Before revision:  These include not only the change in the molecular properties associated with the chemical modifications of DNA and histone proteins - such as methylation and acetylation - but also less conventional ones, such as the physics that governs the URLanization of the genome in cell nuclei.
After revision:  These include not only the change in the molecular properties associated with the chemical modifications of DNA and histone proteins , such as methylation and acetylation - but also less conventional ones, such as the physics that governs the URLanization of the genome in cell nuclei.