Before revision:  In the study of Valleriani }{\it   is assumed to be the mean value of measured translation rate, i.e. the mean value of ratio of the translation number of protein to the lifetime of mRNA. However, in experiments different methods might be used to get \rm tl.
After revision:  In the study of Valleriani }{\it , \omega_{\rm tl is assumed to be the mean value of measured translation rate, i.e. the mean value of ratio of the translation number of protein to the lifetime of mRNA. However, in experiments different methods might be used to get \rm tl.