Before revision:  Indeed, they provide a viewpoint for the most classical time-scales of interest. They namely encompass the time-scale of the life of a single individual, the longer one on which a population can be characterized , the ecological one and at least four nested ones on which selection occurs.
After revision:  Indeed, they provide a viewpoint for the most classical time-scales . Namely, they encompass the timescale typical of the life-expectancy of a single individual, the longer one on which a population can be characterized , the ecological one and at least four nested ones on which selection occurs.