Before revision:  This allows to analyze the influence of a refractory (insensitivity) period following a reversal of movement.
After revision:  However we found that the derived macroscopic model lacks a diffusion term in the density equations, which is necessary to control the number of waves, indicating that a higher order approximation during the derivation is crucial. Upon ad-hoc addition of the diffusion term, we found very good agreement between the age-structured model and the biology. In particular we analyzed the influence of a refractory (insensitivity) period following a reversal of movement.