Original: He uses this to explain general influences from Totonacan and Mixe-Zoquean languages in many other Mesoamerican languages many of which do not have any known history of contact with either of the above-mentioned groups.
Simplified: He uses this to explain general influences from Totonacan and Mixe-Zoquean languages in many other Mesoamerican languages. Many of these languages do not have any known history of contact with either of the above-mentioned groups.