Before revision: The performance of the microscope depends on the state of various tunable knobs , corresponding to distances and orientations of various lenses, whereas the fitness of an organism depends on the state of various phenotypic characters such as body size, beak length and beak depth.
After revision: The performance of the microscope depends on the state of various knobs that can be manipulated , corresponding to distances and orientations of various lenses, whereas the fitness of an organism depends on the state of various phenotypic characters such as body size, beak length and beak depth.