Before revision:  Our experiments also indicated that an asymmetric (polarized) dynein-microbutule on-rate, causes the force imbalance positioning the spindle. GPR-1/2 increases the overall dynein density but also residency time (pulling processivity), although this latter was not polarized .
After revision:  Our experiments also indicated that an asymmetric (polarized) dynein-microbutule on-rate, causes the force imbalance reflecting polarity, and supplements the regulation of mitotic progression via the non-polarized detachment rate .