Before revision:  Although the enzymes involved in multiple glycan substrates indicate cross-talk between modules, the impact of the cross-talk on the module-specific enhancement of glycan synthesis could be confined within a moderate range by transcriptional-level control.
After revision:  Although the enzymes involved in multiple glycan substrates , indicating cross-talk between modules, the impact of the cross-talk on the module-specific enhancement of glycan synthesis could be confined within a moderate range by transcriptional-level control.