Before revision:  Given a distribution of phenotypes on a fitness landscape , we summarize how natural selection moves a population in the direction of a natural gradient , similar to natural gradientdescent, and show how intermediate selection is most informative of the fitness landscape.
After revision:  Optimization of a continuous objective function is analogous to searching for high fitness phenotypes on a fitness landscape , we summarize how natural selection moves a population in the direction of a natural gradient , similar to natural gradientdescent, and show how intermediate selection is most informative of the fitness landscape.