Before revision:  In evolutionary biology, these systems are studied within the general framework of fluctuating environments , where the individual entities are subject to spatio-temporal fluctuations in the environmental conditions. In these environments, evolutionary behavior may emerge that essentially differs from the one observed in standard models .
After revision:  In evolutionary biology, these systems are studied within the general framework of fluctuating environments , where the individual entities are subject to spatio-temporal stochasticity plays a significant role in the evolutionary dynamics .