(E) Cartoon depicting the rearrangement of actin and UapA at the tip of A. nidulans during germling to hyphal maturation. In germlings, dense arrays of actin cables are present in the apex, known as apical actin array (AAA). Actin patches are concentrated, but not restricted, to the apex. In mature hyphae, a core of actin appears in the position of the Spitzenkorper and actin patches are restricted to the subapical endocytic collar (≈2 μm behind the apex). Actin cables are retracted from the apex and shifted to a more distal region of the hyphal tip, the subapical actin web (SAW). In the apex, actin cables are now found associated mostly as sparse arrays across the cell cortex. The localization of UapA during hyphal maturation seems to follow the pattern of actin cables [26,27 