small treefrog similar to k . eiffingeri but with a smaller body size . it has a smaller and weakly developed pollex , has less rugose skin . no female anal flap is present , dorsal warts lack white spinules , its ground color is never greenish , and generally a large dark brown hourglass - like pattern on the back . males have an aggressive vocal behavior . the species has a generalized larval morphology . in contrast to all other known taiwanese rhacophorine frogs , k . idiooticus lays eggs terrestrially near the water ' s edge without constructing a foam nest ( kuramoto and wang 1987 ) .
. . . recently , dubois ( 2005 ) assigned both theloderma and nyctixalus to the tribe philautini ( including also aquixalus , kurixalus , and philautus ) , whereas more recently grosjean et al . ( 2008 ) proposed separation of theloderma and nyctixalus in the new tribe nyctixalini . this assignment and distant phylogenetic position of theloderma and nyctixalus in respect to other members of the subfamily rhacophorinae was also supported by subsequent phylogenetic analyses ( li et al . , 2008 li et al . , , 2009li et al . , , 2013 pyron and wiens , 2011 ) . based on the analysis of mtdna genetic markers , li et al . ( 2008 ) showed sister clade relationships between nyctixalus ( n . . . .
a moderate - sized kurixalus . females average about 41 mm snout - vent length ( range : 27 . 6–46 . 3 mm ) ; males average about 35 mm ( range : 29 . 0–42 . 3 mm ) . iris emerald to light green . two dark brown spots on eyelids , separated from each other and from x - shaped blotch on dorsum . subarticular tubercles on foot rounded and flat . belly and throat white or faintly - speckled . prepollex in males squarish , compressed and expanded . about half - webbed between two outer toes . anterior margin of tadpole dorsal fin extending to body . tadpole heavily dark brown to black pigmented in gular region and on tail muscle . upper lip of tadpole with deep transverse furrow , and prominent ridge extending from upper lip to anterior margin of nostril ( key of tadpole , 3 ) .
. . . three mitochondrial genes and five nuclear genes were surveyed : partial 12s rrna , trna - val , and 16s rrna genes were regarded as a single mitochondrial locus ; the other five nuclear loci included ( i ) a fragment of the brain - derived neurotrophic factor ( bdnf ) ; ( ii ) a fragment of proopiomelanocortin ( pomc ) ; ( iii ) a fragment of the recombination activating gene 1 ( rag - 1 ) ; ( iv ) a fragment of exon 1 of rhodopsin ( rhod ) , and ( v ) a fragment of exon 1 of tyrosinase ( tyr ) . mitochondrial 12s rrna , trna - val , and 16s rrna sequences of kurixalus eiffingeri from ishigaki island ( ryukyu islands ) ( see the last one in supplement table 1 for detailed information ) were newly acquired according to protocols of previous studies ( li et al . , 2008 ; li et al . , 2009 ) . sequences from mitochondrial dna and five nuclear loci were aligned with muscle 3 . 8 . 31 . . .
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frost , d . r . 2014 . amphibian species of the world : an online reference . version 6 ( 27 january 2014 ) . new york , usa . available at : urltoken . ( accessed : 27 january 2014 ) .
justification : listed as least concern in view of its relatively wide distribution , tolerance of a broad range of habitats , presumed large population , and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category .
this species is endemic to taiwan , province of china , and occurs below 2 , 000m asl .
it inhabits shrubland , grassland , paddy fields and nearby pools . it usually breeds in shallow , still waterbodies .
habitat destruction and degradation are potential threats to this species , in particular infrastructure development for human settlement .
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tadpoles are pond - type ( rounded body , small mouth and sinistral spiracle with moderate tail ) . at stages 25 - 27 , body length is 1 . 4 - 1 . 5x longer than wide . body is oval and moderately depressed . eyes and nostrils are dorsolateral . dorsal fin is slightly larger than ventral fin . caudal musculature is slender and tapers gradually to the tip of the tail . papillae are marginal , small , and homogeneous in size ; they form a continuous row across the ventral lip and extend to the sides of the lips . ltrf is 2 : 1 + 1 / 3 at stages earlier than 37 and 2 : 3 + 3 / 3 or 1 : 4 + 4 / 3 at stage 37 . head and body are generally dark brown , sometimes with black spotting . caudal muscle is dark brown . fins are transparent at stage 25 . ( kuramoto and wang 1987 ) .
coloration in life : ground color of light grayish brown , brown , dark brown or yellowish brown ( never greenish ) . the dorsum is characterized by a dark brown stripe . dark brown spots and patches are on the sides of head . a brown bar extends from the nostril to edge of upper lip , and another brown bar extends from the anterior lower edge of the eye to the edge of the upper lip . the upper lip has a few narrow dark bars . an inverted y shape in brown or black extends from a dark cross band between upper eyelids to posterior part of the back . this marking can be y - shaped , h - shaped or x - shaped , or may be lacking . sides of the body have dark brown marbling with some dark spots . anterior throat has dark brown marbling and spotting , as well as some white granules . the belly is dark brown and marbled , scattered with dark spots . forearms have a dark brown cross band . thighs have numerous small dark patches on both the anterior and posterior sides . a dark knee patch is present . the sides of the anal opening are dark brown ; white granules are present on the dorsal and ventral sides of the anal opening ( kuramoto and wang 1987 ) .
is widespread in taiwan ( e . g . , pinglin , mientien - shan , chutung , suao , chushan , chiahsien , anping ) at altitudes from 10 - 750 m asl ( kuramoto and wang 1987 ) . it generally occurs at a lower elevation than the closely related species
; so far the two species are known to overlap only at anping ( kuramoto and wang 1987 ) . habitat includes grassland , paddy fields and shrublands or anywhere near still , shallow water bodies ( stuart et al . 2008 ) .
the breeding season lasts from march - june with males calling from the ground or in low shrubs , often near rain pools ( 5 - 20 cm in depth ) that are surrounded by high vegetation ( grasses or bushes ) . males form large breeding aggregations and calling males confront one another , with one male ceasing to call . calls resemble birds warbling and generally consist of a long series ( 2 - 6 seconds ) of fast , repeating pulses , but other call types include short trills with sharp pulses and short calls with a long pulse . males are aggressive and calling males approach each other , with one male then ceasing to call ( kuramoto and wang 1987 ) .
lays pigmented eggs ( gray - brown animal half , light yellowish gray vegetal half ) with transparent gelatinous coats . ova measured about 1 . 95 mm in diameter and were covered with two jelly layers , with the diameter of the outermost , tough , non - sticky jelly layer about 4 . 09 mm . eggs are laid terrestrially near the water ' s edge or in depressions , with hatching triggered by rainfall , filling the depressions or washing the tadpoles into adjacent ponds and ditches . some egg masses were observed more than 50 cm in horizontal distance from the water . egg masses were often concealed under fallen leaves or other debris , in soil depressions or in crevices behind stones or openings of rat holes . the clutch size averaged 182 eggs . tadpoles of
populations are stable with the major threat being habitat destruction and degradation from human settlements . its range overlaps with many protected areas in taiwan ( stuart et al . 2008 ) .
karyotype : 2n = 26 with eight small pairs and five large pairs of chromosomes ( kuramoto and wang 1987 ) .
is derived from the greek words “idios , ” or peculiar , and
ootocos ,
or egg - laying ( kuramoto and wang 1987 ) .
kuramoto , m . , and wang , c . s . ( 1987 ) . ' ' a new rhacophorid treefrog from taiwan , with comparisons to
stuart , s . , hoffmann , m . , chanson , j . , cox , n . , berridge , r . , ramani , p . , and young , b . ( eds ) ( 2008 ) .
lynx edicions , iucn , and conservation international , barcelona , spain ; gland , switzerland ; and arlington , virginia , usa .
> university of california , berkeley , ca , usa . accessed jul 9 , 2018 .
> university of california , berkeley , ca , usa . accessed 9 jul 2018 .
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lithobates pipiens
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temple treefrog ( frank and ramus , 1995 , compl . guide scient . common names amph . rept . world : 111 ; lue , tu , and hsiang , 1999 , atlas taiwan amph . rept . : 40 ) .
mientien small treefrog ( fei , 1999 , atlas amph . china : 252 ) .
mientien treefrog ( lue , tu , and hsiang , 1999 , atlas taiwan amph . rept . : 40 ) .
please note : these links will take you to external websites not affiliated with the american museum of natural history . we are not responsible for their content .
for access to available specimen data for this species , from over 350 scientific collections , go to vertnet .
copyright © 1998 - 2018 , darrel frost and the american museum of natural history . all rights reserved .
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< p > when browsing through different uniprot proteins , you can use the ‘basket’ to save them , so that you can back to find or analyse them later . < p > < a href = ' / help / basket ' target = ' _ top ' > more . . . < / a > < / p >
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< p > the < a href =
urltoken
> gene ontology ( go ) < / a > project provides a set of hierarchical controlled vocabulary split into 3 categories : < p > < a href = ' / help / gene _ ontology ' target = ' _ top ' > more . . . < / a > < / p >
< p > uniprotkb keywords constitute a < a href =
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> controlled vocabulary < / a > with a hierarchical structure . keywords summarise the content of a uniprotkb entry and facilitate the search for proteins of interest . < p > < a href = ' / help / keywords ' target = ' _ top ' > more . . . < / a > < / p >
< p > information which has been generated by the uniprotkb automatic annotation system , without manual validation . < / p > < p > < a href =
/ manual / evidences # eco : 0000256
> more . . . < / a > < / p >
< p > this section provides information about the protein and gene name ( s ) and synonym ( s ) and about the organism that is the source of the protein sequence . < p > < a href = ' / help / names _ and _ taxonomy _ section ' target = ' _ top ' > more . . . < / a > < / p >
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urltoken
> names and taxonomy < / a > section provides an exhaustive list of all names of the protein , from commonly used to obsolete , to allow unambiguous identification of a protein . < p > < a href = ' / help / protein _ names ' target = ' _ top ' > more . . . < / a > < / p >
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/ manual / evidences # eco : 0000313
> more . . . < / a > < / p >
< p > this subsection of the < a href =
urltoken
> names and taxonomy < / a > section provides information on the name ( s ) of the organism that is the source of the protein sequence . < p > < a href = ' / help / organism - name ' target = ' _ top ' > more . . . < / a > < / p >
< p > this subsection of the < a href =
urltoken
> names and taxonomy < / a > section shows the unique identifier assigned by the ncbi to the source organism of the protein . this is known as the ‘taxonomic identifier’ or ‘taxid’ . < p > < a href = ' / help / taxonomic _ identifier ' target = ' _ top ' > more . . . < / a > < / p >
< p > this subsection of the < a href =
urltoken
> names and taxonomy < / a > section contains the taxonomic hierarchical classification lineage of the source organism . it lists the nodes as they appear top - down in the taxonomic tree , with the more general grouping listed first . < p > < a href = ' / help / taxonomic _ lineage ' target = ' _ top ' > more . . . < / a > < / p >
< p > this section provides information on sequence similarities with other proteins and the domain ( s ) present in a protein . < p > < a href = ' / help / family _ and _ domains _ section ' target = ' _ top ' > more . . . < / a > < / p >
< p > this subsection of the < a href =
urltoken
> family and domains < / a > section describes the position and type of a domain , which is defined as a specific combination of secondary structures organized into a characteristic three - dimensional structure or fold . < p > < a href = ' / help / domain ' target = ' _ top ' > more . . . < / a > < / p >
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/ manual / evidences # eco : 0000259
> more . . . < / a > < / p >
< p > this section displays by default the canonical protein sequence and upon request all isoforms described in the entry . it also includes information pertinent to the sequence ( s ) , including < a href =
urltoken
> length < / a > and < a href =
urltoken
> molecular weight < / a > . < p > < a href = ' / help / sequences _ section ' target = ' _ top ' > more . . . < / a > < / p >
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urltoken
> sequence < / a > section indicates if the < a href =
urltoken
> canonical sequence < / a > displayed by default in the entry is complete or not . < p > < a href = ' / help / sequence _ status ' target = ' _ top ' > more . . . < / a > < / p >
< p > the checksum is a form of redundancy check that is calculated from the sequence . it is useful for tracking sequence updates . < / p > < p > it should be noted that while , in theory , two different sequences could have the same checksum value , the likelihood that this would happen is extremely low . < / p > < p > however uniprotkb may contain entries with identical sequences in case of multiple genes ( paralogs ) . < / p > < p > the checksum is computed as the sequence 64 - bit cyclic redundancy check value ( crc64 ) using the generator polynomial : x < sup > 64 < / sup > + x < sup > 4 < / sup > + x < sup > 3 < / sup > + x + 1 . the algorithm is described in the iso 3309 standard . < / p > < p class =
publication
> press w . h . , flannery b . p . , teukolsky s . a . and vetterling w . t . < br / > < strong > cyclic redundancy and other checksums < / strong > < br / > < a href =
urltoken
> numerical recipes in c 2nd ed . , pp896 - 902 , cambridge university press ( 1993 ) < / a > ) < / p >
< p > this subsection of the ‘sequence’ section is used for sequence fragments to indicate that the residue at the extremity of the sequence is not the actual terminal residue in the complete protein sequence . < p > < a href = ' / help / non _ ter ' target = ' _ top ' > more . . . < / a > < / p >
< p > this section provides links to proteins that are similar to the protein sequence ( s ) described in this entry at different levels of sequence identity thresholds ( 100 % , 90 % and 50 % ) based on their membership in uniprot reference clusters ( < a href =
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> uniref < / a > ) . < p > < a href = ' / help / similar _ proteins _ section ' target = ' _ top ' > more . . . < / a > < / p >
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< p > this subsection of the ‘entry information’ section provides a mnemonic identifier for a uniprotkb entry , but it is not a stable identifier . each reviewed entry is assigned a unique entry name upon integration into uniprotkb / swiss - prot . < p > < a href = ' / help / entry _ name ' target = ' _ top ' > more . . . < / a > < / p >
< p > this subsection of the ‘entry information’ section provides one or more accession number ( s ) . these are stable identifiers and should be used to cite uniprotkb entries . upon integration into uniprotkb , each entry is assigned a unique accession number , which is called ‘primary ( citable ) accession number’ . < p > < a href = ' / help / accession _ numbers ' target = ' _ top ' > more . . . < / a > < / p >
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urltoken
> canonical sequence < / a > are also displayed . < p > < a href = ' / help / entry _ history ' target = ' _ top ' > more . . . < / a > < / p >
< p > this subsection of the ‘entry information’ section indicates whether the entry has been manually annotated and reviewed by uniprotkb curators or not , in other words , if the entry belongs to the swiss - prot section of uniprotkb ( < strong > reviewed < / strong > ) or to the computer - annotated trembl section ( < strong > unreviewed < / strong > ) . < p > < a href = ' / help / entry _ status ' target = ' _ top ' > more . . . < / a > < / p >
< p > this will take you to the blast page where you can edit options < / p > < p > < a href =
/ help / sequence - searches
> more . . < / a > < / p >
we ' d like to inform you that we have updated our privacy notice to comply with europe’s new general data protection regulation ( gdpr ) that applies since 25 may 2018 .
< p > this section provides information on the location and the topology of the mature protein in the cell . < p > < a href = ' / help / subcellular _ location _ section ' target = ' _ top ' > more . . . < / a > < / p >
< p > this section provides information on the quaternary structure of a protein and on interaction ( s ) with other proteins or protein complexes . < p > < a href = ' / help / interaction _ section ' target = ' _ top ' > more . . . < / a > < / p >
< p > this subsection of the < a href =
urltoken
> ' interaction ' < / a > section provides information about the protein quaternary structure and interaction ( s ) with other proteins or protein complexes ( with the exception of physiological receptor - ligand interactions which are annotated in the < a href =
urltoken
> ' function ' < / a > section ) . < p > < a href = ' / help / subunit _ structure ' target = ' _ top ' > more . . . < / a > < / p >
the nucleosome is a histone octamer containing two molecules each of h2a , h2b , h3 and h4 assembled in one h3 - h4 heterotetramer and two h2a - h2b heterodimers . the octamer wraps approximately 147 bp of dna .
< p > this subsection of the ‘family and domains’ section provides information about the sequence similarity with other proteins . < p > < a href = ' / help / sequence _ similarities ' target = ' _ top ' > more . . . < / a > < / p >
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from 蛙蛙哇 ! songs of the frogs of taiwan vol . 1 , released january 6 , 2017
sound artist and field recordist based in taiwan . sound designer for contemporary dance and film . audio documentaries
producer and electroacoustic music composer . works inspired by ethnography , amphibian conservation and marine biology . collaborates with communities and individuals in hakka and austronesian territories .
if you like 蛙蛙哇 ! songs of the frogs of taiwan vol . 1 , you may also like :
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this is brutal . i just discovered leyland kirby , and i think he is a genius . his work does not hide behind empty , pretending stories , but seems to be heavily connected to the mind and its processes indeed . memories come to life and long lost feelings emerge - even feelings that never existed but yet feel familiar . words won ' t be able to capture what happens on your inside when you decide to let yourself be carried away by this sonical wizardry . haunting , stuck as a loop in my head .
supported by 4 fans who also own “蛙蛙哇 ! songs of the frogs of taiwan vol . 1”
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search 123rf with an image instead of text . try dragging an image to the search box .
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1 department of earth and life science , university of taipei . no . 1 , ai - guo west road , taipei , 10048 taiwan
4 department of life sciences , national chung - hsing university , no . 250 , guo - guang road , taichung city , 40227 taiwan
corresponding author : shu - ping wu ( wt . ude . utn @ uwgnipuhs )
this is an open access article distributed under the terms of the creative commons attribution license ( cc by 4 . 0 ) , which permits unrestricted use , distribution , and reproduction in any medium , provided the original author and source are credited .
species were collected by hand , euthanized using a dilute chloretone solution , and fixed in 10 % buffered formalin . frogs were later transferred to 70 % ethanol , and tadpoles were stored in 10 % buffered formalin . in addition to the type specimens described in this study , 343 samples that consisted of
and related taxa were collected from 22 locations throughout the island of taiwan . furthermore , three specimen of
were collected from the type locality . one was from iriomote isle and the other two were from ishigaki isle ( fig .
in addition , the eggs and tadpole morphometric characteristics were measured comprising total length ( tl ) , body length ( bl ) , tail length ( cl ) , tail height ( th ) , tail muscle height ( tm ) , internarial distance ( na ) , distance between eyes ( in ) , and tail muscle width ( mw ) ( altig and mcdiarmid 1999a , b ) . except for tl , bl , and cl , which were measured using dial calipers , tadpoles and eggs were measured under a dissecting microscope with a stage micrometer . developmental stages of tadpoles were as defined by gosner ( 1960 ) . drawings of the tadpoles were done by shw using a dissecting microscope with a camera lucida attachment .
all measurements of morphometric characteristics were taken using a dial caliper under a dissecting microscope , and measurements were rounded to 0 . 1 mm . digital webbing of the adults was recorded using savage and heyer’s formula ( 1997 ) .
t - tests were used to examine whether body size varied by gender within each taxon . an analysis of covariance ( ancova ) method was used to compare the size - adjusted means of morphometric characteristics . morphometric characteristics that satisfied the normality assumption were included in a multivariate principal component analysis ( pca ) based on the correlation matrix of size - standardized measurements ( all measurements divided by svl ) . scatter plots of the scores of the first three factors of pca were used to examine the differentiation among specimens . all of these tests and analyses were applied separately to male and female specimens . the statistical analyses were performed using sigmaplot 12 ( systat software , inc . ) .
frog mating calls were recorded using a digital recorder ( fostex fr - 2le ) and a microphone ( sennheiser me 67 / k6 ) . calls were recorded in the native habitats of these tree frogs , and environmental parameters including temperature and humidity were also recorded . avisoft saslab pro 5 . 2 . 08 ( avisoft bioacoustics ) was used to extract the maximum and minimum frequencies , as well as the width of frequency , the single note duration , and the time interval between notes of the mating calls . a rapid call and a slow call were identified . slow mating calls were compared among the subtypes in a pair - wise manner using wilcoxon - mann - whitney odds ( wmwodds ) calculations ( divine et al . 2013 ) . a bootstrap method was used to calculate the bonferroni corrected confidence intervals of the wmwodds .
polymerase ( super - therm ) , 0 . 5 μl of each primer ( 10 pm / λ ) , 1 μl template dna and ddh
o . thermal cycling was performed on a geneamp 9700 with 5 minutes at 95 °c for pre - denaturing , 35 cycles of 1 minute at 95 °c , 1 minute at 50 °c , 1 minute at 72 °c , and a final extension for 7 minutes at 72°c for both of the co1 and the 16s rrna genes . the amplicons were examined on a 2 % agarose gel for quality and fragment size . then they were purified using a geneaid pcr extraction kit and sequenced on an abi 3730 automated sequencer . estimates of genetic divergence among taxa were calculated using the kimura two - parameter model of correction for multiple substitutions at a site (
) . the transition / transversion ratio was set as 2 : 1 . chromatographs and sequences were examined and edited in bioedit 7 . 0 . 1 . (
) using the branch - and - bound search algorithm with 1000 permutation replicates to generate the null distribution . the fraction of ild null replicates with a significance value greater than the significance value of the ild was recorded . the sequences of the two gene segments were combined into one data set for subsequent analyses . dnasp 5 . 10 (
) was used to compute the population divergence conditions . tcs 1 . 21 (
) was used to reconstruct the minimum spanning network of haplotypes from each genetic population or species . a consensus ml tree was reconstructed by mega 6 (
a general time reversible model with a proportion of invariable sites and a gamma shaped distribution of rates across sites ( gtr + i + g , i = 0 . 4402 , g = 0 . 4519 ) was determined as the best - fitting model for the aligned sequences of the combined dataset using a hierarchical likelihood ratio test performed with the program mrmodeltest 2 . 2 ( nylander 2004 ) . the selected substitution model then was adopted in the reconstruction of the phylogeny by bayesian analysis and neighbor - joining ( nj ) analysis ( saito and nei 1987 ) .
the bayesian tree and the posterior probability distribution were determined using the program mrbayes 3 . 1 ( huelsenbeck and ronquist 2001 , ronquist and huelsenbeck 2003 ) . two independent monte carlo markov chain ( mcmc ) analyses were run simultaneously for 200 , 000 generations and sampled every 100 generations . to summarize the parameters and trees , the first 500 ( 25 % ) parameter values and trees were discarded . the nj analysis was conducted in paup using ml distance . gaps within the alignment were considered as missing values . the mp ( maximum parsimony ) analysis was performed with a heuristic search using 10 random stepwise steps followed by tree bisection reconnection ( tbr ) branch swapping . support for nodes was evaluated by bootstrap analysis ( felsenstein 1985 ) with 1000 replicates of the nj and the mp methods .
partial sequences of mtdna co1 gene were used as haplotypes to examine the genetic structures of the three subtypes . we calculated the fst , nm ( number of immigrants per generation , nm = ( ( 1 / fst ) - 1 ) / 4 ) , nucleotide diversity ( pi ) , and haplotype diversity ( hd ) . these calculations were made to comprehend the divergence and the intensity of gene flow among these taxa and to infer the evolutionary histories experienced by these taxa or their populations ( wright 1965 , avise 1994 ) .
) , collected on ligia timber trail , 1250 m elevation , taitung county , taiwan ( fig .
nchuzool 11311 - 13 collected on 2 august 2005 by hui - ming huang at the type locality ; nchuzool 11431 , asizam 0054 collected on 15 september 2005 by shu - ping wu at the type locality ; nchuzool 11442 ( eggs and tadpoles ) , collected on 7 february 2006 by shu - ping wu at the type locality ; nchuzool 11448 , collected on 16 february 2006 by shu - ping wu at 425 meters above sea level , at antong , hualien county ( fig .
ligia timber trail , 1250 meters above sea level , taitung county , taiwan , republic of china ( fig .
the epithet berylliniris is a compound word formed from beryllin ( l . ) , green - colored , and from iris ( l . ) , iris of the eye , and is treated as a noun in nominative singular in opposition to the generic name .
habitus moderately slender and somewhat flattened , size moderate ( svl 40 . 1 mm ) ; head wider than long ; tip of snout pointed ; snout obtuse in lateral view ; nostril barely visible from above ; canthus rostralis curved , prominent ; loreal region concave , oblique ; interorbital distance 1 . 5 times wider than upper eyelid width ; nostril oval , oblique , closer to tip of snout than to eye ; internarial distance slightly longer than nostril - eye distance ; eye diameter larger than nostril - eye distance ; pupil horizontal ; tympanic region oblique ; diameter of tympanum approximately half of eye diameter ; tympanum distinct , round ; tympanum to eye distance smaller than half tympanum diameter ; supratympanic fold from posterior tip of eye to base of arm ; jaw angle almost to posterior rim of tympanum ; premaxillary and maxillary teeth present ; choana exposed ; vomerine teeth present only on left side ; tooth patch oval , about half of choana diameter . vocal slits near commissure of jaw , slit - like .


the surface pen is an active stylus and digital pen developed by microsoft for its series of surface computing devices .

it is designed to showcase the pen computing capabilities of microsoft 's windows 8/8 .1 and windows 10 operating systems .

surface pen

surface pen surface active stylus capacitive digital pen 384 levels of pressure sen . . .
the original surface pro 3 active pen ( top ) , the surface pro 4 pen ( center ) and the new surface pen .
surface pen , surface stylus pen with 1024 levels of pressure sensitivity and alumin . . .
surface pen , surface stylus pen with 1024 levels of pressure sensitivity and aluminu . . .
* surface pen sold separately for surface pro , surface laptop and surface book 2 .
accurate surface pen , ciscle active surface stylus pen with max 4096 levels of pres . . .
surface pen , fabtek active pen with 1024 pressure levels , compatible with microsoft . . .
surface will then re - discover the surface pen when you hold down the top button of the surface pen for about seven seconds–until the light in the middle of the pen clip starts to flash ( on the original surface pen ) or the light on the flat side of the pen glows white ( on the new surface pen ) .
step 2 : pair your surface pen please note that you can pair the new surface pen with surface pro 4 and surface book as well as with surface 3 or surface pro 3 .
tests in clip studio paint and photoshop consistently showed that the new surface pen greatly outperformed the sp3 pen and did slightly better than the sp4 pen .
devices that ( eventually will ) support new surface pen enhancements : surface 3 , surface pro 3 , surface pro 4 , surface book , surface laptop , and surface studio .
compatibility — surface studio , surface laptop , surface book , surface pro , surface pro 4 , surface pro 3 and surface 3 .
whatever the situation , here’s how to pair a surface pen to your surface device .
compatible with surface 3 , surface pro 3 , surface pro 4 , surface pro and surface book .
surface pen , betop surface active stylus with 1024 levels of pressure sensitivity f . . .
4 surface pen tilt functionality is available now with surface pro . available with other surface devices via windows update .
the microsoft store listing for the surface pen contains the following footnote :
surface pen tilt functionality is available now with surface pro . coming to other surface devices via windows update soon .
here is what owners of previous surface devices should expect with the all - new surface pen .
here’s an easy two - step guide on how to connect your surface pen with your surface 3 , surface pro 3 , surface pro 4 , or surface book .
how does your surface pen help you # dogreatthings ? share with us on twitter via @ surface .
the surface pro 3 , with n - trig pen technology . image : microsoft
the surface pen is practically synonymous with surface at this point . surface pro is all about inking , and microsoft showed off how its latest 2 in 1 does inking with the all - new surface pen .
the new surface pen and microsoft ' s surface dial tool . the dial now works with the new surface pro , too .
the pen runs for $ 49 . 99 usd ( the surface pro 3 comes with one pen already ) , and you can snag the pen in a silver color . the pen utilizes bluetooth 4 . 0 and is 135mm x 9 . 5mm diameter .
the surface pen also now supports tilting for the first time , making shading much easier . the surface pen is backward compatible with the surface 3 , surface pro 3 , surface pro 4 , surface book , and surface studio , so its new features work with those older devices .
perhaps you just bought a new surface tablet and skipped through the set - up instructions for the surface pen . or maybe you upgraded to the new surface pen for your surface pro 3 and need help pairing it .
the redesigned surface pen appears to be a bit longer and sleeker than its predecessor .
for the first time , microsoft will be selling surface pen separately for $ 99 .
due to the focus in the improved surface pen , questions were raised as to whether microsoft had improved the hardware of the surface pen or debuted a new pen like it did with the past generations of surface . it turns out the answer to that question is no , the surface pen for the surface studio is the same pen that we’ve known and loved for the past year and a bit . instead , the surface pen will have an improved performance on the studio due to the technology built into the studio .
connectivity : bluetooth 4 . 0 . user has to manually pair the surface pen to the surface tablet by going to pc > device > bluetooth settings while holding down a button on the surface pen .
a microsoft spokesperson told mspu that “the technology in the pen hasn’t changed , but pen enhancements are linked to the new devices themselves . so [ it is the ] same surface pen but improved performance with the technology built into surface studio . ”
surface pen , active stylus with 1024 levels of of pressure sensitivity for microsof . . .
write and draw naturally with the surface pen that ' s better and faster than ever .
the refurbished surface pen works exclusively with surface pro and surface pro 2 , and makes it easier to draw and take notes in office applications .
surface pen , ciscle digital stylus pen : high - precision 1 . 0mm , with 1024 levels of pressure sensitivity , right click and erase buttons for surface pro ( 2017 ) , surface pro 4 , surface laptop / book _ ( black )
while these advances are coming , they are not here today – at least not completely . in my usage of the new surface pen on the surface studio and surface pro 4 , i noticed an overall increased sensitivity and reduced iaf – there is just less effort needed to use the surface pen . that experience is like the new wacom bamboo ink pen , which is a cheaper alternative to the surface pen .
microsoft surface pen experience the ultimate modern writing experience with surface pen . includes pen tip kit compatible with surface 3 , surface pro 3 , surface pro 4 , surface pro and surface book . experience the ultimate modern writing experience with surface pen . write , draw or mark - up documents , digitally . take notes to quickly capture your thoughts and instantly convert to text for sharing and searching with ease . simply click once to open onenote , or click and hold to instantly access your personal digital assistant , cortana . the surface pen is pairedwith a pen tip kit to help you choose the tip that feels most natural to you and attaches magnetically to your surface so you never leave it behind .
one of the things microsoft focused on with the new surface pen is latency . in order to accurately emulate a pen and paper , the surface pen has to “flow” as effortlessly as an old - fashioned pen . microsoft claims , “the new surface pen more than doubles the accuracy of the previous version , and with latency so low it is virtually imperceptible to the human eye , making it twice as responsive as the apple pencil . ”
the surface pen comes in the new surface color range of platinum , black , burgundy , and cobalt blue , matching the new surface pro type covers and the new surface arc mouse .
as lowensohn himself points out later in his article , the surface pen is good for far more .
microsoft certified surface pen , stylus pens supporting 500 - hour playing time 180 - da . . .
in the following videos i drew my hand across the surface pro 3 , first with its original pen and then with the others . the results are very dramatic between the surface pro 3 and other pens , but much less so between the sp4 pen and the new pen .
surface pen feels as natural as pen on paper ; with precision ink on one end and an eraser on the other . palm block technology ignores the pressure from your hand when it senses you’re using pen . now with 1024 levels of pressure sensitivity and reduced latency , surface pen lets you draw and paint with artistic precision and the control you want .
owners of older surface pcs may wonder what the new surface pen can do for them . here is what you need to know .
are you going to pick up the new surface pro ? or a new surface pen for a previous - gen surface maybe ? let us know in the comments .
when we wrote and drew pictures with the pen , it felt effortless , like using an ink - based pen on a normal sheet of paper .
the most notable difference between the new surface pen and its predecessor is the removal of the clip located near the eraser . that part was a vestige of the original surface pro 3 pen which was not magnetized .
the new surface pen brings significant changes . here are the key features being advertised or that we noticed .
microsoft just lifted the lid on the brand new surface pro , and it comes with some nifty accessories , including a new surface pen .
an aaaa battery is pre - installed in the new surface pen for surface pro 4 and surface book . if it needs to be replaced , follow the steps above .
* surface pen , arc mouse , signature type cover , and dial sold separately . some software sold separately .
apple has a patent for its own pen , called , naturally ipen , but no one knows when , or if , they will come out with their own pen . remember , this is the company that declared pen dead .
unscrew the top of the pen ( the part with the purple button ) .
microsoft has been pushing its pen technology in windows and surface for years . the new surface pro , announced today , includes what microsoft describes as the “fastest pen in the world . ” microsoft’s distinguished engineer and co - inventor of surface , stevie bathiche , excitedly revealed the new surface pen to me in a recent interview . “we did a huge step with the pen , a massive step , ” claims bathiche . “i’m so pumped about this . ”
you click the tip of the pen and the surface pro 4 opens microsoft onenote . much like pretty much every device we have seen at the event , the pen also comes with cortana integration .
the new pen ' s tilt support works exceptionally well on the new surface pro , opening up shading possibilities that were previously inaccessible to surface artists .
the surface also sports pen - based input — fulfilling microsoft’s dream of effectively integrated pen computing in windows , something that began in the 90s . it looks like the surface’s pen is more attuned to the hardware than something like samsung’s galaxy note , which should make it easier to take notes on the glass screen .
call it the great pen wars of 2017 . both microsoft and apple have moved the needle on their respective smart pen computing solutions . so which one is better ?
ever since microsoft introduced its surface tablet pcs , the company has pushed the importance of stylus input using the included pen . accordingly , the surface pro has always included the pen in the box . the company has also made significant improvements to the usefulness of the pen in the most recent major update of windows 10 .
pressure sensitivity : 256 levels of pressure sensitivity ; creates thicker , darker lines the harder you push . the surface pen also incorporates palm block technology to allow you to rest your palm on the screen without interfering with the surface pen’s input .
imagine how much better the pen would look if clad in a molded magnesium cylinder .
never has there been a digital pen like this one . we couldn’t be more excited for surface fans to use it , and to help you coordinate your personal look , surface pen is available in platinum , burgundy , cobalt blue and black .
this video is a technical look at the surface ( pro ) 3 pen up really close ( in macro ) . it compares the pen to previous model n - trig pens like the motion cl900 and fujitsu q702 pens . we then break down the cl900 pen to show you what is inside . we explain the functions of the pen including the purple bluetooth button on the end . at the end of the video we unbox one of the new generation surface 3 coloured pens ( colored ; - ) for those in the usa ) showing that the pen loop now comes in the pen box .
if you are on an older surface , getting the new surface pen is a mixed bag for now , and it ' s best to wait . ( the surface pro 1 and pro 2 used completely different pen tech , so they won ' t see any such improvements . )
i was surprised that when i received the pen it came with additional tips . i enjoy changing the tips and trying them out . pen occasionally disconnects but that’s very rare .
unlike the surface pro 4 , the new surface pro doesn’t include the surface pen — an unfortunate decision on microsoft’s part . the company cited a few reasons for making the pen an option , including the new multi - color scheme , customer feedback that shows that not everyone uses the pen , and the fact that many people who buy the surface pro will be upgrading from previous models . the pens – and the type covers , for that matter – are compatible across the surface 3 , surface pro 3 , surface pro 4 , and surface pro .
the first - generation surface pen arrived alongside the surface pro in 2012 . since then , microsoft’s peripheral has endured several generations of enhancements and refinements up until the release of microsoft’s fifth - generation surface pro 2 - in - 1 last month . the digital pen is now offered as a stand - alone product , supporting the surface book with performance boost , the surface laptop , the surface studio , and surface - branded devices .
clicking the button on a traditional pen ejects the tip so you’re ready to write . pushing the button on a surface pen opens the windows ink workspace so you’re ready to write . the windows ink workspace gives you easy access to sticky notes , sketchpad , screen sketch , and pen - enabled applications .
in my own limited time with the surface pen i saw a big improvement . i couldn’t detect any lag , and the tilt action felt very similar to a wacom pen . i’m not a professional illustrator , so we’ll have to see what creatives think , but microsoft has definitely improved the pen here . although it ' s slightly more refined and lacks the clip at the top , the new surface pen is still not fully circular , as it’s designed to magnetically snap onto the side of surface devices . some creatives will still dislike the shape of the actual surface pen as a result .
keep in mind the surface pen utilizes bluetooth to launch onenote – that’s it . the aaaa battery is crucial for the stylus functions , but the two small batteries at the top of the pen are simply for the ability to launch onenote via the top of the pen . you can turn off bluetooth and the pen will work fine – except that you will not be able to launch onenote .
the no - pen sku appears to be limited to the us only at this time .
“like putting pen to paper or brush to canvas , the new surface pen supports a natural , immersive experience that sets you free to create , learn , and express yourself in completely new ways , ” explains microsoft .
meanwhile , the gilded edition of the surface pen offers a glitzy take on the previously released stylus , but no other differences . it’s set to retail for $ 60 , and comes packaged with the surface pen tip kit that was released late last year .
with surface hub , your teams can collaborate in new ways , with apps optimized for large - screen pen and touch experiences .
surface pro 3 and 4 , surface book and surface studio users will definitely see a difference with the new pen , but whether that difference is worth the $ 100 upgrade price is a different story .
i have used every model of the surface pro since the initial launch , and i have carried around the surface pen since 2012 . i have rarely used it , though , until now . the new surface pen that microsoft rolled out this year alongside the launch of the latest surface pro is a game changer . it may look like the same pen from the outside , but there are significant updates to the surface pen itself , as well as sweeping changes in windows and across microsoft office applications that integrate the device more seamlessly into the computing experience .
another advantage of the pro 4 pen over the pro 3 is that only one aaaa battery is needed to power the [ pro 4 ] pen , compared to the aaaa battery and the three 319 button cells needed to power the pro 3 pen . also , the pro 4 pen connects to the right - hand side of the tablet easier than the pro 3 .
with the surface pro 3 you cannot enjoy some of the functions of the new pen . this includes being limited to 256 levels of sensitivity instead of 1024 and not being able to customize the actions of the pen button .
out of the default loadout , and now selling it separately ( the new pen is $ 99 , while most previous surface pros came bundled with the older model pen , which is still available for around $ 60 ) .
with the surface pen ( $ 50 with the surface 3 ; included with surface pro 3 ) , you can tap the top button on the pen once to open the onenote app — even if your surface is locked . then you can scribble a quick note using the pen , and if you want , turn the tablet off . your notes are saved automatically to your default onenote notebook of choice , which is very cool .
the new surface pen is only as good as the software behind it , which is why we’ve worked with office to create new inking features that take full advantage of the hardware technology within surface pro and surface pen . these include customizable pen tips that roam with you across all your office 365 apps , regardless of device , and amazing support for tilt and shading across all office 365 apps .
the results are impressive . no , not the drawings , those are terrible . what i mean is , the new surface pen , used in tandem with the new surface pro , delivers a very natural experience . in this case , “natural” means that the experience emulates , if not outright duplicates , the experience of doing so with a real pen , pen , or paintbrush on real paper . it doesn’t feel like you’re gliding the surface pen across glass . which of course you are .
not only does surface pro work with on - screen surface dial , it was also designed with the new surface pen , our most natural and responsive pen yet , with over 4096 pressure points , only 21 milliseconds of latency and new tilt functionality . it gives people , from artists to mobile professionals , a more fluid and real - to - life inking experience . over two times more accurate , with four times more pressure sensitivity than the original surface pen , we engineered surface pen to be the fastest pen ever * * * * * . and with the lowest parallax in the industry , we bring your ink and your content closer to your fingertips .
now that your surface pen is read to use , try using it to do things like jotting down notes , taking screenshots , and planning a trip on your surface .
surface pro and surface pro signature type cover are available for pre - order today and will ship on june 15th worldwide ; surface pen will be available in the coming weeks * * * * . surface pro starts at $ 799 . 99 usd .
tips and tricks about how to use microsoft surface pen on surface pro 4 tablet for better productivity . ► subscribe now for daily new tech and gadget videos urltoken how to customize the microsoft surface pen on surface pro 4 to perform specific tasks : urltoken microsoft surface pro 4 - how to take screenshot ( two methods ) : urltoken how to close background running apps on microsoft surface pro 4 how to hard reset or factory reset the microsoft surface pro 4 : urltoken
the pen is available in four colors : black , cobalt blue , platinum , and burgundy .
the next - generation surface pen features increased pressure sensitivity , tilt for artistic shading , and true real - time writing . like putting pen to paper or brush to canvas , the new surface pen supports a natural , immersive experience that sets you free to create , learn , and express yourself in completely new ways . learn more : urltoken audio description video : urltoken
personalise surface pro and express yourself with next - generation tools in rich colours , 3 including new surface arc mouse , * surface pro signature type cover * and surface pen * — with real - time writing and tilt 4 for shading — plus on - screen support for surface dial . *
microsoft redeemed itself after the windows rt fiasco with a sleek , windows 8 . 1 surface 3 tablet that really shines when combined with the optional surface pen and onenote app .
for these and other reasons , microsoft is upgrading its surface pen to a new version that offers 4 times the pressure sensitivity , plus tilt capabilities , just like apple pencil . at its may unveiling , microsoft claimed that the new surface pen , when used in tandem with the new surface pro and its unique “pixelsense accelerator chip” offers the fastest - ever smart pen experience . an obvious jab at apple pencil .
when i try to pair the new pen with my surface 3 , it says connected for a second or two , then it says not connected . pen doesn’t work at all except the purple button on the top will launch onenote .
microsoft’s brand new 12 - inch surface pro 3 tablet is available for purchase and comes with a fantastic stylus called the surface pen . the stylus connects to your surface pro 3 via bluetooth and is powered by batteries . if you are having trouble with the pen’s connectivity , you might have the battery inserted backwards .
compatibility : microsoft’s surface and surface pro 3 2 - in - 1 tablets .
the new microsoft surface pro ( left ) and the older surface pro 4 .
in addition to the hardware updates , microsoft is also adding a variety of surface pen and application specific features that make it easier and better to use . microsoft developed new ink types and added a pencil option to make the output of the surface pen simulate graphite on paper .
if the pen shows as “not connected” in your bluetooth list , double - check that all the latest updates are installed on your surface .
the pen magnet will only loosely attach to the right side of the sp3 , in a small area next to the power connector . in order to securely fasten the pen without the clip , surface pro 3 owners will need to dig up one of the old pen loops that could be attached to that generation ' s type covers .
the new surface pen will be backwards compatible with older surface devices , but that doesn’t mean the new tilt functionality will be available on older devices . microsoft’s new surface pro ships with this new support , and microsoft is planning to add tilt to existing surface studio and surface book devices later this year .
the old pen will work on the new system , so if you don ' t care about tilt and have a pen from a pro 3 , pro 4 , book , or studio , you can continue to use it . one less satisfactory aspect of the new pen is that microsoft no longer bundles it . with the pro 4 , only the cheapest system omitted the pen ; now they all do . worse still , the new pen is more expensive— $ 100 as opposed to $ 60 .
as highlighted last year by josh lowensohn in an article for the verge , almost all of microsoft’s 30 - second ad spots only show the surface pen used to draw circles on the screen . you’d be forgiven then for thinking drawing circles is all the surface pen is good for .
i panicked when i thought the tips from the replacement kit ( right ) wouldn ' t fit inside the new surface pen ' s collar .
within the surface app ( available in the windows store ) if the pressure curve is set at the value shown above , the lower iaf surface pen will produce the lightest strokes possible .
lower the kickstand — now featuring a deeper , next - generation hinge — to place surface pro in studio mode for the perfect writing and drawing angle with the new surface pen . *
but time stands still for no active pen , and apple just announced its own upgrades , and it has ostensibly leapfrogged surface , and surface pen , yet again . what’s interesting is that this didn’t require a new apple pencil . instead , if you use the existing peripheral with a new ( 2017 ) ipad pro and its promotion display technology , you’ll get better performance than surface pro ( 2017 ) with surface pen ( 2017 ) : 20 ms of latency vs . 21 .
but the same can ' t be said for the new surface pen , which offers a major step up to 4 , 096 pressure levels , lower activation force ( ~ 9 grams ) and tilt recognition . on paper at least , this is the pen that surface users have always wanted .
try surface book 2 and the newest surface pro at a microsoft store near you .
cushion your surface pro or surface laptop with a colorful , limited edition marimekko sleeve .
with surface pen and windows ink , nothing comes between you and inspiration . quickly turn thoughts into action and access your inking apps in one place .
overall , the surface pro 3 pen is an excellent tool for note - takers and digital artists looking for a realistic drawing experience on a tablet .
back in may i spoke to jared spataro , general manager of office for microsoft , about the surface pen . he explained that the goal for microsoft was to make using the surface pen with a surface device better than a traditional pen and paper . he told me , “not only does it allow you to do all of the things you can do with paper – it gives you a variety of tools and features that paper can’t provide . ”
express yourself with one of the vibrant pen colors designed to mix and match with your type cover . choose a pen tip that’s right for you . stay with the pre - installed hb tip for writing and drafting , or swap to a 2h tip for detailed drawing , shading and illustrating . choose from one of 4 tips in our surface pen tip kit .
when the pen now appears in the list of bluetooth devices , select it and then choose “pair . ”
surface 3 retains all the other benefits provided by surface pro 3 , like the compatibility with the docking station , surface pen support , and more , but offers them in a much more portable package . what ' s not to like ?
surface pro 4 vs . surface book vs . surface pro 3 : is it worth upgrading and which one is best for you ?
ok , those numbers are close . and they suggest that the performance of apple pencil and the new surface pen are , in fact , roughly identical .
the best days of the new surface pen are still ahead . when microsoft releases its new pen driver and firmware , you will see even more improvements enabled on older surface hardware . that does not mean the $ 99 is still worth it , but that is a personal decision you will need to make .
microsofts surface studio was announced last week to much fanfare . featuring the body of an all in one pc , much like microsoft’s other windows 10 device like the surface book and surface pro 4 , it bucks that staid old norm by allowing users to draw all over the screen with a new surface dial and an improved surface pen .
if you thought styluses were last seen on devices in 1990s , think again . israel ' s n - trig is one of a new breed of pen input companies , recently unseating wacom as microsoft ' s pen of choice on its new surface pro 3 tablets .
the surface pen uses one aaaa battery to feed the digitizer and two small size 319 lithium coin cell batteries , which come pre - installed in the top of the pen , to power the bluetooth section . the aaaa battery is manually installed by the user and can be accidentally inserted backwards , resulting in issues with bluetooth connectivity even though the indicator light on the pen will be on . the indicator light might confuse you into thinking the pen is working fine , when it is not .
alongside the microsoft lumia 950 , lumia 950 xl , and lumia 550 launches , the company also announced the surface pro 4 . microsoft unveiled the surface book as well , its first laptop . both devices will come bundled with the new surface pen .
surface pro 3 pen users should definitely switch to a newer pen . if tilt is important , wait to see if the sp3 will get the necessary update . if tilt doesn ' t matter to your drawing technique , save $ 30 and pick up the bamboo ink .
in australia , another strange situation . the base model surface pro 4 is au $ 1349 . take away the pen and it drops to an astonishing au $ 899 . given that the pen only costs au $ 94 , this au $ 450 saving is rather remarkable .
when all combined with our pixelsense accelerator , the writing experience feels like writing on paper , picking up every expression of the pen and reacting to the subtlest of artistic movements . artists can paint on surface and feel like they are painting on parchment , as surface pen has almost no latency and gives you the ability to do tilt and shading on surface pro . for those at work , you will feel like you are taking notes in your moleskin notebook with your favorite pen .
second , i’m worried about losing the pen . sure , it clicks into the tablet’s power connector with a satisfying snap , but it just sits there , naked , exposed , and easy to pry loose . as a result , i never felt comfortable tossing the surface into the bottomless pit of my backpack when the pen was attached . i believe pen accessories should be sheathed inside their tablets , and not strapped to the side with a hope and a prayer . regardless , if you do lose the surface pen , a replacement costs $ 30 .
there’s just one problem . when apple announced its first ipad pro back in late 2015 , it also launched a companion peripheral , the apple pencil , which duplicates and in many ways exceeds the performance and utility of surface pen . it offers tilt capabilities , for example , where ( the current ) surface pen does not .
digital artists have more to like . the redesigned surface pen boasts 4 , 096 levels of pressure accuracy compared to its predecessors ' 1 , 024 levels , and it has tilt support . this means you can ink with the pen , or angle it and shade in brushstrokes with the side of the nib , just like a real pen would . the new surface pen is also a little longer and sleeker , and it eliminates the clip . microsoft ' s also quite proud of the fact that the new pen virtually eliminates the pen ' s latency ( now just 26 ms ) between when you ink a line on the screen , and when the digital ink actually appears . finally , like the studio , microsoft now offers the option to switch to
enhanced color
from srgb .
one company that doesn ' t support pen — yet — is apple . but that may change , lebovitz said .
try the new surface book 2 and the newest surface pro at a microsoft store near you .
the improved surface pen stylus features increased pressure sensitivity with up to 4 , 096 pressure points ( from 1 , 024 ) , tilt that adjusts the thickness and texture of your ink granularly , based on the angle of your pen , for artistic shading and true real - time writing .
the magnet on the pen is also considerably stronger ; one problem i ' ve found with previous models is that the pen often gets divorced from the system when it ' s in my laptop bag ; a stronger magnet should reduce this .
this custom setup means microsoft can quickly erase , about a 100 - millisecond trail of ink , before the rest of the system catches up , so it improves the latency of the pen . “this is how we’re able to close that gap , ” says bathiche . “we have the fastest pen in the world . ” pressure - sensitivity levels on the new surface pen have increased to 4096 , alongside a 12 - gram activation force . the new surface pen is now double the speed of the previous one , and microsoft claims it’s “twice as responsive as the apple pencil . ”
experience the ultimate modern writing experience with surface pen . write , draw or mark - up documents , digitally . take notes to quickly capture your thoughts and instantly convert to text for sharing and searching with ease . simply click once to open onenote , or click and hold to instantly access your personal digital assistant , cortana . the surface pen is paired with a pen tip kit to help you choose the tip that feels most natural to you and attaches magnetically to your surface so you never leave it behind .
the surface pen is useful for a variety of tasks normally associated with traditional pen and paper , including writing and drawing on the screen ( converted to text ) , highlighting and writing on pdfs in a range of apps like onenote , adobe creative cloud , fresh paint , and drawboard pdf .
pen broke after only a few months . thought battery was dead , but took it to microsoft store just to verify .
in other words , the great pen wars of 2017 are ending with a stalemate . these are both excellent smart pens .
actually the new preview builds will install the sp4 pen drivers , properly . then you can do all the expected tweaks .
the surface pen has been around since the launch of the initial surface pro in 2012 . it was built using technology from wacom , which has been doing digital pens long before microsoft was in the 2 - in - 1 hybrid market . the surface pen provided some unique potential for the tablet - cum - laptop , but it was still more or less a novelty – at least for me .
microsoft has at times been vague about the details of the new surface pen , and the feature set is apparently spread out across devices . my educated hunch is that microsoft is going to push a driver update for the new surface pen in the next few months , but because it is still being worked on the company does not want to commit to specifics . they know this new pen can do more , but just how much more on older hardware is currently fuzzy .
billed as “the most versatile laptop , ” the new surface has 13 . 5 hours of battery life , intel’s seventh - generation “kaby lake” core processor ( the same chip powering microsoft’s $ 999 surface laptop ) , a new alcantara keyboard and a next - generation surface pen .
as per microsoft , surface pro has up to 50 percent better performance and 50 percent better battery life than it predecessor and 35 percent better battery life than an ipad pro . its 12 . 3 - inch pixelsense touch display supports microsoft’s surface dial and the new surface pen .
the new pen has a flat side and the single side switch is much smaller . the slightly longer collar is now entirely plastic , presumably to help conductivity during tilt operations . the new pen is the same length and weight as the prior generation .
the surface pro ' s pen has seen a significant upgrade . it now has 4 , 096 pressure levels ( up from 1 , 024 ) , and microsoft has added tilt support ; you can hold the pen upright to write with a fine point , or you can angle it for shading . a wide range of artists have already made surface pros an important part of their toolkits , and tilt support will further enhance this . ( microsoft has promised to add support for the new tilting pen to the surface pro 4 , surface book , and surface studio through a firmware update , though there ' s no eta yet . )
the surface pen is quite expensive and runs for $ 50 if purchased separately . the pen is cased in aluminum , has over 250 levels of pressure sensitivity , and palm block technology . this means you can write and draw without worrying about resting your hand on the screen and messing up your creation .
as far as looks go , the surface pen closely resembles a typical click pen . it has a good weight of 20 grams , a substantial diameter of 9 . 5 millimeters , and a length of 135 millimeters . the stylus comes in silver , takes batteries , and has bluetooth 4 . 0 .
which i believe they are , having now used both . and in their optimal , most performant scenarios : the new surface pen with the new surface pro , and the apple pencil with a new ( 2017 ) ipad pro .
in other ways , microsoft ' s changed course from prior generations . the new surface pro will go out the door with an intel core m option ( it was a later arrival in the prior generation ) . none of the new surface pro devices will be sold with a surface pen . that has nothing to do with user reluctance to use the pen , microsoft says , but merely reflects that surface owners who choose to upgrade may already own one .
one of those activities is the whiteboard app , which allows for three people to ink on the screen at any one time . the hub ships with a pair of pens , holstered on either side . once you remove a pen from the holster , whiteboard launches . icons appear on the bottom of the screen so you can choose ink color , among other pen options . unlike the surface pen , there are no other buttons on the surface hub pen , though you can “erase” by flipping it over . you won’t find any fancy icons , brush motifs , or digital stickers in whiteboard ; each pen has a choice of what appears to be five colors , and there’s a lasso tool for moving digital ink around on the page .
this will especially come in handy when drawing on the surface pro 3 with the just - announced touch - friendly photoshop cc , along with writing a note in onenote . you can even click the top of the pen and immediately activate your surface pro 3 device and begin taking notes in onenote . double clicking the top of the pen captures a screenshot !
microsoft has announced new premium models of its surface book and surface pro 4 computers , with extra memory and storage to satisfy the needs of power users . the company is also launching a gold version of its surface pen accessory , which will be sold separately from the new hybrids .
the pen also makes it easy to create whiteboard - like drawings and diagrams in onenote . microsoft did a great job with
palm rejection ,
so resting your palm on the tablet screen as you draw with the pen doesn ' t cause problems .
i have a sp3 and a sp4 but have found i prefer to use the sp3 pen on both devices . i honestly can ' t pinpoint exactly why , but i just prefer the feel of the sp3 pen in my hand . personal preference i guess
but microsoft formally made smart pens a core feature of the pc starting with windows xp tablet pc edition in 2002 , and it has been steadily improving the platform ever since . and today , microsoft’s surface pen is routinely cited as the best example of what we now think of as an active smart pen .
to adjust pen pressure you ' ll need to install the surface app from the windows store . for all of my tests i kept pressure settings centered . with the new pen ' s much lower iaf , you will likely need to lower your tip sensitivity as illustrated below in order to see finer strokes .
set next to each other , the surface pro 4 and the new $ 799 surface pro are virtually indistinguishable , especially when matched up with the surface pro 4’s signature type cover . both boast 12 . 3 - inch pixelsense displays , but the new surface pro ( 2017 ) adds a better keyboard , reclines to a surface studio - like 165 degrees , and takes advantage of a new , more sensitive optional surface pen . you ' ll have the choice of either a more traditional type cover keyboard ( $ 129 ) or a new signature type cover with the alcantara fabric used on the surface pro 4 , for $ 159 . the surface pen will cost $ 100 .
my first microsoft surface pen was terrible . it would shut off randomly in the middle of lectures , causing me to miss 20 minutes of lecture trying to fix the dang thing .
the new surface book in full recline ( front ) , with the older surface pro 4 behind it .
the new surface pen is designed to compliment the 2017 surface pro ' s next - generation screen , which takes advantage of unprecedentedly low latency inking thanks to new tech . custom silicon in both the pen and the surface pro screen reduce the latency between your ink strokes and the display to levels
virtually imperceptible
to the human eye , according to microsoft . the company also says the new surface pen is much more responsive than the apple pencil designed for the ipad pro , and it benefits from a boost in pressure sensitivity at 4 , 096 levels , up from 1 , 024 in the previous version .
surface pen offers real buttons that do real things , something i kind of miss with apple pencil . the top of surface pen , for example , functions as an eraser in most apps . and you can press it to launch windows ink workspace . ( or some other app of your choosing . ) there’s also another button on the barrel , which is likewise programmable .
turn bluetooth on . if surface pen appears in the list of discovered devices , it may not be properly paired yet . so , select it from the list and remove the device .
dozens of pen - and - touch enabled devices are on the market already , and more are on the way , leibovitz said .
a lot of devices already included a touch sensor that supports pen , even if those devices currently only support touch ,
he said .
for those devices we have a software solution that can activate the pen component , allowing users to enjoy both .
the collar on the new pen ( left ) is longer and plastic for tilt signal conductivity . the side switch has been shortened .
interestingly enough , microsoft has replaced the wacom digitizer used in earlier surface models with an n - trig stylus . this pen supports 256 points of pressure and there is reportedly no latency or parallax effects . in fact , when you are writing something on the surface pro 3 , it will feel as if there were no distance between the tip of the pen and the screen surface . this means no delay and a natural ink experience .
founded in 1999 , n - trig’s technology is the industry’s only combined active pen and touch solution available today . the company’s solutions support the latest windows and android operating systems – allowing oems to develop slim pen and touch - enabled mobile devices using only a single sensor .
watch your ideas come to life in brilliant colour and razor - sharp resolution on the 12 . 3” pixelsense™ display with a stunning screen that responds to the new surface pen * and touch .
the surface pro will go on sale alongside the surface laptop , which was unveiled at the start of may .
what are the diffrent pen tips for and how can you get the most of the suface pen . quick demo with edge and onenote all the tips & tricks ; in this video can also be applied to the surface pro 3 ( and 2 and 1 ) as long as you use windows 10 and onenote on your machine .
yes . i have both pens and the sp4 pen does stick to the top of the left side but very weakly . it ' s far better sticking it to the charging port . it can still fall off with a shake , but stays on if the sp3 is immobile . the pen also seems more solidly on there if it ' s upside down , for some reason . obviously , with the sp3 in the dock , you can ' t stick the pen to either side of the surface , but the left side of the dock itself is magnetic , so the pen parks there quite nicely .
the ipad pro takes on the surface pro 3 as a productivity tool and , since the stylus ( or pencil ) is an integral part of using these devices , we took a look at how the apple pencil stacks up against microsoft’s surface pen .
one reason pen took a back seat was because it has a number of technical issues that touch doesn ' t : palm rejection , for example , in which the device has to figure out if it is the pen or a user ' s palm resting on the screen .
microsoft ' s new $ 99 surface pen is a steep upgrade from the previous iteration that costs $ 59 ( and which is still available ) . there are quite a few enhancements that may justify the cost for new surface pro owners . but for older surface devices , the value is not too clear .