Before revision:  By studying a simplified model, the delayed response of motors to rapid fluctuations in the cargo  is quantified, allowing for the reduction of the full model to two "characteristic positions " of each of the motor populations .
After revision:  By studying a simplified model, the delayed response of motors to rapid fluctuations in the cargo is quantified, allowing for the reduction of the full model to two "characteristic positions " , a proxy for the net force on the cargo .