zacanthoides is an extinct cambrian genus of trilobite .

it was a nektobenthic predatory carnivore .

its remains have been found in canada ( british columbia , especially in the burgess shale , and newfoundland ) , greenland , mexico , and the united states ( alaska , nevada , utah , vermont , and idaho for which z. idahoensis is named ) .

its major characteristics are a slender exoskeleton with 9 thoracic segments , pleurae with long spines , additional spines on the axial rings , and a pygidium that is considerably smaller than its cephalon .

zacanthoides

in 1887 carl rominger published an engraving of a nearly complete and markedly spiny trilobite and named it embolimus spinosa . in 1908 charles walcott introduced the combination zacanthoides spinosus for the mount stephen species and for a similar trilobite from nevada . the next change came in 1942 , when charles resser at the united states national museum asserted that the mount stephen species was sufficiently distinct that it required a new name . resser chose to honour the man who first formally described many of the common mount stephen trilobites , and zacanthoides romingeri remains the combination in use today .
name : zacanthoides typicalis ( walcott , 1886 ) trilobites order corynexochida , family : zacanthoididae locality : lincoln co . , nevada stratigraphy : glossopleura zone , chisholm shale formation , middle cambrian remarks : collected and photographed by andrew milner . specimen is 3 . 4 cm long including thoracic spine
albertella , albertellina , albertelloides chuchiaspis , danjiangella , delamarina ( / delamarella ) , eozacanthoides , fieldaspis , mendogaspis , mexicaspis , micmaccopsis , panxinella , paralbertella , parkaspis , prozacanthoides , pseudozacanthopsis , ptarmiganoides , qingzhenaspis , stephenaspis , thoracocare , tianshanocephalus , ursinella , vanuxemella ( = vistoia ) , xuzhouia , zacanthoides ( / embolimus ) , zacanthopsina , zacanthopsis .
here is the last piece of the trilobites write up from our
sw site
( stevens way , ashfork , az . ) featuring the interesting pygidiums of the zacanthoides walapai trilobites we found in huge abundance in the bright angel shale . ill post a few nice shots here , and at the end a link to the full ( monstrous ) write up on our paleo web site . thanks for looking !
specimen count 1 site number 55c record last modified 5 jul 2018 geological age paleozoic - cambrian - middle stratigraphy ute ls - spence sh mbr nmnh - paleobiology dept . common name trilobite taxonomy animalia arthropoda trilobita collector spence walcott burling see more items in paleogeneral types : arthropoda arthropoda trilobita type paleobiology place bear lake county , idaho , united states collection date 22 sep 1906 type status paratype type citation resser , c . e . 1939 . smithsonian misc . colln . 97 ( n . 12 ) : 10 , unfig . usnm number pal53435 published name zacanthoides holopygus resser
bathyuriscus rotundatus was first described in the same 1887 publication as several other important mount stephen trilobites . carl rominger initially used the name embolimus rotundata for partial specimens of this trilobite , and named a second similar species in his collection embolimus spinosa ( now known as zacanthoides romingeri ) . in 1908 , walcott revised rominger ' s original species name to yield the combination bathyuriscus rotundatus , still in use today ( walcott , 1908 ) . along with the co - occurring elrathina cordillerae , b . rotundatus is a signature fossil for the middle cambrian bathyuriscus - elrathina zone in the southern canadian rockies .
the most common fossil found in the green shales at our sh locality are trilobite pygidiums for the zacanthoides walapai species . they are joined by assorted cranidiums , thorax and rib segments , and hordes of hyolithids , and a very rare coralomorph . generally low diversity such as this site suggests has been attributed to a stressed environment , with perhaps low food sources , aggressive wave action , or an influx of fresh water from the nearby deltas . this was a shallow sea outbound from the deposits known as the tapeats sandstone which marked shore , delta and beach deposits . combined with the deeper water muav limestone , this trio of formations is known as the tonto group . all three can be found outside the grand canyon to the south in small limited areas such as here , yielding an opportunity to explore the paleo fauna without hiking miles and thousands of feet into the grand canyon .
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jell , p . a . & j . m . adrain . 2003 . available generic names for trilobites .
the list of available names for trilobite genera below is arranged alphabetically by family in the order presented in jell & adrain ( 2003 ) . if you know the genus ( or subgenus * ) of a specimen of trilobite and wish to know its family , superfamily , suborder , and order , you can use the
find
function of your browser ( typically ctrl - f in a windows environment ) to search this list for your genus / subgenus . once you know the order , suborder and superfamily ( in brackets ) and family (
) , you can browse through the nine order fact sheets ( by clicking on the choices below ) and learn more about your trilobite ' s classification . for example , if you search for the trilobite genus
you will find it listed in the family encrinuridae , a member of the order phacopida , suborder cheirurina , superfamily cheiruroidea . clicking on the phacopida fact sheet image below will take you to that order and you can learn more about the constituent families of the superfamily cheiruroidea , and the other genera in the encrinuridae . there are over 180 families of trilobites , and about 5000 genera , which contain the 15 , 000 + described species of trilobites ! in december of 2005 , i visited with dr . jell in brisbane and received permission to offer the full article in pdf format here . you must have adobe acrobat or acrobat reader to open the file . click the citation above , or the link here :
* in the list , subgenera are included and not distinguished from names of genera , since subgenera may be elevated to full genus and are therefore available generic names .
abadiella ( = parabadiella ; = danagouia ) , guangyuanaspis , guangyuania , lunolenus , malongocephalus , shaanxia , sibiriaspis .
acastava , acaste , acastella , acastellina , acastocephala , acastoides , acastopyge , armorigreenops , asteropyge , baniaspis , bellacartwrightia , bradocryphaeus , braunops , breizhops , centauropyge , chimaerastella , coltraneia , comura , cryphina , delocare , deloops , destombesina , dunopyge , echinopyge , erbenochile , ewacaste , feruminops , gourdonia , greenops , gudralisium , hallandclarkeops , harringtonacaste , heliopyge ( = alcaldops ) , hexacosta , hollardops ( = modellops ; = philipsmithiana ) , kayserops , kennacryphaeus , kloucekia , llandovacaste , metacanthina , mimocryphaeus , morocconites , mrakibina , neocalmonia , neometacanthus , paracryphaeus , pelitlina , phacopidina , philonyx , pilletina , protacanthina , pseudocryphaeus , psychopyge , quadratispina , quadrops , radiopyge , rheicops , rhenops , saharops , sanidopyge , scotiella , sokhretia , stummiana , talus , tolkienia , treveropyge , turcopyge , walliserops .
acrocephalella , acrocephalinella , acrocephalites ( = acantholenus ) , acrodirotes , afghancephalites , asturiaspis , brutaspis , cermataspis , decus , diceratocephalina , elatilimbus , ijacephalus , kepisis , mansiella , pseudacrocephalaspina , siligerites , toxotina , trifonella .
acmarhachis ( = cyclagnostus ; = oxyagnostus ; = wanagnostus ) , agnostus ( / battus ; = acutatagnostus ) , aistagnostus , anglagnostus , biciragnostus , connagnostus , distagnostus , eolotagnostus , gymnagnostus , homagnostus , idolagnostus , innitagnostus , ivshinagnostus , kymagnostus , lotagnostus , micragnostus , obelagnostus , oncagnostus ( = eurudagnostus ) , phalacroma ( = platagnostus ) , phalagnostus ( = phalacromina ) , quadrahomagnostus , raragnostus , semagnostus , strictagnostus , trilobagnostus ( = rudagnostus ) .
agasella , agaso , agraulos ( / arion ; / arionius ; = / arionides / arionellus ; = agrauloides ) , batenoides , chittidilla ( = diandongaspidella / diandongaspis ) , chondroparia , clemenella , conagraulos , elankaspis , lenagraulos , litavkaspis , metagraulos , micragraulos , mungyongia , parachittidilla ( = amurticephalus ) , paragraulos , paraplesiagraulos , phymaspis , plesiagraulos , poriagraulos , proampyx , protochittidilla , pseudoeteraspis , pseudoplesiagraulos , qiannanagraulos , shahaspis , skreiaspis , sternbergaspis , taiganella , tetragonocephalus , tholus , tianjingshania , veragraulos , wutaishania .
aldonaia , granutaspis , ideria , perissopyge , planaspis , pumilina , repinaspis , tuvanella ( = eleganolimba ) , tuvanellus , volonellus .
alokistocare ( = pseudoalokistocare ) , alokistocarella , alokistocaropsis , altiocculus , amecephalina , amecephaloides , amecephalus ( = strotocephalus ) , annamitia , arcadiaspis , arellanella , atopiaspis , beldirella , binella , bythicheilus , chancia , chanciaopsis , danzhaiaspis , diaoyaspis , ehmania , ehmaniella ( = anomalocephalus ; = clappaspis ) , elrathia , elrathiella ( = coelaspis ; = glossocoryphus ) , eokaotaia , erdoradites , furia , ganovexopyge ( / scottia ) , huochengella , inglefieldia , jenkinsonia , kailiella , kaotaia , katunicare , kistocare , langqia , lenacare , nelgakia , parapachyaspis , parehmania ( = mcnairia ; = rowia ; = thompsonaspis ) , pedinocephalina , peregrinaspis , plesiamecephalus , proehmaniella , proveedoria , pseudomexicella , schopfaspis , trachycheilus , tympanuella , utaspis .
ajrikina , alataupleura , araiopleura , calycinoidia , caputrotundum , clavatellus , falanaspis , hapalopleura , huamiaocephalus , jegorovaia ( = hermosella ) , jiangxiaspis , orometopus , pagometopus , palquiella , paracalymenemene ( / paracalymene liu ) , plesioparabolina , pyrimetopus , rhadinopleura , seleneceme ( = alsataspis ) , sibiriopleura , skljarella ( = proaraiopleura ) , spirantyx , trigocephalus , yumenaspis , zacompsus .
ammagnostus ( = lispagnostus ; = agnostoglossa ; = tentagnostus ) , hadragnostus ( = formosagnostus ; = kunshanagnostus ) , kormagnostus ( = kormagnostella ; = litagnostoides ) , proagnostus ( = agnostascus ; = paragnostascus ) .
anomocarella ( = psilaspis ; = entorachis ) , eoanomocarella , fissanomocarella , glyphaspis ( = americare ) , hanshania , huayuania , liocare , liopeishania , liopelta , luia , lydiaspis , megalopsis , neoanomocarella , orthodorsum , paranomocarella , peishania , ( = parapeishania ) , peishanoides , plebiellus .
abharella , afghanocare , amginia , anomocare , anomocarina , anomocarioides , anomocariopsis , callaspis , chondranomocare , dilatalimbus , elandaspis , eocatuniella , forchammeria , formosocephalus , fuquania , glyphanellus , glyphaspellus , guizhouanomocare , hanivella , harataspis , hunanaspis , igarkiella , iohomia , irinia , jimanomocare , juraspis , kokuria , kolbinella , kotuia , leichneyella , lomsucaspis , longxumenia , macrotoxus , metanomocare , nadiyella , palella , paracoosia ( = manchurocephalus ) , parakotuia , paranomocare , pjatkovaspellus , qinlingia , rectifrontinella , sachaspis , schoriecare , schoriella , scintilla , sivovella , usovinurus , wutingshania , yongwolia .
antagmella , antagmus , bagradia , bicella , bilimbataia , cambrophatictor , crassifimbra , cyphambon , erzishania ( = oreisator ) , houmengia , katunia , lermontoviella , longshania , luaspides , mantoushania , onchocephalina , onchocephalus ( = litocodia ) , paraantagmus , periomma , plesioperiomma , shilengshuia , sombrerella , wanbeiaspis , xiangqianaspis , xiaofangshangia , xiaomajiella , yuehsienszella .
amorphella , aphelaspidella , aphelaspis ( = proaulacopleura ; = clevelandella ; = labiostria ) , apheloides , dicanthopyge , elegantaspis , erixanium , eugonocare , kobayashella , listroa , litocephalus , maduiya , nganasanella ( = tamaranella ) , notoaphelaspis , olenaspella , olentella , paraphelaspis , pseudaphelaspis , pseudeugonocare , taenicephalites , taenora .
archaeaspis , bradyfallotaspis , fallotaspidella , fritzaspis , geraldinella , selindella . [ profallotaspis , repinaella - see fallotaspididae ]
anataphrus , araiocaris , asaphellus ( = asaphelloides ; = asaphoon ; = hemigyraspis ; = megalaspidella ; = plesiomegalaspis ) , asaphus ( = schizophorus ) , atopasaphus , aulacoparia , aulacoparina , australopyge , baltiites , banqiaoites , basilicus ( = basiliella ; = carinobasiliella ; = dolerobasilicus / basilicoides ; = mekynophrys ; = parabasilicus ) , bellefontia , birmanitella , birmanites ( = opsimasaphus ) , bohemopyge ( / ptychocheilus ) , borogothus , brachyaspis , branisaspis , burminresia , charabaia , chengkouella , dubovikites , ectenaspis , ekeraspis , emanuelaspis , emanuelina , eoasaphus ( / anorina ) , eoisotelus , erdelia ( / maja ) , estoniites , fuyunia , gerasaphes , gog , gogiura , golasaphus , griphasaphus , guohongjunia , hazarania , heraspis , hoekaspis , homalopyge , hunjiangites , hunnebergia , huochengia , iduia , isabelinia , isotella , isoteloides , isotelus ( = homotelus ) , isyrakella , isyrakopeltis , kainisiliellina , kayseraspis , klabavia , kobayashia , lachnostoma , lamanskytes , lapidaria , leningradites , liomegalaspides , lisogorites ( = trigonoaspis ; = tangyaia ) , liushuicephalus , lonchobasilicus ( = sinomegalaspis ) , lycophron , megalaspidella , megalaspides ( = lannacus ) , megasaphus , megatemnoura , megistaspidella ( = spinopyge ) , megistaspis ( / megalaspis ; = megistaspinus ; = rhinaspis ) , merlinia , metaptychopyge , metayuepingia , mioptychopyge , mischynogorites , nahannia , neoasaphus ( = trematophoris ; = multiasaphus ; = postasaphus ; = subasaphus ) , neopeltis , nerudaspis , nileoides , ningkianites , niobe , niobella ( = metoptogyrus ) , niobides , niobina , nobiliasaphus ( = pamirotchechites ) , norasaphites , norasaphus , norinia , notopeltis , ogmasaphus , ogyginus , ogygiocarella , ogygiocaris , ogygitella , ogygites , ogygitoides , onchometopus , parabellefontia , paramegalaspis ( = dolerasaphus ) , paramegistaspis ( / varvaspis ) , paraptychopyge , paratamdaspis , parayuepingia , penchiopsis , platyptychopyge , plectasaphus , plesiyuepingia , popovkiaspis , popovkites , praecoparia , presbynileus ( / paranileus ) , priceaspis ( = fitzroyaspis ) , proasaphus , promegalaspides , protopresbynileus ( / pseudonileus hintze , 1953 ) , protoptychopyge , proxiniobe , pseudoasaphinus , pseudoasaphoides , pseudoasaphus , pseudobasilicoides , pseudobasilicus , pseudobasiliella , pseudobasiloides , pseudogriphasaphus , pseudogygites , pseudomegalaspis , pseudoptychopyge , pseudoptyocephalus , psilocephalina , psilocephalops , ptychopyge , ptyocephalus ( = kirkella ) , rhinoferus ( = lawiaspis ; = ropschiaspis ) , sanbernardaspis , shergoldina , stegnopsis , stenorhachis , suriaspis , tchukeraspis , thysanopyge ( = basilicoides ) , trigonocerca , trigonocercella , tsaidamaspis , valdaites , vogdesia , volchovites , xenasaphus , xenostegium , xinanocephalus , , yuepingioides , zhenganites ( = eosoptychopyge ) , zoraspis , zuninaspis .
anomocarellius , asaphiscus , blainia , blainiopsis , blountiella , blountina , canotiana ( = williamsina ) , cinnella , ? conoides , dunderburgella , edithiella , eoasaphiscus hajrullina , eokaninia ( / kaniniella sivov ) , eoproetus , erbenia , eteraspis , iniotoma , kaninia ( / kaniniella sivov ; = dolgaia ) , kaniniella kobayashi , lioparia ( / lorentzia , / pseudoliostracina ; = emmrichella ; = liaoyangaspis ) , luyanhaoaspis ( / luaspis peng et al . , 1995 ) , mindycrusta , paraorlovia , vega , verkholenella .
aulacopleura ( / arethusa / arethusina ; = paraaulacopleura ) , aulacopleuroides , beggaspis , coignops , cyphaspides , cyphaspis ( = novakaspis ) , dixiphopyge , harpidella ( = rhinotarion ) , latecephalus , malimanaspis ( = goodsiraspis ) , maurotarion ( = goniopleura ; = branisella ; = tricornotarion ) , namuropyge ( = coignouina ) , otarion ( = aulacopleurella ; = conoparia ; = otarionella ) , otarionides , protocyphaspides , pseudotrinodus , songkania , tilsleyia .
acidiphorus ( = goniotelina ; = goniotelus / goniurus ) , aksuaspis , bathyurellus , bathyurus , benthamaspis ( = oculomagnus ) , bolbocephalus , catochia , ceratopeltis , eleutherocentrus , ermanella , gignopeltis , grinnellaspis ( / actinopeltis poulsen ) , hadrohybus , jeffersonia ( = bathyurina ) , licnocephala ( = domina ) , lutesvillia , madaraspis , peltabellia ( = biolgina ) , petigurus , platyantyx , ? proscharyia , psephosthenaspis ( = aponileus ; = ludvigsenella ) , pseudoolenoides , punka , rananasus , randaynia , raymondites , sinobathyurus , strigigenalis , uromystrum .
bigotina , bigotinella , bigotinops , bulaiaspis , hupetina , neobigotina , ouijjania , pruvostina , serrania .
acutimetopus , asiagena , australosutura , brachymetopella , brachymetopus ( / brachymetopina ; = iriania ) , cheiropyge ( = suturikephalion ) , conimetopus , cordania , eometopus , loeipyge , mystrocephala , proetides , radnoria , spinimetopus .
anchiopella , andinacaste , australoacaste , australops , awaria , bainella ( = paradalmanites ; = paranacaste ) , belenops , bouleia ( = dereimsia ) , calmonia , chiarumanipyge , clarkeaspis , cryphaeoides , curuyella , deltacephalaspis , eldredgeia , feistia , hadrorachus , jujuyops , kozlowskiaspis , malvinella , malvinocooperella , metacryphaeus , oosthuizenella , palpebrops , parabouleia , paracalmonia ( / proboloides ) , pennaia , phacopina , plesioconvexa , plesiomalvinella , prestalia , probolops , punillaspis , renniella , romanops , schizostylus , talacastops , tarijactinoides ( = bolivianaspis ) , tibagya ( / schizopyge ) , tormesiscus , typhloniscus , vogesina , wolfartaspis .
calodiscus ( / goniodiscus ; = brevidiscus ) , chelediscus , korobovia , neocobboldia ( / cobboldia ; = margodiscus ) , pseudocobboldia , sinodiscus ( = tologoja ) .
alcymene , apocalymene , arcticalymene , calymene ( / calymena / calymaena / calymmene / calymmena ) , calymenella , calymenesun , colpocoryphe ( = thoralocoryphe ) , dekalymene , diacalymene , flexicalymene , gravicalymene , limbocalymene , linguocalymene , liocalymene , metacalymene , neseuretinus , neseuretus ( = synhomalonotus ) , nipponocalymene , onnicalymene , papillicalymene , paracalymene , platycalymene ( = sulcocalymene ) , pradoella , protocalymene , reacalymene , reedocalymene , salterocoryphe , sarrabesia , spathacalymene , sthenarocalymene , tapinocalymene , thelecalymene , vietnamia .
acheilops , agelagma ( ? = paradistazeris ) , buttsia , buttsiella , catillicephala ( / cephalocoelia ) , catillicephalites , coephalocoeliaspis , cryptoderaspis , distazeris , galeaspis , lajishanaspis , madarocephalus , matania , onchonotellus ( = onchonotina ; = guotangia ; = seletella ) , onchonotopsis , onchonotus , pemphigaspis ( = hallaspis ) , peracheilus ( = acheilus raymond ) , qilianshania , stenochilina , theodenisia ( / denisia ; = calculites ; = mannschreekia ) , triarthropsis , tumidulaspis , tuojiangella , urbanaspis , waergangia , welleraspis ( = avonaspis ) , yukonaspis .
bonneterrina ( = holstonia ; = piedmontia ) , carinamala , cedaria , cedarina , henadoparia , jimachongia , vernaculina .
? ajacicrepida , asiocephalus , boschchekulia , cataplotaspis , ceratopyge , cermatops , charchaqia ( = aplotaspis ) , diceratopyge ( = paraceratopyge ) , dichelepyge ( = bicornipyge ) , dipleuropyge , guozia , haniwoides ( = yuepingia ) , hedinaspis , hunanopyge , hysterolenus ( = ruapyge / hectoria ) , kaltykelina , kaufmannella ( / kaufmannia ) , kogenium , lopnorites , macropyge ( = haniwapyge , = lichapyge ; = macropygella ) , mansuyella , nannopeltis , neohedinaspis , onychopyge ( = prionopyge ) , proceratopyge , promacropyge ( = aksapyge ) , pseudohysterolenus , pseudoyuepingia ( = iwayaspis ; = sayramaspis ) , sinoproceratopyge , tamdaspis ( = psiloyuepingia ) , tropidopyge , wannania , xiaodaositunia .
benxiella , changshania ( = metachangshania ; = prochangshania ) , changshanocephalus , kazelia ( = kazellina ) , mecophyrs , narinosa , parachangshania , paramenomonia , paraqingshuiheella ( = qingshuiheella ) , pseudowentsuia , suribongia , wentsuia .
aksayaspis , cheilocephalus ( = pseudolisania ; = zhalangtania ) , emsurella , lecanoaspis , macelloura , oligometopus ( = bernicella ) , parakoldinia , pseudokingstonia , pseudokoldinia .
acanthoparypha , actinopeltis hawle & corda , anasobella , ancyginaspis , apollonaspis , arcticeraurinella , areia , areiaspis , azyptyx , barrandeopeltis , borealaspis ( = alreboaspis ) , bornholmaspis , bufoceraurus , ceraurinella ( = bartoninus ) , ceraurinium , ceraurinus ( = remipyga ) , cerauromeros , cerauropeltis , ceraurus ( = eoceraurus ) , cheirurus , chiozoon , contracheirurus , courtessolium , crotalocephalides , crotalocephalina ( / gibbocephalus ; = mezocrotalus ) , crotalocephalus ( = cerauroides ; = pilletopeltis / boeckia pillet ) , cyrtometopella , cyrtometopus , deiphon , didrepanon , eccoptochile , eccoptochiloides , forteyops , foulonia , gabriceraurus , geracephalina , hadromeros , hammannopyge , hapsiceraurus , heliomera , heliomeroides , holia ( = ainoa ) , hyrokybe ( = shiqiania ) , junggarella , kawina ( = cydonocephalus ) , kolymella , krattaspis , ktenoura , laneites , lehua , leviceraurus , nieszkowskia , onycopyge , osekaspis , pandaspinapyga , paraceraurus , parasphaerexochus ( = mayopyge ) , parayoungia ( = ichiyamella ) , parisoceraurus , pateraspis , patomaspis , placoparina , pompeckia , proromma , protocerauroides , pseudocheirurus , pseudosphaerexochus ( = zethus ) , radiurus , ratinkaspis , reraspis , skelipyx , sphaerexochus ( = korolevium ; = onukia ; = parvixochus ) , sphaerocoryphe ( = ellipsocoryphe ; = hemisphaerocoryphe ) , stubblefieldia , sycophantia , turantyx , valongia , whittakerites , xylabion , xystocrania ( = xialiangshania ) , youngia , zazvorkaspis .
zhang & lin in w . zhang et al . , 1980a [ redlichiida redlichina redlichioidea ]
aragotus , bathynotus ( = pagura ) , bathynotellus , belliceps , chengkouaspis , elegestina , inella , pseudoresserops , terechtaspis ( = nellina ) .
zhu in w . zhang et al . , 1980a [ ptychopariida ptychopariina ? ellipsocephaloidea ]
acanthomicmacca ( = chengkouia ; = jaskovitchella ; = myopsomicmacca ) , bidjinella , changyangia , micmacca , turkestanella , wenganella , xiuqiella , zacanthellina , zhenbaspis ( = yankongia ; = zhenxiongaspis ) .
aspidagnostus ( = biragnostus ) , clavagnostus ( = tomorhachis ; = culipagnostus ; = stigmagnostus ; = acanthagnostus ; = leptagnostus ; = paraclavagnostus ) , triadaspis , utagnostus .
condylopyge ( / paragnostus ; = fallagnostus ) , miraculaspis , pleuroctenium ( = dichagnostus ) .
bailiaspis , bailiella ( = liaotungia ; = liocephalus ; = tangshihella ) , cainatops ( = cornucoryphe ) , conocoryphe ( / conocephalites ; = conocephalus ; = couloumania ) , ctenocephalus , elyx ( / eryx ) , hartella , parabailiella , tchaiaspis .
buitella , catuniella , conokephalina ( = lobocephalina ; = ruzickaia / lobocephalus ) , gorskia , maspakites , meisteraspis , meisterella , miranda , oirotella , suludella , westergaardella .
abakania , acontheus ( = aneucanthus ; = aneuacanthus ) , bonnaspis , chatiania ( = parachatiania ) , clavigellus , corynexochella , corynexochina , corynexochus ( = karlia ) , eochatiana , eocorynexochus , hartshillia , hartshillina , milaspis , miranella , olinaspis , sanaschtykgolia , shivelicus , trinia .
bagongshania , beikuangaspis , cayupania , coosella ( = wilsonella ) , coosia , coosina , coosinoides , crepicephalina ( = mesocrepicephalus ) , crepicephalus , hsuchuangia , idioura , kasatchaspis , neimonggolaspis , perforina , pseudocrepicephalus , sinocoosella , sinocrepicephalus , sneedvillia , temnoura ( = asteromajia ) , tetraceroura , uncaspis , zaozhuangaspis .
amicus , aspidaeglina , circulocrania , cyclopyge ( / egle / aeglina ) , degamella , ellipsotaphrus , emmrichops , gastropolus ( = lisogoraspis ) , girvanopyge ( = cremastoglottos ; = gamops ; = nanlingia ) , heterocyclopyge ( = selenoptychus ) , microparia ( = gallagnostoides ) , novakella ( = incisopyge ) , paramicroparia , phylacops , pricyclopyge ( = bicyclopyge ) , prospectatrix , psilacella , quadratapyge , sagavia , symphysops , waldminia , xenocyclopyge .
anchiopsis , andreaspis , argentopyge , banilatites , bessazoon , blanodalmanites , chacomurus , chattiaspis , coronura , corycephalus , crozonaspis , dalmanites ( / dalmania ; = guaranites ; = hausmannia etheridge & mitchell ; = heliocephalus / malvernia ; = makaspis ; = ommokris ) , dalmanitina , dalmanitoides , dalmaniturus , daytonia , deloites , delops , destombesites , dreyfussina ( = prephacopidella ) , duftonia , eodalmanitina , eudolatites , fenestraspis , forillonaria , francovichia , furacopyge , gamonedaspis , glyptambon , guichenia , huntoniatonia ( / huntonia ) , kasachstania , lygdozoon , malladaia , morgatia , mucronaspis ( = guaykinites ) , mytocephala ( = mirops ) , neoprobolium , odontocephalus , odontochile ( / hausmannia hall & clark ) , ormathops , pericopyge , phalangocephalus , preodontochile , prodontochile , prosocephalus , retamaspis , reussiana , roncellia , schoharia , songxites , struveria , synphoria ( / eocorycephalus ; / neosynphoria ) , synphoroides , thuringaspis , toletanaspis , trypaulites , vokovicia , zeliszkella , zlichovaspis ( = devonodontochile ; = spinodontochile ) .
? adelogonus , ariaspis , bergeronites ( = spinopanura ) , blackwelderia ( = parablackwelderia ) , blackwelderioides , chiawangella , cyrtoprora , damesella ( = haibowania ; = eodamesella ) , damesops ( = meringaspis ; = paradamesops ) , dipentaspis , dipyrgotes , drepanura , duamsannella , fengduia , guancenshania , ? hercantyx , histiomona , jiawangaspis , liuheaspis , metashantungia , neodamesella , palaeadotes ( = pseudobergeronites ) , paradamesella ( = falkopingia ) , parashantungia , pingquania ( = oxygonaspis ) , pionaspis , protaitzehoia , pseudoblackwelderia , shantungia , stephanocare , taihangshania , taitzehoia , teinistion ( = dorypygella ) , xintaia , yanshanopyge .
anopocodia , aulacodigma , cyclolorenzella , diceratocephalus , fenghuangella ( = cyclolorenzellina ) , hwangjuella , jiangnania , tangshihlingia , tholifrons ( = paraphoreotropsis ) , torifera , xiangia .
berkeia , blandicephalus , briscoia , camaraspoides , dikelocephalus , elkia , goumenzia , hoytaspis , iranella , kasachstanaspis , monocheilus , olimus , osceolia , parabriscoia , patalolaspis , princetonella ( / calyptomma ) , pterocephalops rasetti , randicephalus , stigmacephalus , walcottaspis .
celmus ( = crotalurus ; = ischyrophyma ) , dimeropyge ( / haploconus ) , dimeropygiella , glaphurella , ischyrotoma , pseudohystricurus .
amginoerbia , botomella ( = sayanella ) , chakasskia , chakasskiella , compsocephalus ( / lepidocephaloides ) , densocephalus , dilataspis , dinesus , erbia ( = paratollaspis ) , erbiella , erbina , erbiopsidella , erbiopsis , ghwaiella , paraerbia , piriforma , pokrovskiella , proerbia , pseudoerbia , pseudoerbiopsis , rondocephalus , tingyuania , tollaspis , tumulina .
aethedionide , digrypos , dionide ( / dione ; / polytomurus ; = dionidepyga ; = trigrypos ) , dionideina , dionidella , huangnigangia , paradionide , tongxinaspis , trinucleoides .
agnostotes , baltagnostus ( ? = trilagnostus ) , denagnostus , diplagnostus ( = enetagnostus ; = tasagnostus ) , dolichagnostus , iniospheniscus , linguagnostus ( = cristagnostus ) , machairagnostus , nahannagnostus , oedorhachis , oidalagnostus ( = ovalagnostus ) , pseudagnostus ( = litagnostus ; = plethagnostus ; = pseudagnostina ; = rhaptagnostus ; = sulcatagnostus ; = xestagnostus ) , pseudoglyptagnostus ( = glyptagnostotes ) , pseudorhaptagnostus ( = neoagnostus ; = euplethagnostus ; = hyperagnostus ; = tarayagnostus ; = calagnostus ) , trisulcagnostus ( = tririmagnostus ) .
acrocephalina , alekcinella , anemocephalus , anuloides , apachia ( = apachilites ) , bellaspidella , bellaspis , beothuckia , burnetiella ( / burnetia ) , calocephalites , chalfontia , conaspis , crusoiina , deckera , dellea ( = eshelmania ) , delleana , didwudina , dokimocephalus , fastigaspis , glyptometopsis , glyptometopus , iddingsia ( = plataspella ) , jingxiania , kindbladia , kiowaia , kyphocephalus , lorrettina , obrucheviaspis , pinctus , plakhinella , pseudosaratogia , puanella , ritella , saimachia , sulcocephalus , taenicephalina , tatulaspis , tchuostachia , whittingtonella , wilsonarella , wuhuia ( = deadwoodia ) , yangweizhouia .
dolerolenus ( / olenopsis ; = malungia ) , giordanella , granolenus , paramalungia .
aegunaspis amphoton ( = eurodeois ; = amphotonella ; = paramphoton ; = sunia ) , anoria , asperocare , athabaskia , athabaskiella , atypicus , basanellus , bathyuriscidella , bathyuriscus ( = orria ; = orriella ; = wenkchemnia ) , borovikovia , centonella , chilometopus , chilonorria , clavaspidella , corynexochides , deiradonyx , dolicholeptus , dolichometopsis , dolichometopus , drozdoviella , erratobalticus , ezhuangia , fuchouia ( = parafuchouia ; = pseudofuchouia ) , glossopleura ( = sonoraspis ) , ? granularaspis ( / granularia ) , guraspis , ? hanburia , hemirhodon , horonastes , itydeois , kannoriella , klotziella , lianhuashania , mendospidella , neopoliellina , parapoliella , poliella ( = bornemannia ) , poliellaspidella , poliellaspis , poliellina , politinella , polypleuraspis , prosymphysurus , pseudamphoton , ptarmigania , saimixiella , sestrostega , shanghaia , sinijanella , suvorovaaspis , undillia , zhenpingaspis .
atdabanella , basocephalus , bonnaria , bonnia , bonniella , bonnima , bonnioides , bonniopsis , dorypygaspis , dorypyge , dorypygina , dorypygoides , duyunia , fordaspis , hicksia , holteria , jiuquania , kharausnurica , kootenia ( = notasaphus ) , kooteniella ( = babakovia ) , kooteniellina , kootenina , liokootenia , mengzia , metakootenia , namiolenoides , neolenus , ogygopsis ( = taxioura ) , olenoides , paraolenoides , popigaia , prokootenia , protypus ( = bicaspis ) , pulvillaspis , rabutina , saryaspis , shipaiella , strettonia , tabatopygellina , tadjikia , tienzhuia , tolanaspis .
alacephalus , edelsteinaspis , gelasene , keeleaspis , labradoria ( = sinolenus ) , labradorina , laticephalus , litaspis , nehanniaspis , neoredlichina , nodiceps , paleofossus , polliaxis , torosus , venosus .
acadolenus , alueva , antatlasia , argunaspis , asiatella , bergeroniaspis , bergeroniellus , blayacina , brevitermierella ( = paratermierella ) , cambrunicornia , catadoxides , charaulaspis , chorbusulina , comluella , culmenaspis , ellipsocephalus ( = germaropyge ) , ellipsostrenua , glabrella , hamatolenus , hupeolenus , issafeniella , kadyella , kameschkoviella , kijanella , kingaspidoides ( = elatius ) , kingaspis ( = mesetaia ) , krolina , kymataspis , latikingaspis , latouchia , latuzella , lermontovia , limataceps , limouolenus , lotzeia , lusatiops ( = jalonella ) , mohicana , myopsolenus ( = collyrolenus ) , myopsostrenua , nelegeria , olekmaspis , ornamentaspis , orodes , ourikaia , paramicmacca , paraprotolenella , pauliceps , planolimbus , protagraulos , protaldonaia , protolenella , protolenus ( / bergeronia ; = matthewlenus ) , pruvostinoides , pseudoasiatella , pseudokadyella , pseudolenus , pseudoprotolenella , ptychoparopsis ( = berabichia ) , rinconia , sailycaspis , sectigena , strenuaeva ( = hindermeyeria ) , strenuella , tadakoustia , termieraspis , termierella , thoralaspis , timnaella , triangulaspis ( = acutaspis ; = angustaeva ; = plenudiscus ; = triangullina ) , yeshanaspis .
chariocephalus , dartonaspis , drumaspis , dunderbergia , dytremacephalus , elburgia , elvinaspis , elvinia ( = moosia ) , elviniella , elvinioides , elyaspis , enshia , irvingella ( = irvingellina ; = parairvingella ; = komaspis ) , jessievillia , kujandina , maladioides , maladiopsis , megadundabergia , metisaspina , onchopeltis , paraenshia , parakomaspis , pesaia , protemnites ( = prismenaspis ) , pseudomaladioides , pseudosaukia , qingshuihella , schmidtaspis , yunlingia .
aegrotocatellus , alwynulus , atractocybeloides , atractopyge ( = cybelella ) , avalanchurus , balizoma , batocara ( = pacificurus / australurus ) , bevanopsis , billevittia , brianurus ( / briania ) , celtencrinurus , coronaspis , coronocephalus ( = coronocephalina ; = senticucullus ) , cromus ( = encrinuraspis ) , curriella , cybele ( = cybelina ) , cybeloides , cybelurus ( = miracybele ) , dayongia , deacybele , dindymene ( = cornovica ) , distyrax , dnestrovites , elsarella , encrinuroides , encrinurus ( = saoria ) , eodindymene , erratencrinurus , fragiscutum , frammia , frencrinuroides , johntempleia , kailia , koksorenus , langgonia , lasaguaditas , libertella , lyrapyge , mackenziurus , mitchellaspis ( / mitchellia ) , nucleurus , oedicybele ( = dindymenella ; = jemtella ) , paracybeloides , paraencrinurus , parakailia , perirehaedulus , perryus , physemataspis , plasiaspis , prophysemataspis , prostrix , rongxiella , sinocybele , staurocephalus , stiktocybele , struszia , tewonia , walencrinuroides , wallacia .
dawsonia ( = aculeodiscus ; = metadiscus ) , eodiscus ( = spinodiscus ; = deltadiscus ) , helepagetia , kiskinella , macannaia , opsidiscus ( / aulacodiscus ) , pagetia ( = eopagetia ; = mesopagetia ) , pagetides ( = discomesites ) , sinopagetia .
alanisia , chulanolenus , coreolenus , eomalungia , estaingia ( = hsuaspis , = pseudichangia ; = zhuxiella ; = sematiscus ; = strenax ) , hupeia , ichangia , longmenshania , longxianaspis , madianaspis , mundocephalina , ningxiaspis , olekmanellus , paraichangia , pararaia ( = proichangia ; = tannuolaspis ) , protolenoides , shangsiaspis , shifangia , shiqihepsis , sichuanolenus , subeia , szechuanolenus , yinshanaspis .
acrocephalaspina , altaiaspis , ? amzasskiella ( = triplacephalus ) , archaeuloma , baikadamaspis , bilacunaspis , butyrinia , crucicephalus , dolgeuloma ( / rosovaspis / psedoacrocephalites rosova ) , duplora , euduplora , euloma ( = calymenopsis ) , guizhoucephalina , iveria , karataspis , ketyna ( = kujandaspis ) , lateuloma , limpeina , loparella , lopeuloma , luyanhaoia , miaeuloma , ? natmus , pareuloma ( = gansucephalina ) , pesaiina , plecteuloma , probilacunaspis , proteuloma ( = mioeuloma ) , pseudoacrocephalites maksimova , sanduspis , spineuloma , stigmatoa .
bandalaspis , bayfieldia , corbinia , eurekia , leocephalus , lochmanaspis , magnacephalus , maladia , tostonia .
amplifallotaspis , choubertella , daguinaspis ( = eodaguinaspis ; = epidaguinaspis ) , eofallotaspis , fallotaspis , lenallina , parafallotaspis , pelmanaspis , profallotaspis , repinaella , wolynaspis . [ profallotaspis & repinaella placed in archaeaspididae by some workers ]
bornemannaspis , gigantopygus , parayiliangella , pseudoyiliangella , yilliangella ( = palaeoaspis ) , yilliangellina , zhangshania .
arraphus , bohemoharpes ( = declivoharpes ; = unguloharpes ) , bowmania , brachyhipposiderus , conococheaguea , dolichoharpes , dubhglasina ( = australoharpes ; = sinoharpes ) , entomaspis ( = hypothetica ) , eoharpes ( / harpina ) , eotrinucleus , harpes ( = helioharpes ; = reticuloharpes ) , heterocaryon , hibbertia ( / platyharpes ; = harpesoides ; = metaharpes ; = paraharpes ; = thorslundops ; = wegelinia ) , kathrynia , kielania ( = lowtheria ) , lioharpes ( = fritchaspis ) , palaeoharpes , scotoharpes ( = aristoharpes ; = selenoharpes ) .
chencunia , dictyocephalites , fissocephalus , harpides , harpidoides , kitatella , loganopeltis , loganopeltoides , metaharpides , paraharpides , pscemiaspis .
delgadella ( = alemtejoia ; = delgadodiscus ; = delgadoia ; = pagetiellus ; = pentagonalia ) , dicerodiscus , hebediscus , luvsanodiscus , natalina ( = limbadiscus ) , neopagetina ( / pagetina ) , parapagetia ( = planodiscus ) , tchernyshevioides .
andalusiana , baltobergstroemia , callavia ( = cephalacanthus / callavalonia ; = cobboldus ) , cambropallas , elliptocephala ( / georgiellus , / ebenezeria ) , holmia ( = esmeraldina ) , holmiella , iyouella , kjerulfia , palmettaspis , postfallotaspis , schmidtiellus ( / schmidtia ) .
dasometopus , holocephalina ( = carausia ) , holocephalites , meneviella ( / menevia , / salteria / errinys ) , sdzuyella .
arduennella , brongniartella ( = pamirotellus ; = portaginus ) , burmeisterella , burmeisteria , digonus , dipleura , eohomalonotus ( / brongniartiasalter ) , homalonotus , huemacaspis , iberocoryphe , kerfornella , leiostegina , parahomalonotus , plaesiacomia , platycoryphe ( = liangshanaspis ) , scabrella , trimerus .
asaphopsoides ( = dainellicauda ; = xiangxiia ) , ciliocephalus , dactylocephalus , dikella , dikelocephalopsis , dikelokephalina , dikelus , hungaia ( = acrohybus ) , hungioides ( = argentinops ) , leimitzia , meitanopsis , songtaoia , warendia , xiushanopsis .
amblycranium , etheridgaspis , flectihystricurus , genalaticurus , glabretina , guizhouhystricurus , hillyardina ( = metabowmania ) , hintzecurus , ? holubaspis ( / holubia ) , hyperbolochilus , hystricurus ( = vermilionites ) , ibexicurus , lavadamia , nyaya , omuliovia , pachycranium , paenebeltella , parahystricurus , paraplethopeltis , politicurus , psalikilopsis , psalikilus , rollia , rossicurus , tanybregma , ? taoyuania ( = batyraspis ) , tasmanaspis , tersella .
aguilarella , arrhenaspis , brabbia , comanchia , duibianaspis , elviraspis , langyashania , lauzonella , levisella , loganellus ( = highgatea ) , maladioidella ( = kuruktagella ; = cedarellus ) , noelaspis , patronaspis , psalaspis , pyttstrigis , saratogia ( = idahoia ; = meeria ) , shitaia , valtoressia , wafangia , wilbernia , zhuitunia .
alloillaenus , bumastoides , dysplanus , ectillaenus ( = wossekia ) , harpillaenus , hyboaspis , illaenus ( / cryptonymus ; = actinolobus ; = deucalion ; = svobodapeltis ) , nanillaenus , ninglangia , octillaenus , ordosaspis , parillaenus , platillaenus , ptilillaenus , quadratillaenus , snajdria , spinillaenus , stenopareia , trigoncekovia , ulugtella , vysocania , wuchuanella , zbirovia , zdicella , zetillaenus .
ambonolium , illaenurus , lecanopyge , minicephalus , olenekella , platydiamesus , polyariella , rasettaspis , rasettia ( / platycolpus ) , resseraspis , tatonaspis yurakia .
catinouyia , eoinouyia , huainania , inouyia , parahuainania , parainouyia , parajialaopsis , parawuania , plesiowuania , proinouyia , pseudinouyia .
cyphoniscus , effnaspis , hanzhongaspis , holdenia ( / tiresias ) , isocolus ( / astyages ) , kielanella , liangshanocephalus , paratiresias , pradesia , pseudopetigurus , taimyraspis , thoralocolus , triarthroides .
argasalina , bathyuriscellus , bathyuriscopsis , daldynia , gibscherella , jakutus , janshinicus , jucundaspis , judaiella , kobdus , lenaspis , malykania , manaspis , prouktaspis , uktaspis , vologdinaspis .
anhuiaspis , ceronocare , donggouia , eokaolishania , eomansuyia , eotingocephalus , hapsidocare , hemikaolishania , kabutocrania , kaolishania , kaolishaniella , liaotropis , mansuyia , mansuyites ( = parapalacorona ) , ? mimana , palacorona , palemansuyia , parakaolishania , paramansuyella ( / paramansuyia ) , peichiashania , prolloydia , shidiania , taianocephalus , tangjiaella , tingocephalus , tugurellum , wayaonia .
acheilus clark , ankoura , blountia ( = homodictya ; = protillaenus ; = stenocombus ) , brachyaspidion ( / brachyaspis miller , 1936 ) , bynumia , bynumina , calvipelta , clelandia ( / harrisia ; = bynumiella ) , ithycephalus , kingstonella , kingstonia ( = ucebia ) , kingstonioides , komaspidella ( = buttsina ; = ataktaspis ) , larifugula , maryvillia , pugionicauda , saonella , shuizuia , wanwanaspis , wanwanoglobus , yanzhuangia .
aedotes , aethochuangia , agerina ( = otarionellina / otarionella ) , alloleiostegium , ampullatocephalina , annamitella ( = bathyuriscops ; = endoaspis / wutingia ; = proetiella ; = monella ) , aspidochuangia , baoshanaspis , brackebuschia ( = bodenbenderia ; = hexianella ) , cholopilus , chosenia ( = leiostegioides ) , chuangia ( / schantungia ; = parachuangia ; = pterochuangia ) , chuangiella , chuangina , chuangioides , chuangiopsis , chuangites , constrictella , eochuangia , euleiostegium , evansaspis , gonicheirurus , iranaspis , iranochuangia , jinanaspis , kepisis , leiostegium ( = endocrania ) , leptochuangia , linguchuangia , lloydia , madaoyuites , manitouella , marcouella , meropalla , paraaojia , paraleiostegium , paraonychopyge , paraszechuanella , perischodory , plethopeltella , pseudocalymene ( = eucalymene ) , pseudoleiostegium , reubenella , sailoma , shanchengziella , sobovaspis , szechuanella , tinaspis , xinhuangaspis , yaopuia , yarmakaspis , yinjiangia .
acidaspidella , acidaspides , acidaspidina , archikainella , belovia , bestjubella , brutonia , colossaspis , eoacidaspis , lichakephalus , lichokephalina , metaacidaspis , paraacidaspis , usoviana .
acanthopyge ( = euarges ) , akantharges , allolichas , amphilichas ( / paralichas / platymetopus ; = acrolichas ; = kerakephalichas ; = tetralichas ) , apatolichas , arctinurus ( / oncholichas ; / platynotus ; / pterolichas ) , autoloxolichas , borealarges , ceratarges ( / arges ) , ceratolichas , conolichas ( = cypholichas ) , craspedarges , dicranogmus , dicranopeltis ( = dicranopeltoides ; = nonix ; = raymondarges ; / trachylichas ; = tsunyilichas ) , echinolichas , eifliarges , gaspelichas , hemiarges ( = choneilobarges ) , homolichas , hoplolichas ( = cyranolichas ) , hoplolichoides , jasperia , leiolichas , lichas ( = apolichas ; = autolichas ) , lobopyge ( = belenopyge ) , lyralichas , mephiarges , metaleiolichas , metalichas , metopolichas ( / metopias ; = holoubkovia ; = macroterolichas ) , neolichas , nipponarges , ohleum , oinochoe , otarozoum , paraleiolichas , perunaspis ( = nitidulopyge ) , platylichas ( = lingucephalichas ) , probolichas , pseudotupolichas ( = arctinuroides ) , radiolichas ( = diplolichas ; = septidenta ) , richterarges , rontrippia , terataspis , terranovia , trimerolichas , trochurus ( = corydocephalus ; = plusiarges ; = makromuktis ) , uralichas ( = bohemolichas ; = platopolichas ) , uripes .
dazhuia , eoshengia ( = baojingia ) , extrania , klimaxocephalus , lisania ( = aojia ) , megalisania , metalisania , paralisaniella , paraojia , parashengia , platylisania , quandraspis , redlichaspis ( = lisaniella ) , rinella , shengia , xichuania .
amquia , arcuolimbus , deiracephalus ( / asteraspis ) , genevievella ( = placosema ; = nixonella ; = torridella ) , llanoaspidella , llanoaspis , metisaspis , nahannicephalus , paracedaria ( / pilgrimia ) , rogersvillia , sacha , stenelymus , tagenarella .
amiaspis , bolaspidellus , calymenidius , caulaspina , caulaspis , durinia , glaphyraspis ( = raaschella ) , graciella , hawkinsaspis ( / hawkinsia ) , interalia , kuraspis , kuraspoides , lazarenkiura , letniites , lonchocephalus ( = bucksella ) , monosulcatina , neoglaphyraspis , nordia , olegaspis , prolonchocephalus , pseudotalbotina , quebecaspis , raaschellina , talbotina , terranovella , trymataspis , weeksina .
damiaoaspis , eujinnania , inouyops , inoyellaspis , jiangjunshania , lonchinouyia , lorenzella , paralorenzangella ( / paralorenzella q . z . zhang ) , paralorenzella luo , paraporilorenzella , porilorenzella ( = jinnania ) , pseudolorenzella , ptyctolorenzella , zhongweia .
angsiduoa , hualongia , mapania . mapanopsis , metanomocarella , paramapania , pseudomapania , quitacetra , quitalia .
anemocephalops , crepichilella , glyphopeltis , holmdalia , ithyektyphus , lecanopleura , loulania , marjumia , modocia ( = armonia ; = metisia ; = perioura ; = semnocephalus ) , nasocephalus , nericella , nericia , pearylandia , petruninaspis , schyilaspis , shickshockia , syspacheilus .
balderia , biaverta , bolaspidella ( = deissella ; = howellaspis ) , bridgeia , coenaspis , coenaspoides , deltophthalmus , dresbachia , hysteropleura ( = apedopyanus ) , josina , knechtelia , menomonia ( = densonella / millardia ) , tavsenia , verditerrina .
churkinia , conomicmacca , enantiaspis , fuminaspis , hongshiyanaspis , metadoxides ( = anadoxides ) , minusinella , onaraspis , pratungusella .
arthrorhachis ( = metagnostus ; = girvanagnostus ) , chatkalagnostus ( = oculagnostus ) , corrugatagnostus ( = segmentagnostus ; = granulatagnostus ; = cenagnostus ) , diplorrhina ( = mesospheniscus ; = quadragnostus ; = pseudoperonopsis ) , dividuagnostus ( = pezizopsis ) , galbagnostus , geragnostella , geragnostus ( = geratrinodus ; = neptunagnostella ) , granuloagnostus , homagnostoides , novoagnostus , trinodus .
fuzhouwania , hardyia , lunacrania ( = paranumia ) , parakoldinioidia ( = macroculites ; = missisquoia ; = rhamphopyge ; = tangshanaspis ) , pseudokoldinioidia , tasmanocephalus .
jinxiaspis , liaoningaspis , liaoningella , metalioparella , monkaspis ( = kushanopyge ; = paraliaoningaspis ) , nomadinis , proliaoningaspis , walcottaspidella .
chondrinouyina , erkelina , inouyina , juliaspis , kassinius , namanoia , sinoschistometopus , tarynaspis .
buenaspis , corcorania , liwia ( / livia ) , misszhouia , naraoia , soomaspis , tarricoia , tegopelte .
ascionepea , ferenepea , folliceps , loxonepea , nepea , penarosa ( = trinepea ) .
buenellus , cambroinyoella , cirquella , limniphacos , nevadella , nevadia , plesionevadia , pseudojudomia , sdzuyomia .
aocaspis , barrandia , berkutaspis , borthaspidella , bumastides , elongatanileus , homalopteon , illaenopsis ( = eurymetopus ; = procephalops ; = rokycania / pseudobarrandia ) , kodymaspis , ? lakaspis , neopsilocephalina , nileus ( = remopleuridioides ) , parabarrandia , parabumastides , paranileus , peraspis , petrbokia , platypeltoides ( / platypeltis ) , poronileus , psilocephalinella ( / psilocephalus / psilocephalina / borthaspis ) , shenjiawania , symphyroxochus , ? symphysurina ( = symphysurinella ; = symphysuroides ) , symphysurus , troedssonia , varvia .
cedaraspis , garbiella , hardyoides ( = norwoodina ) , holcacephalus , levisaspis , norwoodella , norwoodia ( = whitfieldina ) , paranorwoodia , xenocheilos .
acanthalomina , acidaspis , anacaenaspis ( = bruxaspis ) , apianurus , archaeopleura , boedaspis , borkopleura , brutonaspis , calipernurus , ceratocara , ceratocephala ( = bounyongia ; = onchaspis ; / trapelocera ) , ceratocephalina , ceratonurus , chlustinia , dalaspis , diacanthaspis , dicranurus , dudleyaspis , edgecombeaspis , eoleonaspis ( = bojokoralaspis ) , exallaspis , gaotania , globulaspis , hispaniaspis , isoprusia ( = mauraspis ) , ivanopleura , kettneraspis ( = grossia ) , koneprusia , laethoprusia , leonaspis ( = acanthaloma ) , meadowtownella , miraspis ( = elbaspis ) , ningnanaspis , odontopleura , orphanaspis , periallaspis , primaspis , proceratocephala ( = drummuckaspis ) , radiaspis ( = xanionurus ; = charybdaspis ) , rinconaspis , selenopeltis ( = languedopeltis ; = polyeres ) , selenopeltoides , sinespinaspis , stelckaspis , taemasaspis ( = gondwanaspis ; = snoderaspis ) , uriarra , whittingtonia .
angustolenellus , arcuolenellus , biceratops , bolbolenellus , bristolia , fremontella , fritzolenellus , gabriellus , laudonia , lochmanolenellus , mesolenellus , mesonacis ( = fremontia ) , mummaspis , nephrolenellus , olenelloides , olenellus ( / barrandia ) , paedeumias , peachella , teresellus , wanneria .
acerocare , acerocarina ( / cyclognathus ) , aciculolenus , anaximander , angelina ( = keidelaspis ) , apoplanias , asilluchus , baikonuraspis , balnibarbi , bienvillia ( = diatemnus ; = mendoparabolina ) , boeckaspis ( / boeckia brögger ; = sphaerophthalmella ) , bondarevites , bulbolenus , chekiangaspis , cloacaspis , ctenopyge , cyclognathina , danarcus , desmetia , eoctenopyge , euonchonotina , eurycare , granitzia , hancrania , helieranella , highgatella , huangshiaspis , hunanolenus , hypermecaspis ( = spitsbergaspis ) , inkouia ( = agalatus ) , isidrella , jujuyaspis ( = alimbetaspis ) , leiobienvillia , leptoplastides ( = andesaspis ; = beltella ; = chunkingaspis ; = parabolinopsis ; = rampartaspis ) , leptoplastus , leurostega , magnomma , mesoctenopyge , moxomia , neoolenus , neoparabolina , nericiaspis , olenus ( = simulolenus ) , orkekeia , parabolina ( / odontopyge ) , parabolinella , parabolinina , parabolinites , paraolenus , paraplicatolina , peltocare , peltura ( / anthes ; = anopocare ) , pelturina , plicatolina , plicatolinella , porterfieldia , prohedinella , protopeltura , psilocara , remizites , rhodonaspis , saltaspis , shihuigouia , sphaerophthalmus , svalbardites , talbotinella , triarthrus ( / brongniartia eaton ) , ? ullaspis , westergaardia ( = sphaerophthalmoides ) , westergaardites , wujiajiania .
delinghaspis , kontrastina , nidanshania , ordosia , paralevisia , plesioinouyella , poshania , pseudotaitzuia , taitzuia , taitzuina , tylotaitzuia , wanshania , xundiania .
arthricocephalus ( = arthricocephalites ; = protoryctocara ; = oryctocarella ) , balangia , barklyella , cheiruroides ( = inikanella ) , curvoryctocephalus , duodingia , duyunaspis , eoryctocephalus , euarthricocephalus , feilongshania , haliplanktos , hunanocephalus , kunshanaspis , lancastria ( = changaspis ; = chienaspis ; = goldfieldia ; = paraoryctocephalops ; = pseudolancastria ) , metabalangia , metarthricocephalus , microryctocara , neocheiruroides , opsiosoryctocephalus , oryctocara , oryctocephalina , oryctocephalites , oryctocephaloides , oryctocephalops , oryctocephalus ( = vinakainella ) , oryctometopus , ovatoryctocara , paleooryctocephalus , parachangaspis , paracheiruroides , protoryctocephalus , sandoveria , shabaella , taijiangocephalus , teljanzella , tonkinella , udjanella .
arcifimbria , bienella , datsonia , girandia , idamea , lichengaspis , lotosoides , oreadella , pagodia , pagodioides , phoreotropis , prochuangia , ptychopleurites ( / ptychopleura ; = aposolenopleura ; = punctularia ) , sagitaspis , sagitoides , seletoides , wittekindtia .
alataurus , bajangoliaspis , enammocephalus , ferralsia , gigoutella , habrocephalus , hoffetella , latipalaeolenus , megapalaeolenus , palaeolenella , palaeolenides , palaeolenus , resimopsis , schistocephalus , torgaschina , ulakhanella , validaspis .
hemibarrandia ( / pseudonileus kobayashi , 1951b ) , ottenbyaspis , panderia ( / rhodope ) , pogrebovites .
badainjaranaspis , metapianaspis , papyriaspis , pianaspis , prohedinaspis , prohedinia ( = tosotychia ) , sanduhedinaspis , wandelella , wangcunia , wudangia .
abdulinaspis , apomodocia , boestrupia , croixana , jasmundia , kendallina ( / kendallia ) , minkella , orygmaspis , parabolinoides ( = bernia ) , pedinocephalus , pesaiella , roksaspis , stigmacephaloides , taenicephalops , taenicephalus ( = bemaspis ; = maustonia ) , weishania .
acadoparadoxides ( / entomolithus / entomostracites ; = eoparadoxides ) , anabaraceps , anabaraspis , bajanaspis , baltoparadoxides , phanoptes ( = eccaparadoxides ; = macrocerca ) , hydrocephalus ( = phlysacium ; = rejkocephalus ) , paradoxides ( = vinicella ) , plutonides ( / plutonia ) , primoriella , schagonaria , schoriina .
archaeagnostus , cotalagnostus , eoagnostus , gratagnostus , hypagnostus ( = spinagnostus ; = cyclopagnostus ; = breviagnostus ; = metahypagnostus ) , lisogoragnostus ( = abagnostus ; = scanagnostus ) , micagnostus , peronopsella , peronopsis ( / mesagnostus ; = euagnostus ; = acadagnostus ; = axagnostus ; = itagnostus ) , sphaeragnostus , svenax .
acernaspis ( = eskaspis ; = otadenus ; = murphycops ) , acuticryphops , adastocephalum , afrops , ainasuella , altaesajania , ananaspis , angulophacops , arduennops , atopophacops , babinops , boeckops , burtonops , chotecops ( = cordapeltis ) , cryphops ( / gortania / microphthalmus ) , cultrops , denckmannites ( / denckmannia ) , dianops , dienstina , drotops , ductina , echidnops , echinophacops , eldredgeops , eocryphops , eophacops ( / pterygometopidella ; = bullicephalus ) , geesops , hypsipariops , illaenula , kainops , liolophops , lochkovella , nandanaspis , nephranomma , nephranops , nyterops , omegops , orygmatos , paciphacops , pedinopariops , phacopidella ( / glockeria ) , phacops , plagiolaria , portlockia , prokops , rabienops , reedops , rhinophacops , rhinoreedops , sambremeuaspis , signatops , somatrikelon , spinicryphops , struveaspis , struveops , tangbailaspis , teichertops , toxophacops , trimerocephalus ( / eutrimerocephalus ) , viaphacops , weyerites , zaplaops , zhusilengops .
eopharostoma , paivinia , pharostomina ( = colpocoryphoides ) , prionocheilus ( = pharostoma ) , ptychometopus , thulincola ( = pharostomaspis ) , xuanenia .
ludvigsen & westrop in ludvigsen et al . , 1989 [ ptychopariida ptychopariina ptychoparioidea ]
aphelotoxon ( = ponumia ) , cliffia , drabia , phylacterus ( = liostracinoides ) , westonaspis .
anacheiruraspis , anacheirurus , chashania , demeterops , emsurina , koraipsis , landyia , macrogrammus , metapilekia , metapliomerops , parapilekia , pilekia , pliomeroides , pseudopliomera , seisonia , sinoparapilekia , victorispina .
arapahoia ( = hesperaspis ) , exigua ( = brassicicephalus ) , koldinia , koldiniella , kuljumbina , leiocoryphe , meniscocoryphe , notaiella , plethopeltides , plethopeltis ( = plethometopus ; = enontioura ) , plethopeltoides ( = kulyumbopeltis ) , poriplethopeltis , pseudokoldinella , samgonus ( = lampropeltis ; = lampropeltastes ) , semicyclocephalus , stenopilus , strotocephala , tolstotchichaspis .
anapliomera , benedettia , canningella , colobinion , coplacoparia , cybelopsis , ectenonotus , encrinurella , evropeites , gogoella , guizhoupliomerops , hawleia , hintzeia , humaencrinuroides , ibexaspis , josephulus , kanoshia , leiostrototropis , liexiaspis , ngaricephalus , obliteraspis , ovalocephalus ( = hammatocnemis ; = paratzuchiatocnemis ) , parahawleia , parapliomera , perissopliomera , placoparia , pliomera ( / amphion ) , pliomerella , pliomeridius , pliomerina ( / pliomeraspis ) , pliomerops , protoencrinurella , protopliomerella , protopliomerops ( = stototropis ) , pseudocybele , pseudomera , quinquecosta , rossaspis , strotactinus , tesselacauda , tienshihfuia , tzuchiatocnemis .
acanthocephalus alomataspis , chelidonocephalus , dananzhuangaspis , daopingia , deltocephalus , derikaspis , dignaceps , eoasaphiscellus ( / eoasaphiscus lu & yuan ) , eymekops ( = kolpura ) , farsia , gangdeeria , gloria , grandioculus ( / megalophthalmus ; = honania ) , guankouia , hadraspis , hatangia , heukkyoella , holanshania , honanaspis , hsiaoshia , huaibeia , hundwarella ( = anomocaraspis ) , hunjiangaspis , iranochresterius , iranoleesia ( / irania ; = michaspis ; = heyelingella ) , itcheriella , jiangsuaspis , jiangsucephalus , jiangsuia , jiubaspis , jixianella ( / jixiania ) , koptura ( = parakoptura ) , kopungiella , kutsingocephalus , leichuangia , lioparella ( = zhuozishania ) , luliangshanaspis , luonanocephalus , manchuriella , maotunia , memmatella , miaobanpoia , paofeniellus , paragangdeeria , parazhongtiaoshanaspis , plectrocrania , plesigangdeeria , proasaphiscina , proasaphiscus , pseudanomocarina , pseudonericella , raduginella , severina , shanxiella , shuangshania , stella , sudanomocarina , szeaspis ( = spitiaspis ) , tankhella , tengfengia , teratokoptura , tylotaspis , ulania , urjungaspis , wutaishanaspis , xiangshania , xinglongia , yanshania , yujinia , zhaishania , zhongtiaoshanaspis .
? anecocephalus , beigongia , bromella , camaraspis , cernuolimbus , cilia , dikelocephalioides , dikelocephalites , dingxiangaspis , flabellocephalus , guangxiaspis , housia ( = housiella ) , jubileia , labiostrella , labiostrina ( = abia ) , longlingaspis , lunacephalus , luotuolingia , morosa , parahousia , paramaladioidella , pauciella , pedinaspis , pedinocephalites , pelicephalus , petalocephalus , plesiocilia , prehousia , proapatokephaloides , prodikelocephalites , pterocephalia ( / pterocephalus ; = hederacauda ) , pterocephalina , pterocephalopsinus ( / pterocephalops lin & zhang in zhu et al . , 1979 ) , pulchricapitus ( = reaganaspis ) , qilianaspis , sigmocheilus , stenambon , strigambitus , tiantouzhania , tumicephalus , uxunella , xiaoshiella , yingziaspis , yokusenia , yushugouia , zhenania , zhuangliella .
achatella , bolbochasmops , calliops , calyptaulax ( = ligometopus ; = homalops ) , carinopyge , ceratevenkaspis , chasmops , denella , elasmaspis , eomonorachus , estoniops , evenkaspis , ingriops , isalaux , isalauxina , keilapyge , liocnemis , monorakos , oculichasmops , oelandiops , parevenkaspis , podowrinella , pterygometopus , rollmops , ruegenometopus , sceptaspis , schmidtops , scopelochasmops , toxochasmops , tricopelta , truncatometopus , upplandiops , uralops , valdariops , vironiaspis , volkops , yanhaoia .
allobodochus , criotypus , goniagnostus , lejopyge ( / miagnostus ) , myrmecomimus , onymagnostus ( = agnostonymus ) , pseudophalacroma , ptychagnostus , ( = triplagnostus ; = huarpagnostus ; = solenagnostus ; = pentagnostus ; = aristarius ; = aotagnostus ; = acidusus ; = canotagnostus ; = zeteagnostus ) , schismagnostus , tomagnostella , tomagnostus .
alborsella , changia ( = coreanocephalus ; = quadraticephalus ; = fengshania ) , eoptychaspis , eowuhuia , euptychaspis , idiomesus , kathleenella , keithia , keithiella , macronoda ( = promesus ) , plectrella , proricephalus , ptychaspis ( = asioptychaspis ) , saukioides ( / pseudosaukia ; / jeholaspis ) , sunwaptia , wilcoxaspis .
achlysopsis , altikolia , altitudella , amecephalites , asthenopsis , austinvillia , balangcunaspis , bashania , bathyholcus , bathyocos , billingsaspis , binodaspis ( = xilingxia ) , blairella , bolaspidina , brunswickia , bulkuraspis , caborcella , callidaspina , callidaspis , cathayanella , champlainia , chengshanaspis , chinghisicus , chunghwaella , conopolus , dananzhuangia , deltina , douposiella ( = tongshania ) , elrathina , entsyna , eodouposiella , eokochaspis , eosoptychoparia , eospencia , eurostina , finecrestia , gaotanaspis , gaphuraspis , gedongaspis , gunnia ( = ellotia ; = yiliangaspis ) , hadrocephalites , hadrokraspedon , hamptonella , hejinaspis , hemicricometopus , hewenia , holasaphus , horbusonia , illtydaspis , jangudaspis , jialaopsis , jianchangia , jimaoshania , jiumenia , kermanella , kochaspis ( / palaeocrepicephalus ) , kochiella ( = eiffelaspis ) , kochiellina , kochina , kounamkites , kunmingaspis ( = benxiaspis ) , laminurus , laoyingshania , lianglangshania , loriella , luguoia , luxella , lyriaspis , majiangia , manailina , meitania , metisella , mexicella , monanocephalus , mopanshania , mrassina , mufushania , nangaocephalus , nangaoia , nangaops , nanoqia , nassovia , nelsonia , neokochina , nyella , olenekina , onchocephalites , ontoella , orienturus , orlovia , orloviella , pachyaspidella , pachyaspis , palmeraspis , panacus , paraeosoptychoparia , paragunnia , paramecephalus ( = parahiolites ) , paraperiomma , paraplagiura , parapoulsenia , parashuiyuella , paraziboaspis , perimetopus , periommella , piazella , pingluaspis , piochaspis , plagiura ( = ruichengella ; = plagiurella ) , pokrovskayaspis , poulsenella , poulsenia , poulseniella , probowmania , probowmaniella ( = proshantungaspis ) , probowmanops , proliostracus , promeitania , ? protohedinia , pseudoliostracus , psilostracus , ptychoparella ( = eoptychoparia ; = syspacephalus ; = elrathina ) , ptychoparia ( = agraulopsis ; = ptychoparioides ) , qiaotouaspis , qingshuiheia , reedus , regina , regius , runnania , salankanaspis , sanhuangshania , sanwania , schistometopus , semisphaerocephalus , seriaspis , shanganella , shantungaspis , shuiyuella , sinoptychoparia , spencella , spencia ( = stauroholcus ) , stoecklinia , sujaraspis , suluktella , taijiangia , taniaspidella , ? townleyella , trachyostracus , trigonyangaspis ( / trigonaspis ) , tukalandaspis , ? ulrichaspis , variopelta , vermontella , vica , volocephalina , wanhuaia , weijiaspis , wuhaina , xiangshanaspis , xingrenaspis ( = spitella ; = danzhaina ; = wuxunaspis ) , yaoyiayuella , yohoaspis , yuknessaspis , ziboaspidella , ziboaspis .
ampyx ( / brachyampyx ) , ampyxella , ampyxina , ampyxinella , ampyxoides , anisonotella , bulbaspis , caganaspis , carinocranium , cerampyx , cnemidopyge , collis , edmundsonia , ellsaspis , endymionia ( / endymion ) , globampyx , jiuxiella ( = miboshania ) , kanlingia , lonchodomas , malinaspis , malongullia ( = ampyxinops ) , mendolaspis , metalonchodomas , miaopopsis , nambeetella , nanshanaspis , parabulbaspis , parampyx , pseudampyxina , pytine , raphioampyx , raphiophorus , raymondella ( / reedaspis ) , rhombampyx , salteria , sinampyxina , sinoluia , taklamakania ( = xinjiangia ) .
breviredlichia , chaoaspis , chengjiangaspis , conoredlichia , elganellus , eoredlichia ( = archaeops ; = saukiandops ; = galloredlichia ; = pararedlichia ) , hesa , iglesiella , irgitkhemia , jingyangia , kepingaspis , kuanyangia , latiredlichia , lemdadella , leptoredlichia ( = paraleptoredlichia ) , maopingaspis , metaredlichia , mianxianella , nebidella , neoredlichia , ningqiangaspis , olgaspis , pachyredlichia , parawutingaspis , parazhenbaspis , pseudoredlichia , pseudowutingaspis , pteroredlichia ( = spinoredlichia ) , redlichia ( / hoeferia ; = mesodema ; = dongshania ) , redlichops , sapushania , sarassina , sardaspis , sardoredlichia , syndianella , tolbinella , ushbaspis ( = metaredlichioides ) , wengangaspis , wutingaspis , xela , xenoredlichia , yorkella , zhanglouia .
aktugaiella , amphitryon ( / caphyra ; = brachypleura ) , aotiaspis , apatokephalina , apatokephaloides , apatokephalops ( = aristokainella ; = wanliangtingia ) , apatokephalus , apiflabellum , arator , artokephalus , atratebia , auricula , binervus , blosyropsis , cavia , deanokephalus , diplapatokephalus , dislobosaspis , eoapatokephalus , eorobergia , haniwa , hastiremopleurides , hexacopyge , hualongella , hukasawaia , hypodicranotus , ivshinaspis , jiia , jingheella , jinshaella , kainella , kainellina , kainelloides , lacorsalina , lingukainella , lohanpopsis , loshanella , lulongia , makbelaspis , mendosina , menoparia , naustia , oculeus , poletaevia , portentosus , praepatokephalus , proapatokephalops , pseudokainella ( = elkanaspis ; = parakainella ; = fatocephalus ) , pugilator , remopleurella , remopleurides , remopleuridiella , richardsonaspis , richardsonella ( = lakella ; = protapatokephalus ) , robergia , robergiella , scinocephalus , sculptaspis , sculptella , sigmakainella , spinacephalus , taishania , teratorhynchus , tibikephalus , tramoria , yosimuraspis ( = eoyosimuraspis ; = metayosimuraspis ) .
cyamella ( / cyamops ; = paracyamella ) , hanjiangaspis , isbergia , madygenia ( = pseudobirmanites ) , protarchaeogonus , rorringtonia ( = analocaspis ; = chenaspis ; = trigonoproetus ) , solariproetus .
australaspis , clariondia , despujolsia , dolerolichia , eops , ezhimia , longianda , pareops , perrector ( = rawops ) , planocephalus , pseudosaukianda , realaspis , resserops , richterops ( = marsaisia ) , saukianda .
anderssonella ( / anderssonia , / sunina ) , calvinella , caznaia , danzhaisaukia , diemanosaukia , eosaukia ( = scolosaukia ) , galerosaukia , guangnania , hamashania , lichengia , linguisaukia , liquania , lophoholcus , lophosaukia , mareda , metacalvinella , mictosaukia , mictosaukioidia , paracalvinella , pileaspis , platysaukia , prosaukia ( = stenosaukia ) , pseudocalvinella , saukia , saukiella , sinosaukia , stigmaspis , tellerina , thailandium , wedekindiaspis ( / wedekindia ) .
acanthopleurella , changchowilla , conophrys , elaphraella , eoshumardia , gaoloupingia , koldinioidia ( = akoldinioidia ) , kuruktagaspis , kweichowilla , leioshumardia , leoforteyia , liriamnica , parashumardia , puanocephalus , shumardia , shumardoella ( / shumardella ) , shumardops , stigmametopus , tarimella ( = yinganaspis ) , thomondia , trianguraspis .
abakanopleura , acanthometopus , acrocephalaspis , aiaiaspis , aidarella , aikhaliella , albansia , aldanaspis , badulesia , bigranulella , bijelina , braintreella , canotaspis , catasolenopleura , changqingia ( = austrosinia ) , colliceps , conicephalus , crusoia , daldynaspis , datongites , denaspis , eilura , ejinaspis , erratojincella , foveatella , gonzaloia , gushanaspis , herse ( / solenopleurina ; = parasolenopleura westergard ) , huzhuia , hyperoparia , jiagouia , jincella , kabuqiia , kaipingella , karagandoides , keguqinia , lashushania , levisia , lingyuanaspis , liosolenopleura , maiaspis , manublesia , markhaspis , mataninella , menocephalites ( = parataitzuia ) , minupeltis , mukrania , munija , neoacrocephalites , neosolenopleurella , nilegna , ninaspis , notocoryphe , paracrocephalites ( / arctaspis ) , paramenocephalites ( = solenoparina ) , parasolenoparia , parasolopleurena ( / parasolenopleura poletaeva ) , parayabeia , pardailhania , perneraspis ( / perneria ) , pingluia , plesisolenoparia , proavus , protrachoparia , pseudosolenoparia , reillopleura , rimouskia , rina , sao ( = acanthocnemis ; = acanthogramma , = crithias ; = endogramma ; = enneacnemis ; = goniacanthus ; = micropyge ; = monadina ; = monadella ; = selenosema ; = staurogmus ; = tetracnemis ) , shanghaiaspis , sohopleura , solenoparia , solenoparops , solenopleura , solenopleurella , solenopleuropsis , squarrosoella , suyougouia , tabalqueia , tangwangzhaia , tjungiella , trachoparia , usumunaspis , velieuxia , wafangdiania , xianfengia , yabeia .
alceste , altaepeltis , amphoriops , ancyropyge , andegavia ( = sagittapeltis ) , arctipeltis , australoscutellum , avascutellum ( = ctenoscutellum ; = rutoscutellum ) , bojoscutellum ( = holomeris ) , boreoscutellum , breviscutellum , bronteopsis ( = homoglossa ) , brontocephalina , brontocephalus , bubupeltina , bumastella , bumastus , calycoscutellum , cavetia , cekovia , chichikaspis , chugaevia , ciliscutellum , cornuscutellum , craigheadia , cybantyx , decoroscutellum , delgadoa , dentaloscutellum , dulanaspis , ekwanoscutellum , eobronteus , eokosovopeltis ( = heptabronteus ) , eoscutellum , exastipyx , excetra , failleana ( = opsypharus ) , flexiscutellum , goldillaenoides , goldillaenus , hallanta , hidascutellum , illaenoides , illaenoscutellum , izarnia , japonoscutellum , kirkdomina , kobayashipeltis , kolihapeltis , kosovopeltis ( = heptabronteus ) , kotysopeltis , lamproscutellum , leioscutellum , ligiscus , liolalax ( / lalax ) , litotix , meitanillaenus , meridioscutellum , meroperix , metascutellum , microscutellum , mulciberaspis , neoscutellum , octobronteus ( = stoermeraspis / stoermeria ) , opoa , ottoaspis , paracybantyx , paralejurus , paraphillipsinella ( / phillipsella ; = protophillipsinella ) , perischoclonus , phillipsinella , planiscutellum ( = protoscutellum ) , platyscutellum , poroscutellum , protobronteus , protostygina , pseudoeobronteus , pseudostygina , quyuania , radioscutellum , raymondaspis ( / warburgella raymond ) , rhaxeros ( / rhax ) , sangzhiscutellum , scabriscutellum ( / dicranactis ) , scutellum ( = bronteus / brontes ; = goldfussia / brontes ; = goldius ) , septimopeltis , spiniscutellum , stygina , styginella , tenuipeltis , thaleops ( = hydrolaenus ) , theamataspis , thomastus , thysanopeltella , thysanopeltis , tosacephalus , turgicephalus , unicapeltis , uraloscutellum , waisfeldaspis , weberopeltis , xyoeax .
asaphellina , asaphopsis , omeipsis , pacootella , renhuaia , taihungshania ( = miquelina ) , tungtzuella .
carolinites ( = dimastocephalus ; = keidelia ; = tafnaspis ) , fialoides , goniophrys , oopsites , opipeuterella ( = ompheter ; = opipeuter ) , paraphorocephala , phorocephala ( = carrickia ) , ? pyraustocranium , telephina ( / telephus ) , telephops .
erediaspis , meteoraspis ( = greylockia ; = coleopachys ) , prometeoraspis , tricrepicephalus ( / paracrepicephalus ) .
anebolithus , australomyttonia , bancroftolithus , bergamia ( = bohemaspis ; = brandysops ; = cochliorrhoe ) , bettonolithus , botrioides , broeggerolithus ( = ulricholithus ) , costonia , cryptolithoides , cryptolithus , deanaspis , declivolithus , decordinaspis , eirelithus , famatinolithus , furcalithus , guandacolithus , gymnostomix , hanchungolithus ( = ichangolithus ; = yinjiangolithus ) , huenickenolithus , incaia , jianxilithus , lloydolithus , lordshillia , marekolithus , marrolithoides , marrolithus , ? microdiscus , myinda , myindella , myttonia , nankinolithus , ningkianolithus ( = ceratolithus ; = hexianolithus ) , novaspis , onnia , paratretaspis , paratrinucleus , parkesolithus , pragolithus , protoincaia , protolloydolithus , reedolithus , reuscholithus , salterolithus ( = smeathenia ) , stapeleyella , telaeomarrolithus , tetrapsellium , tretaspis , trinucleus ( / edgellia ) , whittardolithus , xiushuilithus , yinpanolithus .
alberticoryphe , astycoryphe , bojocoryphe , buchiproetus , centriproetus , cornuproetus , cyrtosymboloides , dalarnepeltis , dalejeproetus , decoroproetus ( ogmocnemis ; = proetidella , warburgaspis ) , denemarkia , diademaproetus , dipharangus , eopiriproetus , erbenicoryphe , eremiproetus ( = natatoraspis ; = dufresnoyiproetus ) , gracilocoryphe , gruetia , guilinaspis , hollardia , ignoproetus , interproetus , koneprusites , krohbole , lardeuxia , laticoryphe , lepidoproetus , linguaproetus , lodenicia , longicoryphe , macroblepharum , miriproetus , nagaproetus , paraeremiproetus , paralardeuxia , paralepidoproetus , parvigena , perakaspis , phaetonellus , phaseolops , piriproetoides , piriproetus , pribylia , prionopeltis ( / phaetonides / phaeton ) , prodrevermannia , proetina , proetopeltis , pterocoryphe , pteroparia , quadratoproetus , rabuloproetus , ranunculoproetus , remacutanger , richteraspis , rokycanocoryphe , sculptoproetus , slimanella , spinoproetus , stenoblepharum ( = viruanaspis ) , tafilaltaspis , tropicoryphe , tropidocoryphe , vicinoproetus ( / vicinopeltis ) , voigtaspis , wolayella , xiphogonium ( = trautensteinproetus ) , zetaproetus .
blandiaspis , dictyella , esseigania , guluheia , jiwangshania , leiaspis , lonchopygella , paradictyites , shergoldia , taipakia , tsinania ( = dictyites , / dictya ) , zhujia .
s . zhang in w . zhang et al . , 1980a [ agnostida eodiscina eodiscoidea ]
tsunyidiscus ( = mianxiandiscus ; = liangshandiscus ; = emeidiscus ; = hupeidiscus ; = shizhudiscus ; = guizhoudiscus ) .
amginouyia , antagmopleura ( = poljakovia ) , chondragraulina , chondagraulos , okunevaella , utia .
abakolia ( = costadiscus ) , acidiscus , acimetopus , analox , bathydiscus , bolboparia , cephalopyge , cobboldites , leptochilodiscus ( = kerberodiscus ) , litometopus , mallagnostus ( = ladadiscus ; ? = jinghediscus ) , meniscuchus , ninadiscus , oodiscus , runcinodiscus , semadiscus , serrodiscus ( = paradiscus ) , stigmadiscus , tannudiscus , weymouthia .
eotaitzuia , houmaia , inouyella , jixianaspis , latilorenzella ( = wuania ) , lophodesella , megagraulos , pseudosolenopleura , rencunia , ruichengaspis , wuanoides , wuhushania , yunmengshania .
drepanopyge , drepanuroides ( = xishuiella ) , hongjunshaoia , longduia , meitanella , paokannia , parapaokannia , parayinites , pseudopaokannia , qingkouia ( = paradrepanuroides ) , yinites , yunnanaspidella , yunnanaspis .
alaskadiscus , egyngolia ( = mongolodiscus ) , ekwipagetia , hebediscina ( = szechuanaspis ; = zhenbadiscus ) , lenadiscus , yukonia , yukonides .
elicicola , luaspis , pensacola , wangzishia , wenganlenus , yunnanocephalus ( / pseudoptychoparia ) .
agnostina ( family uncertain ) agnostogonus , armagnostus , blystagnostus , delagnostus , dignagnostus , gallagnostus , glaberagnostus ( = toragnostus ) , grandagnostus , hastagnostus , leiagnostus ( = phoidagnostoides ; = ciceragnostus ) , megagnostus , peratagnostus ( = monaxagnostus ) , phaldagnostus , phoidagnostus , plurinodus , skryjagnostus , valenagnostus .
redlichina ( family uncertain ) akbashichia , fandianaspis , iolgia , micangshania , xingzishania .
family uncertain aethia , agyrenella , aligerites , altiplanelaspis , amechilus , araeocephalus , arglina , aulacopleurina , bifodina , bijaspis , carmon , cayastaia , chishanheella , costapyge , crossoura , curiaspis , deltacare , dipharus , discagnostus , dolgaiella , entsyna , eoampyx , eocheirurus , eulomella , fabulaspis , faciura , fengtienia , gdowia , gladiatoria , gonioteloides , goycoia , hospes , hybocephalus , idiorhapha , irvingelloides , ishpella , isidreana , jonotus , juriietella , kirengina , kleptothule , loxoparia , loxopeltis , lusampa , macnairides , maximovella , metaprodamesella , muchattellina , nanshihmenia , neoprodamesella , oenonella , paraxenocephalus , peculicephalina , pichyklen , pliomerellus , prodamesella ( = metaprodamesella ; = neoprodamesella ) , pseudoclelandia , pseudodipharus , pseudosalteria , pseudosarkia , punctaspis , qijiangia , resseria , robroyia , sakhaspidella , spizharaspis , subitella , sukhanaspis ( = kerbinella ) , tersiceps , thymurus , toxotis , trapezocephalina , typhlokorynetes , yinaspis , yoyarria .
family indeterminate aguaraya , anthracopeltis , aytounella , cancapolia , chalchaquiana , chambersiellus , columbicephalus , cuchulain , cuyanaspis , desmus , diplozyga , eilidh , ellesides , epumeria , glossicephalus , hagiorites , hoekaspiella , hsiaella , huaquinchaia , huilichia , kilmahogia , leptopilus , levinia , melopetasus , mendodiscus , meneghinella , menocephalus , mial , neilsoniella , neochilonorria , neotaenicephalus , nicoljarvius , pegelina , perthiellus , pichunia , pichynturia , piliolites , pseudolevinia , querandinia , tambakia , tchabdania , triarthrella , yanquetruzia .
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parent taxon : zacanthoididae according to p . a . jell and j . m . adrain 2003
full reference : c . e . resser . 1939 . the spence shale and its fauna . smithsonian miscellaneous collections 97 ( 12 ) : 1 - 29
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this trilobite website is designed to have several unique characteristics , all presented to provide visitors with an entertaining and hopefully enlightening experience with these amazing ancient arthropods . each of our featured trilobite localities has been chosen for its particular geographic or historic significance . as an example , it was decided to launch the site by focusing upon a locality within new york state , the home of the american museum of natural history . the rochester shale has long been renowned for its diverse silurian fauna , including arctinurus boltoni , which in 1825 was only the second trilobite species described from new york state . we will also visit many of the world ' s most prominent trilobites locations… from the hot deserts of morocco , to the rolling hills of china , to the high altiplano of bolivia . our intent here is to provide a more detailed overview of exactly how and where many of our featured specimens were found . the gallery of trilobites will display some of the very best specimens that have been collected in localities around the globe . we will do our best to present specimens possessing unique scientific import , as well as those that are simply magnificent examples of their specific species . in the trilobite files , we will present detailed reports that delve into the myriad mysteries of the trilobite world . from the first trilobites , to the uniqueness of trilobite eyes , to the amazing tale told by the widespread distribution of the trilobite paradoxides , all these issues - - and many more - - will be properly addressed and “filed” in this website section . despite the fact that this website is designed and maintained by amateur trilobite enthusiasts , we will always try to make sure that scientific accuracy is prioritized .
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bathyuriscus rotundatus ( ummp 4884 ) . presumed moult , lacking free cheeks . specimen length = 45 mm . specimen dry – direct light . trilobite beds on mount stephen .
trilobites are extinct euarthropods , probably stem lineage representatives of the mandibulata , which includes crustaceans , myriapods , and hexapods ( scholtz and edgecombe , 2006 ) .
bathyuriscus – a variation of the earlier trilobite genus name bathyurus , originally based on the greek bathys , “deep , ” and the greek oura , “tail , ” thus , a trilobite with a deep tail .
rotundatus – from the latin rotundus , “round , ” presumably alluding to the rounded outline of the dorsal shield .
type status under review – ummp 4884 ( 9 specimens ) , university of michigan museum of paleontology , ann arbor , michigan , usa .
burgess shale and vicinity : bathyuriscus adaeus walcott , 1916 , from several localities higher in the bathyuriscus - elrathina zone on mount stephen , mount odaray , and park mountain .
other deposits : other species of bathyuriscus have been described from numerous localities elsewhere in the cambrian of north america .
the trilobite beds and other localities on mount stephen . fossil ridge in sections stratigraphically below the walcott quarry .
hard parts : adult dorsal exoskeletons may be up to 5 cm long and are narrowly oval in outline , with a semicircular cephalon , a thorax of nine segments ending in blade - like tips with short spines , and a semicircular pygidium without spines .
the long glabella reaches almost to the anterior cephalic border ; the posterior portion is narrow and parallel - sided , while the anterior third expands rapidly forward . there are four pairs of lateral glabellar furrows , with the two front pairs angled forward and the posterior pair directed obliquely back . the eyes are relatively long and lie close to the glabella . broad free cheeks are extended back into short genal spines . the pygidium is slightly smaller than the cephalon , with a well - defined narrow axial lobe of five rings and a terminal piece ; four pairs of pygidial ribs are usually visible . the exoskeleton is mostly smooth externally , but very well preserved specimens may show faint anastomosing ridges on the free cheeks .
extremely common in the mount stephen trilobite beds , where it rivals ogygopsis klotzi in abundance .
bathyuriscus rotundatus was a mobile epibenthic trilobite . because we have no direct evidence of limb structure , its feeding habits are uncertain . it may have been a deposit feeder and opportunistic scavenger . like ogygopsis , bathyuriscus may occur as fully intact individuals ( probably carcasses ) , with the free cheeks missing , inverted , or rotated ( presumed moults ) , and as scattered pieces . some show evidence of healed injuries that may be predation scars ( rudkin , 2009 ) .
rasetti , f . 1951 . middle cambrian stratigraphy and faunas of the canadian rocky mountains . smithsonian miscellaneous collections , 116 ( 5 ) : 1 - 277 .
rominger , c . 1887 . description of primordial fossils from mount stephens , n . w . territory of canada . proceedings of the academy of natural sciences of philadelphia , 1887 : 12 - 19 .
rudkin , d . m . 2009 . the mount stephen trilobite beds , pp . 90 - 102 . in j . - b . caron and d . rudkin ( eds . ) , a burgess shale primer - history , geology , and research highlights . the burgess shale consortium , toronto .
scholtz , g . and edgecombe , g . d . 2006 . the evolution of arthropod heads : reconciling morphological , developmental and palaeontological evidence . development genes and evolution , 216 : 395 - 415 .
walcott , c . d . 1888 . cambrian fossils from mount stephens , northwest territory of canada . american journal of science , series 3 : 163 - 166 .
walcott , c . d . 1908 . mount stephen rocks and fossils . canadian alpine journal , 1 : 232 - 248 .
walcott , c . d . 1916 . cambrian geology and paleontology iii . cambrian trilobites . smithsonian miscellaneous collections , 64 ( 5 ) : 303 - 456 .
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