Before revision:  In this article, we describe how to encode animal behavior data as different types of multilayer networks, and we link recently-developed multilayer methods to individual-, group-, population-, and evolutionary-level questions in behavioral ecology. We outline a variety of examples for how to apply multilayer methods in behavioral ecology research, including examples that demonstrate how taking a multilayer approach can alter inferences about social structure and the positions of individuals within such a structure.
After revision:  In this article, we describe how to encode animal behavior data as different types of multilayer networks, and we link recently-developed multilayer methods to individual-, group-, population-, and evolutionary-levels. We give examples for how to apply multilayer methods in behavioral ecology research, including examples that demonstrate how taking a multilayer approach can alter inferences about social structure and the positions of individuals within such a structure.