Before revision:  Additionally, taking into account the cell-intrinsic force generation on apical and basal sides , which are also broadly observed in morphogenesis, homeostasis, and carcinogenesis, we found apically/basally directed cell delaminations and pseudostratified structures, which could universally explain mechanical regulations of epithelial geometries in both physiology and pathophysiology{\it   .
After revision:  Additionally, taking into account the cell-intrinsic force generation on apical and basal sides . Although our analyses are based on a simple mechanical model, our results are in accordance with observations of epithelial monolayers{\it .