Before revision:  However, we still lack a systematic understanding of the neural mechanism of how animals perceive, maintain, and use time intervals ranging from hundreds of milliseconds to multi-seconds in working memory and appropriately combine time information with spatial information processing and decision making  .
After revision:  However, we still lack a systematic understanding of the neural mechanism of how animals perceive, maintain, and use time intervals ranging from hundreds of milliseconds to multi-seconds in working memory , the combination of time information with spatial information processing and decision making .