Before revision: Background: The correct positioning of the mitotic spindle during the asymmetric division of the nematode C. elegans zygote relies on the combination of centering and corticalpulling forces. These forces, revealed by centrosome anaphase oscillations, are regulated through the dynamics of force generators, related to mitosis progression.
After revision: Background: The correct positioning of the mitotic spindle during the asymmetric division of the Caenorhabditis elegans nematode zygote, the polarity cues distribution and daughter cell fates depend on the correct positioning of the mitotic spindle which results from both centering and cortical pulling forces. Revealed by spindle rocking, these pulling forces are regulated by the force generator dynamics, which are related to mitosis progression.