Before revision:  This congruence is explained by the agreement of bifurcations in dynamical-systems theory between evolution and development, where slowly varying gene-expression levels work as emergent control parameters. In terms of the gene regulation networks, this congruence is understood as the successive addition of downstream modules, either as feedforward or feedback , while the upstream feedforward network shapes the boundary condition for the downstream dynamics, based on the maternal morphogen gradient.
After revision:  This congruence is explained by the upstream feedforward network shapes the boundary condition for the downstream dynamics, based on the maternal morphogen gradient.