Before revision:  This model not only recapitulates monoallelic OR expression, but also elucidates how the olfactory system maximizes and maintains the diversity of OR expression . The model has multiple predictions validated by existing experimental results , and particularly underscores cooperativity and synergy as a general design principle for multi-objective optimization in biology 
After revision:  This model not only recapitulates monoallelic OR expression, but also elucidates how the olfactory system maximizes and maintains the diversity of OR expression , and has multiple predictions validated by existing experimental results , and particularly underscores cooperativity and synergy as a general design principle for multi-objective optimization in biology