Before revision: The footprints left enough evidence for researchers to determine that the crab probably fell into the lagoon upside-down, righted itself, and started walking before succumbing to the anoxic conditions of the water before dying.
After revision: The footprints left enough evidence for researchers to determine that the creature probably fell into the lagoon upside-down, righted itself, and started walking before succumbing to the anoxic conditions of the water before dying.