Before revision:  Under certain limits this general expression of energy is reduced to the worm-like chain (WLC) model . Using this simplified energy in the strong-stretching limit, we obtain analytical solutions for all the cumulants of the end-to-end distance projected to the force direction. Finally we apply our results to recent experimental observations that revealed the hitherto unknown periodic cytoskeleton structure of axons and measured the longitudinal fluctuations , and show how the comparison between our results and experiments limits possible local configurations and dynamics of the spectrin tetramers in the axonal cytoskeleton  .
After revision:  Under certain limits this general expression of energy is reduced to the worm-like chain (WLC) model to the network level and investigate the situation when the system is stretched by strong forces (weakly bending limit). We specifically studied recent experimental observations that revealed the hitherto unknown periodic cytoskeleton structure of axons and measured the longitudinal fluctuations , and show how the comparison between our results and experiments limits possible local configurations and dynamics of the spectrin tetramers in the axonal cytoskeleton .