Before revision: Two possibilities were considered: 1) That a social trait might reliably operate straightforwardly via social context in species where genetic relatives are usually concentrated in a local home area where they were born ("viscous populations"); 2) That genetic detection mechanisms ("supergenes") might emerge that go beyond statistical correlations, and reliably detect actual genetic relatedness between the social actors using direct "kin recognition".
After revision: Two possibilities were considered: 1) That a social trait might reliably operate straightforwardly via social context in species where close relatives are usually concentrated in a local home area where they were born ("viscous populations"); 2) That genetic detection mechanisms ("supergenes") might emerge that go beyond statistical correlations, and reliably detect actual genetic relatedness between the social actors using direct "kin recognition".