Original: From 1842 to 1848 he studied physics, chemistry, botany, mineralogy, philosophy and medicine at the University of Giessen, where he graduated in 1848 with a dissertation entitled Beiträge zur Hall'schen Lehre von einem excitomotorischen Nervensystem (Contributions to the Hallerian Theory of an Excitomotor Nervous System).
Simplified: From 1842 to 1848 he studied physics, chemistry, botany, mineralogy, philosophy and medicine at the University of Giessen.