title: Cats (musical)

text: Madame Tussauds New York features wax figures of several characters from the musical, including one of Grizabella that sings "Memory" through the use of projection mapping technology. Similarly, a wax figure of Rumpleteazer is displayed at the Panoptikum wax museum in Hamburg, Germany.


title: Cats (musical)

text: A "Cats" postage stamp was issued by the United States Postal Service in 2000 as part of its Celebrate the Century series. The musical was chosen as one of fifteen "outstanding artifacts, events and activities" from the 1980s to be commemorated with its own stamp design. Beginning in March 2019, the Rinkai Line in Tokyo, Japan, uses two songs from the musical for its train departure melodies at the Ōimachi Station; the train to Ōsaki Station uses a jingle of "Memory", while the train to Shin-Kiba Station uses a jingle of "Skimbleshanks: The Railway Cat".


title: Cats (musical)

text: The visual effects of the 2019 film adaptation were mocked in the 2022 Disney film "", in a scene where the titular characters notice two alley cats resembling the ones in the film as they venture through an "uncanny valley" part of town.


title: Cats (musical)

text: The stage production of "Cats" has undergone several revisions since its London opening. When the show transferred to Broadway, several cuts and rewrites were made with the intention of appealing more to an American audience. Additionally, a song entitled "Cat Morgan Introduces Himself" was cut during initial development. Lloyd Webber performed this song at the show's 6,138th Broadway performance, when it broke the record to become the longest-running Broadway show.


title: Cats (musical)

text: The "Growltiger's Last Stand" sequence has been changed multiple times over the course of the show's history. In the original London production, the "last duet" for Growltiger and Griddlebone was a setting for an unpublished Eliot poem, "The Ballad of Billy M'Caw". For the original Broadway production, the Ballad was replaced with "In Una Tepida Notte", a parody of Italian opera with more slapstick humour. This new version was eventually incorporated into all other productions of "Cats".


title: Cats (musical)

text: "Growltiger's Last Stand" has been criticised as being racially offensive. The original lyrics, taken directly from the Eliot poem it is based on, included the ethnic slur "Chinks" and this was later replaced with the word "Siamese". The number also originally involved the cast putting on "Asian accents" to portray the Siamese cats.


title: Cats (musical)

text: In the 1998 video version, the entire scene featuring Growltiger was cut. By 2016, "Growltiger's Last Stand" had been removed completely from the US and UK productions of the show. The 2019 Vienna Revival included the song, but since its closing in June 2022, "Growltiger's Last Stand" is now not present in any replica production of Cats.


title: Cats (musical)

text: In the original London production, Mungojerrie and Rumpleteazer were characters in their own right and sang their eponymous song themselves as a singsong-style duet. When the show transferred to Broadway, the song was instead sung in the third-person, with Mungojerrie and Rumpleteazer as puppets being magically controlled by Mr. Mistoffelees. Their number was also rewritten to be faster and more upbeat, alternating between vaudeville-style verses and a "manic patter" section. Eventually, the Broadway version of the song was rewritten to allow Mungojerrie and Rumpleteazer to once again sing their own song as full characters.


title: Cats (musical)

text: The 2014 London revival introduced several modernisations to the show. Rum Tum Tugger was reworked from a ladies-man rockstar to a breakdancing street cat. His eponymous musical number was also turned into a rap. The 2015 Australian tour and 2015 Paris production also used the new version of the character; however, the 2016 Broadway revival did not.


title: Cats (musical)

text: The 2016 Broadway revival featured new choreography by Andy Blankenbuehler, who introduced more hip hop and cool jazz elements to the movements and dances. Blankenbuehler's choreography for the ensemble numbers did not differ too much from the original by Lynne, but significant changes were made in several solo numbers, including "The Rum Tum Tugger" and "Mr. Mistoffelees".


title: UFC 276

text: UFC 276: Adesanya vs. Cannonier was a mixed martial arts event produced by the Ultimate Fighting Championship that took place on July 2, 2022, at the T-Mobile Arena in Paradise, Nevada, part of the Las Vegas Metropolitan Area, United States.


title: UFC 276

text: The event was held during the UFC's 10th annual International Fight Week. The 2022 UFC Hall of Fame induction ceremony also took place during the week.


title: UFC 276

text: A UFC Middleweight Championship bout between current champion Israel Adesanya and Jared Cannonier headlined the event.


title: UFC 276

text: A UFC Featherweight Championship trilogy bout between current champion Alexander Volkanovski and former champion Max Holloway took place at this event. The pairing first met at UFC 245, where Volkanovski defeated Holloway via unanimous decision to capture the title. Their second meeting took place at UFC 251, with Volkanovski successfully defending the title via split decision. They were originally expected to meet for the third time at UFC 272, but Holloway withdrew due to an aggravated previous injury.


title: UFC 276

text: A women's flyweight bout between former Strikeforce and UFC Women's Bantamweight Champion Miesha Tate and former Invicta FC Bantamweight Champion (also former UFC Women's Flyweight Championship challenger) Lauren Murphy was expected to take place at UFC on ESPN 36. However, the bout was moved to this event due to unknown reasons. In turn, a week before the event, Murphy pulled out after she tested positive for COVID-19. The bout was then rescheduled to .


title: UFC 276

text: A middleweight bout between Uriah Hall and André Muniz took place at the event. The pairing was originally scheduled to meet as but Hall withdrew for unknown reasons.


title: UFC 276

text: A women's flyweight bout between former UFC Women's Flyweight Championship challenger Jessica Eye and Casey O'Neill was expected to take place at the event. However, O'Neill withdrew in late April due to a torn ACL and was replaced by Maycee Barber.


title: UFC 276

text: Sean Strickland was expected to face former Glory Middleweight and Light Heavyweight Champion Alex Pereira in a middleweight bout at UFC 277. However, the promotion decided to move the pairing to this event.


title: UFC 276

text: A lightweight bout between Bobby Green and Jim Miller was previously booked twice, but was canceled on both occasions: first at UFC 172 in April 2014, when Green pulled out due to an elbow injury. Green also pulled out of the second booking at UFC 258 in February 2021, when he collapsed after the weigh-ins and was deemed unfit to compete. They were then expected to meet at this event. In turn, Green withdrew again for unknown reasons and was replaced by former UFC Lightweight Championship challenger Donald Cerrone in a welterweight bout. Miller and Cerrone previously met 8 years prior at which Cerrone won by second round knockout.


title: UFC 276

text: The following fighters received Crypto.com "Fan Bonus of the Night" awards paid in bitcoin of US$30,000 for first place, US$20,000 for second place, and US$10,000 for third place.


title: Tarantula

text: Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. , 1,040 species have been identified, with 156 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although many other members of the same infraorder (Mygalomorphae) are commonly referred to as "tarantulas" or "false tarantulas". Some of the more common species have become popular in the exotic pet trade. Many New World species kept as pets have setae known as urticating hairs that can cause irritation to the skin, and in extreme cases, cause damage to the eyes.


title: Tarantula

text: Like all arthropods, the tarantula is an invertebrate that relies on an exoskeleton for muscular support. Like other Arachnida, a tarantula's body comprises two main parts, the prosoma (or cephalothorax) and the opisthosoma (or abdomen). The prosoma and opisthosoma are connected by the pedicel, or pregenital somite. This waist-like connecting piece is actually part of the prosoma and gives the opisthosoma a wide range of motion relative to the prosoma.


title: Tarantula

text: Tarantula sizes can range from as small as the size of a BB pellet to as large as a dinner plate when the legs are fully extended. Depending on the species, the body length of tarantulas ranges from about , with leg spans of . Leg span is determined by measuring from the tip of the back leg to the tip of the front leg on the opposite side. Some of the largest species of tarantula may weigh over ; the largest of all, the goliath birdeater ("Theraphosa blondi") from Venezuela and Brazil, has been reported to attain a weight of and a leg-span up to , males being longer and females greater in girth. The fang size of this tarantula reaches a maximum of .


title: Tarantula

text: "Theraphosa apophysis" (the pinkfoot goliath) was described 187 years after the goliath birdeater, so its characteristics are not as well attested. "T. blondi" is generally thought to be the heaviest tarantula, and "T. apophysis" has the greatest leg span. Two other species, "Lasiodora parahybana" (the Brazilian salmon birdeater) and "Lasiodora klugi", rival the size of the two goliath spiders.


title: Tarantula

text: Most species of North American tarantulas are brown. Elsewhere, species have been found that variously display cobalt blue ("Cyriopagopus lividus"), black with white stripes ("Aphonopelma seemanni"), yellow leg markings ("Eupalaestrus campestratus"), metallic blue legs with vibrant orange abdomen and green prosoma ("Chromatopelma cyaneopubescens"). Their natural habitats include savanna, grassland such as in the pampas, rainforest, desert, scrubland, mountains, and cloud forest. They are generally classed among the terrestrial types. They are burrowers that live in the ground.


title: Tarantula

text: Tarantulas are becoming increasingly popular as pets and some species are readily available in captivity.


title: Tarantula

text: Tarantulas can be confused with other members of the order Mygalomorphae, such as trapdoor spiders, funnel-web spiders and purseweb spiders. They can also be confused with some members of the order Araneomorphae such as the Lycosidae family. There are multiple ways to identify a tarantula. First the hairs: in the Americas most tarantulas have urticating hairs, though some, such as the "Hemirrhagus" genus, lack these. The hairs are usually more noticeable than with most other spiders. Another is the size, as tarantulas tend to be bigger, but this is again not a failproof way. They also don't use their webs for hunting, instead using them as building material or tripwire.


title: Tarantula

text: One of the most decisive ways to tell is by looking at their fangs. Tarantula fangs face downwards, as opposed to those of true spiders, which face each other, allowing them to make pincerlike motions. They also own two book lungs, as opposed to true spiders which only have one. Their lifespan is also longer than most spiders.


title: Tarantula

text: The spider originally bearing the name "tarantula" was "Lycosa tarantula", a species of wolf spider native to Mediterranean Europe. The name is derived from the southern Italian town of Taranto. The term "tarantula" was subsequently applied to almost any large, unfamiliar species of ground-dwelling spider, in particular to the Mygalomorphae and especially the New World Theraphosidae. Compared to tarantulas, wolf spiders are not particularly large or hairy, and so among English speakers in particular, usage eventually shifted in favour of the Theraphosidae, even though they are not closely related to wolf spiders at all, being in a different infraorder.


title: Tarantula

text: The name "tarantula" is also incorrectly applied to other large-bodied spiders, including the purseweb spiders or atypical tarantulas, the funnel-webs (Dipluridae and Hexathelidae), and the dwarf tarantulas. These spiders are related to tarantulas (all being mygalomorphs) but fall into different families from them. Huntsman spiders of the family Sparassidae have also been termed "tarantulas" because of their large size, when, in fact, they are not related. Instead, huntsman spiders belong to the infraorder Araneomorphae.


title: Tarantula

text: Many theraphosid genera have names, either accepted or synonymous, containing the element "pelma". This can be traced back to Carl Ludwig Koch in 1850, who in describing his new genus "Eurypelma" wrote, "" (). German arachnologists use the word to refer to the tarsus (the last article of a spider's leg). Translations of into Latin use the word . Hence in English arachnological terminology, Koch meant 'the scopula of the base of the tarsus very wide'. "Eury-" is derived from the Greek (), meaning 'wide', while () means 'the sole of the foot', paralleling Koch's use of (in modern spelling). Thus "Eurypelma" literally means 'wide footsole'; however, arachnologists have conventionally taken "pelma" in such names to refer to the scopula, so producing the meaning 'with a wide scopula'.


title: Tarantula

text: Other genus names or synonyms that Estrada-Alvarez and Cameron regard as having 'footsole' or 'scopula' meanings include:


title: Tarantula

text: Later, particularly following genus names published by R.I. Pocock in 1901, the element "pelma" appears to have become synonymous with 'theraphosid'. For example, the author of "Cardiopelma" writes, "" ('Cardiopelma refers to the female genitalia that evoke the shape of a heart'), with no reference to either 'footsole' or 'scopula'. Names interpreted in this way include:


title: Tarantula

text: Tarantulas of various species occur throughout the United States, Mexico, in Central America, and throughout South America. Other species occur variously throughout Africa, much of Asia (including the Ryukyu Islands in southern Japan), and all of Australia. In Europe, some species occur in Spain, Portugal, Turkey, southern Italy, and Cyprus.


title: Tarantula

text: Some genera of tarantulas hunt prey primarily in trees; others hunt on or near the ground. All tarantulas can produce silk; while arboreal species typically reside in a silken "tube tent", terrestrial species line their burrows with silk to stabilize the burrow wall and facilitate climbing up and down. Tarantulas mainly eat large insects and other arthropods such as centipedes, millipedes, and other spiders, using ambush as their primary method of prey capture. Armed with their massive, powerful chelicerae tipped with long, chitinous fangs, tarantulas are well-adapted to killing other large arthropods. The biggest tarantulas sometimes kill and consume small vertebrates such as lizards, mice, bats, birds, and small snakes.


title: Tarantula

text: The eight legs, the two chelicerae with their fangs, and the pedipalps are attached to the prosoma. The chelicerae are two double-segmented appendages located just below the eyes and directly forward of the mouth. The chelicerae contain the venom glands that vent through the fangs. The fangs are hollow extensions of the chelicerae that inject venom into prey or animals that the tarantula bites in defense, and they are also used to masticate. These fangs are articulated so that they can extend downward and outward in preparation to bite or can fold back toward the chelicerae as a pocket knife blade folds back into its handle. The chelicerae of a tarantula completely contain the venom glands and the muscles that surround them, and can cause the venom to be forcefully injected into prey.


title: Tarantula

text: The pedipalpi are two six-segmented appendages connected to the prosoma near the mouth and protruding on either side of both chelicerae. In most species of tarantulas, the pedipalpi contain sharp, jagged plates used to cut and crush food often called the coxae or maxillae. As with other spiders, the terminal portions of the pedipalpi of males function as part of their reproductive system. Male spiders spin a silken platform (sperm web) on the ground onto which they release semen from glands in their opisthosoma. Then they insert their pedipalps into the semen, absorb the semen into the pedipalps, and later insert the pedipalps (one at a time) into the reproductive organ of the female, which is located in her abdomen. The terminal segments of the pedipalps of male tarantulas are moderately larger in circumference than those of a female tarantula. Male tarantulas have special spinnerets surrounding the genital opening. Silk for the sperm web of the tarantula is exuded from these special spinnerets.


title: Tarantula

text: A tarantula has four pairs of legs and two additional pairs of appendages. Each leg has seven segments, which from the prosoma out are: coxa, trochanter, femur, patella, tibia, tarsus and pretarsus, and claw. Two or three retractable claws at the end of each leg are used to grip surfaces for climbing. Also on the end of each leg, surrounding the claws, is a group of bristles, called the scopula, which help the tarantula to grip better when climbing surfaces such as glass. The fifth pair is the pedipalps, which aid in feeling, gripping prey, and mating in the case of a mature male. The sixth pair of appendages is the chelicerae and their attached fangs. When walking, a tarantula's first and third legs on one side move at the same time as the second and fourth legs on the other side of its body. The muscles in a tarantula's legs cause the legs to bend at the joints, but to extend a leg, the tarantula increases the pressure of haemolymph entering the leg.


title: Tarantula

text: Tarantulas, like almost all other spiders, have their primary spinnerets at the end of the opisthosoma. Unlike most spider species in the infraorder Araneomorphae, which includes the majority of extant spider species, and most of which have six, tarantula species have two or four spinnerets. Spinnerets are flexible, tube-like structures from which the spider exudes its silk. The tip of each spinneret is called the spinning field. Each spinning field is covered by as many as 100 spinning tubes through which silk is exuded. As the silk is pulled out of the spinnerets, the shear forces cause proteins in the silk to crystallize, transforming it from a liquid to a solid thread.


title: Tarantula

text: The tarantula's mouth is located under its chelicerae on the lower front part of its prosoma. The mouth is a short, straw-shaped opening that can only suck, meaning that anything taken into it must be in liquid form. Prey with large amounts of solid parts, such as mice, must be crushed and ground up or predigested, which is accomplished by coating the prey with digestive juices secreted from openings in the chelicerae.


title: Tarantula

text: The tarantula's digestive organ (stomach) is a tube that runs the length of its body. In the prosoma, this tube is wider and forms the sucking stomach. When the sucking stomach's powerful muscles contract, the stomach is increased in cross-section, creating a strong sucking action that permits the tarantula to suck its liquefied prey up through the mouth and into the intestines. Once the liquefied food enters the intestines, it is broken down into particles small enough to pass through the intestine walls into the hemolymph (blood stream), where it is distributed throughout the body. After feeding, the leftovers are formed into a small ball by the tarantula and thrown away. In a terrarium, they often put them into the same corner.


title: Tarantula

text: A tarantula's central nervous system (brain) is located in the bottom of the inner prosoma. A tarantula perceives its surroundings primarily via sensory organs called setae (bristles or spines, sometimes referred to as hairs). Although a tarantula has eight eyes like most spiders, touch is its keenest sense, and in hunting, it primarily depends on vibrations given off by the movements of its prey. A tarantula's setae are very sensitive organs and are used to sense chemical signatures, vibrations, wind direction, and possibly even sound. Tarantulas are also very responsive to the presence of certain chemicals such as pheromones.


title: Tarantula

text: The eyes are located above the chelicerae on the forward part of the prosoma. They are small and usually set in two rows of four. Most tarantulas are not able to see much more than light, darkness, and motion. Arboreal tarantulas generally have better vision compared with terrestrial tarantulas.


title: Tarantula

text: All types of tarantulas have two sets of book lungs (breathing organs); the first pair is located in a cavity inside the lower front part of the abdomen near where the abdomen connects to the cephalothorax, and the second pair is slightly farther back on the abdomen. Air enters the cavity through a tiny slit on each side of and near the front of the abdomen. Each lung consists of 15 or more thin sheets of folded tissue arranged like the pages of a book. These sheets of tissue are supplied by blood vessels. As air enters each lung, oxygen is taken into the blood stream through the blood vessels in the lungs. Needed moisture may also be absorbed from humid air by these organs.


title: Tarantula

text: A tarantula's blood is unique (not only in appearance); an oxygen-transporting protein is present (the copper-based hemocyanin), but not enclosed in blood cells such as the erythrocytes of mammals. A tarantula's blood is not true blood, but rather a liquid called hemolymph (or haemolymph). At least four types of hemocytes, or hemolymph cells, are known.


title: Tarantula

text: The tarantula's heart is a long, slender tube located along the top of the opisthosoma. The heart is neurogenic as opposed to myogenic, so nerve cells instead of muscle cells initiate and coordinate the heart. It pumps hemolymph to all parts of the body through open passages often referred to as sinuses, and not through a circular system of blood vessels. If the exoskeleton is breached, loss of hemolymph will kill the spider unless the wound is small enough that the hemolymph can dry and close it.


title: Tarantula

text: Despite their large size and fearsome appearance and reputation, tarantulas themselves are prey for many other animals. The most specialized of these predators are large members of the wasp family Pompilidae such as the wasp "Hemipepsis ustulata". These wasps are called "tarantula hawks". The largest tarantula hawks, such as those in the genus "Pepsis", track, attack, and kill large tarantulas. They use olfaction to find the lair of a tarantula. The wasp must deliver a sting to the underside of the spider's cephalothorax, exploiting the thin membrane between the basal leg segments. This paralyzes the spider, and the wasp then drags it back into its burrow before depositing an egg on the prey's abdomen. The wasp then seals the spider in its burrow and flies off to search for more hosts. The wasp egg hatches into a larva and feeds on the spider's inessential parts, and as it approaches pupation, it consumes the remainder. Other arthropods, such as large scorpions and giant centipedes, are also known to prey on tarantulas.


title: Tarantula

text: Tarantulas are also preyed upon by a wide variety of vertebrates. Many of these, including lizards, frogs, birds, snakes and mammals, are generalist predators of all kinds of large arthropods. Mammals that have been known to prey on tarantulas, such as the coati, kinkajou, and opossum in the New World, and mongooses and the honey badger in the Old World, are often immune to the venom of their arthropod prey.


title: Tarantula

text: Humans also consume tarantulas for food in their native ranges. They are considered a delicacy in certain cultures (e.g. Venezuela and Cambodia). They can be roasted over an open fire to remove the bristles (described further below) and then eaten.


title: Tarantula

text: Tarantulas have evolved specialized bristles, or setae, to defend themselves against predators. Besides the normal bristles covering the body, some tarantulas also have a dense covering of irritating bristles called urticating hairs, on the opisthosoma, that they sometimes use as protection against enemies. These bristles are present on most New World species, but not on any specimens from the Old World. Urticating hairs are usually kicked off the abdomen by the tarantula, but some may simply rub the abdomen against the target, like the genus "Avicularia". These fine bristles are barbed and serve to irritate. They can be lethal to small animals such as rodents. Some people are sensitive to these bristles, and develop serious itching and rashes at the site. Exposure of the eyes and respiratory system to urticating hairs should be strictly avoided. Species with urticating hairs can kick these bristles off; they are flicked into the air at a target using their back pairs of legs. Tarantulas also use these bristles for other purposes, such as to mark territory or to line their shelters (the latter such practice may discourage flies from feeding on the spiderlings). Urticating hairs do not grow back, but are replaced with each molt. The intensity, number, and flotation of the bristles depends on the species of tarantula.


title: Tarantula

text: To predators and other enemies, these bristles can range from being lethal to simply being a deterrent. With humans, they can cause irritation to eyes, nose, and skin, and more dangerously, the lungs and airways, if inhaled. The symptoms range from species to species, from person to person, from a burning itch to a minor rash. In some cases, tarantula bristles have caused permanent damage to human eyes.


title: Tarantula

text: Some setae are used to stridulate, which makes a hissing sound. These bristles are usually found on the chelicerae. Stridulation seems to be more common in Old World species.


title: Tarantula

text: All tarantulas are venomous. Although their venom is not deadly to humans, some bites cause serious discomfort that might persist for several days. In general, the effects of the bites of all kinds of tarantula are not well known. While the bites of many species are known to be no worse than a wasp sting, accounts of bites by some species are reported to be very painful and to produce intense spasms that may recur over a period of several days; the venom of the African tarantula "Pelinobius muticus" also causes strong hallucinations. For "Poecilotheria" species, researchers have described more than 20 bites with the delayed onset of severe and diffuse muscle cramps, lasting for several days, that in most cases resolved completely with the use of benzodiazepines and magnesium. In all cases, seeking medical aid is advised. Because other proteins are included when a toxin is injected, some individuals may suffer severe symptoms due to an allergic reaction rather than to the venom. Such allergic effects can be life-threatening. Additionally, the large fangs of a tarantula can inflict painful puncture wounds, which can lead to secondary bacterial infections if not properly treated.


title: Tarantula

text: Before biting, a tarantula may signal its intention to attack by rearing up into a "threat posture", which may involve raising its prosoma and lifting its front legs into the air, spreading and extending its fangs, and (in certain species) making a loud hissing by stridulating. Tarantulas often hold this position for longer than the duration of the original threat. Their next step, without biting, may be to slap down on the intruder with their raised front legs. If that response fails to deter the attacker, the tarantulas of the Americas may next turn away and flick urticating hairs toward the pursuing predator. The next response may be to leave the scene entirely, but especially if no line of retreat is available, their final response may also be to whirl suddenly and bite. Some tarantulas are well known to give "dry bites", i.e., they may defensively bite some animal that intrudes on their space and threatens them, but they do not pump venom into the wound.


title: Tarantula

text: New-world tarantulas—those indigenous to the Americas—have bites that generally pose little threat to humans (other than causing localized pain). Most of them are equipped with urticating hairs on their abdomens, and almost always throw these barbed bristles as the first line of defense. These bristles irritate sensitive areas of the body and especially seem to target curious animals that may sniff these bristles into the mucous membranes of the nose. Some species have more effective urticating bristles than others. The goliath birdeater is known for its particularly irritating urticating bristles. They can penetrate the cornea, so eye protection should be worn when handling such tarantulas.


title: Tarantula

text: Old World tarantulas have no urticating bristles and are more likely to attack when disturbed. They often have more potent, medically significant venom, and are faster and much more nervous and defensive than New World species.


title: Tarantula

text: Some dangerous spider species are related to tarantulas and are frequently confused with them. A popular urban legend maintains that deadly varieties of tarantula exist somewhere in South America. This claim is often made without identifying a particular spider, although the "banana tarantula" is sometimes named. A likely candidate for the true identity of this spider is the dangerous Brazilian wandering spider ("Phoneutria fera") of the family Ctenidae, as it is sometimes found hiding in clusters of bananas and is one of several spiders called "banana spiders". It is not technically a tarantula, but it is fairly large (4- to 5-inch legspan), somewhat ″hairy″, and is highly venomous to humans. Another dangerous type of spiders that have been confused with tarantulas are the Australian funnel-web spiders. The best known species of these is the Sydney funnel-web spider ("Atrax robustus") a spider that is aggressive, highly venomous, and (prior to the development of antivenom in the 1980s) was responsible for numerous deaths in Australia. These spiders are members of the same infraorder as tarantulas, Mygalomorphae. Some Australians use the slang term "triantelope" (a corruption of the incorrect term tarantula, which is also used) for large, ″hairy″, and harmless members of the huntsman spider family, which are often found on interior household walls and in automobiles.


title: Tarantula

text: Some tarantula species exhibit pronounced sexual dimorphism. Males tend to be smaller (especially their abdomens, which can appear quite narrow) and may be dull in color when compared to their female counterparts, as in the species "Haplopelma lividum". Mature male tarantulas also may have tibial hooks on their front legs, which are used to restrain the female's fangs during copulation. Males typically have longer legs than the females.


title: Tarantula

text: A juvenile male's sex can be determined by looking at a cast exuvia for epiandrous fusillae or spermathecae. Females possess spermathecae, except for the species "Sickius longibulbi" and "Encyocratella olivacea". Males have much shorter lifespans than females because they die relatively soon after maturing. Few live long enough for a postultimate molt, which is unlikely in natural habitats because they are vulnerable to predation, but has happened in captivity, though rarely. Most males do not live through this molt, as they tend to get their emboli, mature male sexual organs on pedipalps, stuck in the molt. Most tarantula fanciers regard females as more desirable as pets due to their much longer lifespans. Wild-caught tarantulas are often mature males because they wander out in the open and are more likely to be caught.


title: Tarantula

text: Like other spiders, tarantulas have to shed their exoskeleton periodically as they grow, a process called molting. A young tarantula may do this several times a year as a part of the maturation process, while full-grown specimens only molt once a year or less, or sooner, to replace lost limbs or lost urticating hairs. It is visibly apparent that molting is imminent when the exoskeleton takes on a darker shade. If a tarantula previously used its urticating hairs, the bald patch turns from a peach color to deep blue. The tarantula also stops feeding and becomes more lethargic during this time.


title: Tarantula

text: While most Tarantulas species take between two to five years to reach sexual maturity, some species can take up to 10 years. Upon reaching adulthood, males typically have an 18-month period left to live so immediately go in search of a female mate. Although females continue to molt after reaching maturity, male rarely do again once they reach adulthood. Those that do often can become stuck during the molting process due to their sexual organs and die.


title: Tarantula

text: Female can live for 30 to 40 years. "Grammostola rosea" spiders, which eat once or twice a week, have lived up to 20 years in captivity. Some have survived on water alone for up to two years.


title: Tarantula

text: After reaching sexual maturity, a female tarantula normally mates and lays eggs once per year, although they do not always do so.


title: Tarantula

text: As with other spiders, the mechanics of intercourse are quite different from those of mammals. Once a male spider reaches maturity and becomes motivated to mate, he weaves a web mat on a flat surface. The spider then rubs his abdomen on the surface of this mat, and in so doing, releases a quantity of semen. He may then insert his pedipalps (short, leg-like appendages between the chelicerae and front legs) into the pool of semen. The pedipalps absorb the semen and keep it viable until a mate can be found. When a male spider detects the presence of a female, the two exchange signals to establish that they are of the same species. These signals may also lull the female into a receptive state. If the female is receptive, then the male approaches her and inserts his pedipalps into an opening in the lower surface of her abdomen, the opisthosoma. After the semen has been transferred to the receptive female's body, the male swiftly leaves the scene before the female recovers her appetite. Although females may show some aggression after mating, the male rarely becomes a meal.


title: Tarantula

text: Females deposit 50 to 2,000 eggs, depending on the species, in a silken egg sac and guard it for six to eight weeks. During this time, the females stay very close to the egg sacs and become more aggressive. Within most species, the females turn the egg sac often, which is called brooding. This keeps the eggs from deforming due to sitting in one position too long. The young spiderlings remain in the nest for some time after hatching, where they live off the remains of their yolk sacs before dispersing.


title: Tarantula

text: Linnaeus placed all spiders in a single genus, "Aranea". In 1802, Charles Athanase Walckenaer separated mygalomorph spiders into a separate genus, "Mygale", leaving all other spiders in "Aranea". However, "Mygale" had already been used in 1800 by Georges Cuvier for a genus of mammals (in Greek, "mygale" means "shrew"). Accordingly, in 1869, Tamerlan Thorell used the family name "Theraphosoidae" (modern Theraphosidae) for the mygalomorph spiders known to him, rather than "Mygalidae" (as used, for example, by John Blackwall). Thorell later split the family into a number of genera, including "Theraphosa".


title: Tarantula

text: A 2019 phylogenomic study recognized 12 subfamilies, one (Ischnocolinae) known not to be monophyletic.


title: Tarantula

text: The relationship between the subfamilies found in the study is shown in the following cladogram. The dual placing of Ischnocolinae is highlighted.


title: Tarantula

text: All the species that possess urticating hairs and have been seen to use them in bombardment behavior are placed in the "bombardier clade", although not all species in the included subfamilies possess such hairs (all "Schismatothelinae" lack them as do most "Psalmopoeinae" genera). It is not clear whether the possession of urticating hairs was an ancestral trait of the clade, and has been lost in some species, or whether it represents multiple gains. Foley "et al." suggested that the second hypothesis appeared to be better supported.


title: Tarantula

text: Although fossils of mygalomorph spiders date back to the Triassic, only two specimens have been found so far which can be convincingly assigned to the Theraphosidae. One is from Dominican Republic amber; the other is from Chiapas (Mexico) amber. Both these ambers are quite young, being Miocene in age or about 16 million years old.


title: Tortoise

text: Tortoises () are reptiles of the family Testudinidae of the order Testudines (Latin: "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like other members of the suborder Cryptodira, they retract their necks and heads directly backward into the shell to protect them.


title: Tortoise

text: Tortoises can vary in size with some species, such as the Galápagos giant tortoise, growing to more than in length, whereas others like the Speckled cape tortoise have shells that measure only long. Several lineages of tortoises have independently evolved very large body sizes in excess of 100 kg, including the Galapagos giant tortoise and the Aldabra giant tortoise. They are usually diurnal animals with tendencies to be crepuscular depending on the ambient temperatures. They are generally reclusive animals. Tortoises are the longest-living land animals in the world, although the longest-living species of tortoise is a matter of debate. Galápagos tortoises are noted to live over 150 years, but an Aldabra giant tortoise named Adwaita may have lived an estimated 255 years. In general, most tortoise species can live 80–150 years.


title: Tortoise

text: Differences exist in usage of the common terms turtle, tortoise, and terrapin, depending on the variety of English being used; usage is inconsistent and contradictory. These terms are common names and do not reflect precise biological or taxonomic distinctions.


title: Tortoise

text: The American Society of Ichthyologists and Herpetologists uses "turtle" to describe all species of the order Testudines, regardless of whether they are land-dwelling or sea-dwelling, and uses "tortoise" as a more specific term for slow-moving terrestrial species. General American usage agrees; turtle is often a general term (although some restrict it to aquatic turtles); tortoise is used only in reference to terrestrial turtles or, more narrowly, only those members of Testudinidae, the family of modern land tortoises; and terrapin may refer to turtles that are small and live in fresh and brackish water, in particular the diamondback terrapin ("Malaclemys terrapin"). In America, for example, the members of the genus "Terrapene" dwell on land, yet are referred to as box turtles rather than tortoises.


title: Tortoise

text: British usage, by contrast, tends not to use "turtle" as a generic term for all members of the order, and also applies the term "tortoises" broadly to all land-dwelling members of the order Testudines, regardless of whether they are actually members of the family Testudinidae. In Britain, terrapin is used to refer to a larger group of semiaquatic turtles than the restricted meaning in America.


title: Tortoise

text: Australian usage is different from both American and British usage. Land tortoises are not native to Australia, and traditionally freshwater turtles have been called "tortoises" in Australia. Some Australian experts disapprove of this usage—believing that the term tortoises is "better confined to purely terrestrial animals with very different habits and needs, none of which are found in this country"—and promote the use of the term "freshwater turtle" to describe Australia's primarily aquatic members of the order Testudines because it avoids misleading use of the word "tortoise" and also is a useful distinction from marine turtles.


title: Tortoise

text: Most species of tortoises lay small clutch sizes, seldom exceeding 20 eggs, and many species have clutch sizes of only 1–2 eggs. Incubation is characteristically long in most species, the average incubation period are between 100 and 160.0 days. Egg-laying typically occurs at night, after which the mother tortoise covers her clutch with sand, soil, and organic material. The eggs are left unattended, and depending on the species, take from 60 to 120 days to incubate. The size of the egg depends on the size of the mother and can be estimated by examining the width of the cloacal opening between the carapace and plastron. The plastron of a female tortoise often has a noticeable V-shaped notch below the tail which facilitates passing the eggs. Upon completion of the incubation period, a fully formed hatchling uses an egg tooth to break out of its shell. It digs to the surface of the nest and begins a life of survival on its own. They are hatched with an embryonic egg sac which serves as a source of nutrition for the first three to seven days until they have the strength and mobility to find food. Juvenile tortoises often require a different balance of nutrients than adults, so may eat foods which a more mature tortoise would not. For example, the young of a strictly herbivorous species commonly will consume worms or insect larvae for additional protein.


title: Tortoise

text: The number of concentric rings on the carapace, much like the cross-section of a tree, can sometimes give a clue to how old the animal is, but, since the growth depends highly on the accessibility of food and water, a tortoise that has access to plenty of forage (or is regularly fed by its owner) with no seasonal variation will have no noticeable rings. Moreover, some tortoises grow more than one ring per season, and in some others, due to wear, some rings are no longer visible.


title: Tortoise

text: Tortoises generally have one of the longest lifespans of any animal, and some individuals are known to have lived longer than 150 years. Because of this, they symbolize longevity in some cultures, such as Chinese culture. The oldest tortoise ever recorded, and one of the oldest individual animals ever recorded, was Tu'i Malila, which was presented to the Tongan royal family by the British explorer James Cook shortly after its birth in 1777. Tu'i Malila remained in the care of the Tongan royal family until its death by natural causes on May 19, 1965, at the age of 188.


title: Tortoise

text: The Alipore Zoo in India was the home to Adwaita, which zoo officials claimed was the oldest living animal until its death on March 23, 2006. Adwaita (also spelled Addwaita) was an Aldabra giant tortoise brought to India by Lord Wellesley, who handed it over to the Alipur Zoological Gardens in 1875 when the zoo was set up. West Bengal officials said records showed Adwaita was at least 150 years old, but other evidence pointed to 250. Adwaita was said to be the pet of Robert Clive.


title: Tortoise

text: Harriet was a resident at the Australia Zoo in Queensland from 1987 to her death in 2006; she was believed to have been brought to England by Charles Darwin aboard the "Beagle" and then on to Australia by John Clements Wickham. Harriet died on June 23, 2006, just shy of her 176th birthday.


title: Tortoise

text: Timothy, a female spur-thighed tortoise, lived to be about 165 years old. For 38 years, she was carried as a mascot aboard various ships in Britain's Royal Navy. Then in 1892, at age 53, she retired to the grounds of Powderham Castle in Devon. Up to the time of her death in 2004, she was believed to be the United Kingdom's oldest resident.


title: Tortoise

text: Many species of tortoises are sexually dimorphic, though the differences between males and females vary from species to species. In some species, males have a longer, more protruding neck plate than their female counterparts, while in others, the claws are longer on the females.


title: Tortoise

text: The male plastron is curved inwards to aid reproduction. The easiest way to determine the sex of a tortoise is to look at the tail. The females, as a general rule, have smaller tails, dropped down, whereas the males have much longer tails which are usually pulled up and to the side of the rear shell.


title: Tortoise

text: The brain of a tortoise is extremely small. Red-footed tortoises, from Central and South America, do not have an area in the brain called the hippocampus, which relates to emotion, learning, memory and spatial navigation. Studies have shown that red-footed tortoises may rely on an area of the brain called the medial cortex for emotional actions, an area that humans use for actions such as decision making.


title: Tortoise

text: In the 17th century, Francesco Redi performed an experiment that involved removing the brain of a land tortoise, which then proceeded to live six months. Freshwater tortoises, when subjected to the same experiment, continued similarly, but did not live so long. Redi also cut the head off a tortoise entirely, and it lived for 23 days.


title: Tortoise

text: Tortoises are found from southern North America to southern South America, around the Mediterranean basin, across Eurasia to Southeast Asia, in sub-Saharan Africa, Madagascar, and some Pacific islands. They are absent from Australasia. They live in diverse habitats, including deserts, arid grasslands, and scrub to wet evergreen forests, and from sea level to mountains. Most species, however, occupy semiarid habitats.


title: Tortoise

text: Many large islands are or were characterized by species of giant tortoises. Part of the reason for this is that tortoises are good at oceanic dispersal. Despite being unable to swim, tortoises are able to survive long periods adrift at sea because they can survive months without food or fresh water. Tortoises have been known to survive oceanic dispersals of more than 740 km. Once on islands tortoises faced few predators or competitors and could grow to large sizes and become the dominant large herbivores on many islands due to their low metabolic rate and reduced need for fresh water compared to mammals.


title: Tortoise

text: Today there are only two living species of giant tortoises, the Aldabra giant tortoise on Aldabra Atoll and the several species/subspecies of Galapagos giant tortoise found on the Galapagos Islands. However, until recently giant tortoises could be found on nearly every major island group, including the Bahamas, the Greater Antilles (including Cuba and Hispaniola), the Lesser Antilles, the Canary Islands, Malta, the Seychelles, the Mascarene Islands (including Mauritius and Reunion), and Madagascar. Most of these tortoises were wiped out by human arrival. Many of these giant tortoises are not closely related (belonging to different genera such as "Megalochelys", "Chelonoidis", "Centrochelys", "Aldabrachelys", "Cylindraspis", and "Hesperotestudo"), but are thought to have independently evolved large body size through convergent evolution. Giant tortoises are notably absent from Australasia and many south Pacific islands, but the distantly related meiolaniid turtles are thought to have filled the same niche. Giant tortoises are also known from the Oligocene-Pliocene of mainland North America, South America, Europe, Asia, and Africa, but are all now extinct, which is also attributed to human activity.


title: Tortoise

text: Tortoises are generally considered to be strict herbivores, feeding on grasses, weeds, leafy greens, flowers, and some fruits. However, hunting and eating of birds has been observed on occasion. Pet tortoises typically require diets based on wild grasses, weeds, leafy greens and certain flowers. Certain species consume worms or insects and carrion in their normal habitats. Too much protein is detrimental in herbivorous species, and has been associated with shell deformities and other medical problems. Different tortoise species vary greatly in their nutritional requirements.


title: Tortoise

text: Communication in tortoises is different from many other reptiles. Because they are restricted by their shell and short limbs, visual communication is not a strong form of communication in tortoises. Tortoises use olfactory cues to determine the sex of other tortoises so that they can find a potential mate. Tactile communication is important in tortoises during combat and courtship. In both combat and courtship, tortoises use ramming to communicate with other individuals.


title: Tortoise

text: This species list largely follows Turtle Taxonomy Working Group (2021) and the Turtle Extinctions Working Group (2015).


title: Tortoise

text: In Hinduism, Kurma () was the second Avatar of Vishnu. Like the Matsya Avatara, Kurma also belongs to the Satya Yuga. Vishnu took the form of a half-man, half-tortoise, the lower half being a tortoise. He is normally shown as having four arms. He sat on the bottom of the ocean after the Great Flood. A mountain was placed on his back by the other gods so they could churn the sea and find the ancient treasures of the Vedic peoples.


title: Tortoise

text: In September, 1968, two Russian tortoises became the first animals to fly to and circle the Moon. Their Zond 5 mission brought them back to Earth safely.


title: Tortoise

text: Teenage Mutant Ninja Turtles is an entertainment franchise. The four anthropomorphic turtle brothers trained in ninjitsu are shown to fight evil in New York City.


title: Coelacanth

text: The coelacanths ( ) are fish belonging to the order Actinistia that includes two extant species in the genus "Latimeria": the West Indian Ocean coelacanth ("Latimeria chalumnae"), primarily found near the Comoro Islands off the east coast of Africa, and the Indonesian coelacanth ("Latimeria menadoensis"). The name "coelacanth" originates from the Permian genus "Coelacanthus", which was the first scientifically named coelacanth.


title: Coelacanth

text: Coelacanths follow the oldest-known living lineage of Sarcopterygii (lobe-finned fish and tetrapods), which means they are more closely related to lungfish and tetrapods (which includes amphibians, reptiles, birds and mammals) than to ray-finned fish. They are found along the coastline of Indonesia and in the Indian Ocean. The West Indian Ocean coelacanth is a critically endangered species.


title: Coelacanth

text: The oldest known coelacanth fossils are over 410 million years old. Coelacanths were thought to have become extinct in the Late Cretaceous, around 66 million years ago, but were discovered living off the coast of South Africa in 1938.


title: Coelacanth

text: The coelacanth was long considered a "living fossil" because scientists thought it was the sole remaining member of a taxon otherwise known only from fossils, with no close relations alive, and that it evolved into roughly its current form approximately 400 million years ago. However, several more recent studies have shown that coelacanth body shapes are much more diverse than previously thought.


title: Coelacanth

text: The word "Coelacanth" is an adaptation of the Modern Latin "Cœlacanthus" ("hollow spine"), from the Greek κοῖλ-ος (, "hollow") and ἄκανθ-α (, "spine"), referring to the hollow caudal fin rays of the first fossil specimen described and named by Louis Agassiz in 1839, belonging to the genus "Coelacanthus". The genus name "Latimeria" commemorates Marjorie Courtenay-Latimer, who discovered the first specimen.


title: Coelacanth

text: The earliest fossils of coelacanths were discovered in the 19th century. Coelacanths, which are related to lungfishes and tetrapods, were believed to have become extinct at the end of the Cretaceous period. More closely related to tetrapods than to the ray-finned fish, coelacanths were considered transitional species between fish and tetrapods. On 23 December 1938, the first "Latimeria" specimen was found off the east coast of South Africa, off the Chalumna River (now Tyolomnqa). Museum curator Marjorie Courtenay-Latimer discovered the fish among the catch of a local fisherman. Courtenay-Latimer contacted a Rhodes University ichthyologist, J. L. B. Smith, sending him drawings of the fish, and he confirmed the fish's importance with a famous cable: "Most Important Preserve Skeleton and Gills = Fish Described."


title: Coelacanth

text: Its discovery 66 million years after its supposed extinction makes the coelacanth the best-known example of a Lazarus taxon, an evolutionary line that seems to have disappeared from the fossil record only to reappear much later. Since 1938, West Indian Ocean coelacanth have been found in the Comoros, Kenya, Tanzania, Mozambique, Madagascar, in iSimangaliso Wetland Park, and off the South Coast of Kwazulu-Natal in South Africa.


title: Coelacanth

text: The Comoro Islands specimen was discovered in December 1952. Between 1938 and 1975, 84 specimens were caught and recorded.


title: Coelacanth

text: The second extant species, the Indonesian coelacanth, was described from Manado, North Sulawesi, Indonesia in 1999 by Pouyaud et al. based on a specimen discovered by Mark V. Erdmann in 1998 and deposited at the Indonesian Institute of Sciences (LIPI). Erdmann and his wife Arnaz Mehta first encountered a specimen at a local market in September 1997, but took only a few photographs of the first specimen of this species before it was sold. After confirming that it was a unique discovery, Erdmann returned to Sulawesi in November 1997 to interview fishermen and look for further examples. A second specimen was caught by a fisherman in July 1998, which was then handed to Erdmann.


title: Coelacanth

text: "Latimeria chalumnae" and "L. menadoensis" are the only two known living coelacanth species. Coelacanths are large, plump, lobe-finned fish that can grow to more than and weigh around . They are estimated to live up to 100 years, based on analysis of annual growth marks on scales, and reach maturity around the age of 55; the oldest known specimen was 84 years old at the time of its capture in 1960.


title: Coelacanth

text: Even though their estimated lifetime is similar to humans, gestation can last 5 years, which is 1.5 years more than the deep-sea frilled shark, the previous record holder. They are nocturnal piscivorous drift-hunters. The body is covered in ctenoid elasmoid scales that act as armor. Coelacanths have eight fins – two dorsal fins, two pectoral fins, two pelvic fins, one anal fin and one caudal fin. The tail is very nearly equally proportioned and is split by a terminal tuft of fin rays that make up its caudal lobe. The eyes of the coelacanth are very large, while the mouth is very small. The eye is acclimatized to seeing in poor light by rods that absorb mostly short wavelengths. Coelacanth vision has evolved to a mainly blue-shifted color capacity. Pseudomaxillary folds surround the mouth and replace the maxilla, a structure absent in coelacanths. Two nostrils, along with four other external openings, appear between the premaxilla and lateral rostral bones. The nasal sacs resemble those of many other fish and do not contain an internal nostril. The coelacanth's rostral organ, contained within the ethmoid region of the braincase, has three unguarded openings into the environment and is used as a part of the coelacanth's laterosensory system. The coelacanth's auditory reception is mediated by its inner ear, which is very similar to that of tetrapods and is classified as being a basilar papilla.


title: Coelacanth

text: Coelacanths are a part of the clade Sarcopterygii, or the lobe-finned fishes. Externally, several characteristics distinguish coelacanths from other lobe-finned fish. They possess a three-lobed caudal fin, also called a trilobate fin or a diphycercal tail. A secondary tail extending past the primary tail separates the upper and lower halves of the coelacanth. ctenoid elasmoid scales act as thick armor to protect the coelacanth's exterior. Several internal traits also aid in differentiating coelacanths from other lobe-finned fish. At the back of the skull, the coelacanth possesses a hinge, the intracranial joint, which allows it to open its mouth extremely wide. Coelacanths also retain an oil-filled notochord, a hollow, pressurized tube which is replaced by a vertebral column early in embryonic development in most other vertebrates. The coelacanth's heart is shaped differently from that of most modern fish, with its chambers arranged in a straight tube. The coelacanth's braincase is 98.5% filled with fat; only 1.5% of the braincase contains brain tissue. The cheeks of the coelacanth are unique because the opercular bone is very small and holds a large soft-tissue opercular flap. A spiracular chamber is present, but the spiracle is closed and never opens during development. Also unique to extant coelacanths is the presence of a "fatty lung" or a fat-filled single-lobed vestigial lung, homologous to other fishes' swim bladders. The parallel development of a fatty organ for buoyancy control suggests a unique specialization for deep-water habitats. There are small and hard—but-flexible—plates around the vestigial lung in adult specimens, though not around the fatty organ. The plates most likely had a regulation function for the volume of the lung. Due to the size of the fatty organ, researchers assume that it is responsible for the kidney's unusual relocation. The two kidneys, which are fused into one, are located ventrally within the abdominal cavity, posterior to the cloaca.


title: Coelacanth

text: In 2013, a research group published the genome sequence of the coelacanth in the scientific journal "Nature".


title: Coelacanth

text: Due to their lobed fins and other features, it was once hypothesized that the coelacanth might be the most recent shared ancestor between terrestrial and marine vertebrates. But after sequencing the full genome of the coelacanth, it was discovered that the lungfish is the most recent shared ancestor. Coelacanths and lungfish had already diverged from a common ancestor before the lungfish made the transition to land.


title: Coelacanth

text: Another important discovery made from the genome sequencing is that the coelacanths are still evolving today. While phenotypic similarity between extant and extinct coelacanths suggests there is limited evolutionary pressure on these organisms to undergo morphological divergence, they are undergoing measurable genetic divergence. Despite prior studies showing that protein coding regions are undergoing evolution at a substitution rate much lower than other tetrapods (consistent with phenotypic stasis observed between extant and fossil members of the taxa), the non-coding regions subject to higher transposable element activity show marked divergence even between the two extant coelacanth species. This has been facilitated in part by a coelacanth-specific endogenous retrovirus of the Epsilon retrovirus family.


title: Coelacanth

text: According to the fossil record, the divergence of coelacanths, lungfish and tetrapods is thought to have occurred during the Silurian. Over 100 fossil species of coelacanth have been described. The oldest identified coelacanth fossils are around 420-410 million years old, dating to the early Devonian. Coelacanths were never a diverse group in comparison to other groups of fish, and reached a peak diversity during the Early Triassic (252-247 million years ago), coinciding with a burst of diversification between the Late Permian and Middle Triassic. Most Mesozoic coelacanths belong to the order Latimerioidei, which contains two major subdivisions, the marine Latimeriidae, which contains modern coelacanths, as well as the extinct Mawsoniidae, which were native to brackish, freshwater as well as marine environments.


title: Coelacanth

text: Paleozoic coelacanths are generally small (~30–40 cm in length), while Mesozoic forms were larger. Several specimens belonging to the Jurassic and Creataceous mawsoniid coelcanth genera "Trachymetopon" and "Mawsonia" likely reached or exceeded in length, making them amongst the largest known fishes of the Mesozoic, and amongst the largest bony fishes of all time.


title: Coelacanth

text: The most recent fossil latimeriid is "Megalocoelacanthus dobiei", whose disarticulated remains are found in late Santonian to middle Campanian, and possibly earliest Maastrichtian-aged marine strata of the Eastern and Central United States, the most recent mawsoniids are "Axelrodichthys megadromos" from early Campanian to early Maastrichtian freshwater continental deposits of France., as well as an indeterminate marine mawsoniid from Morocco, dating to the late Maastrichtian A small bone fragment from the European Paleocene has been considered the only plausible post-Cretaceous record, but this identification is based on comparative bone histology methods of doubtful reliability.


title: Coelacanth

text: Living coelacanths have been considered "living fossils" based on their supposedly conservative morphology relative to fossil species; however, recent studies have expressed the view that coelacanth morphologic conservatism is a belief not based on data. Fossils suggest that coelacanths were most morphologically diverse during the Devonian and Carboniferous, while Mesozoic species are generally morphologically similar to each other.


title: Coelacanth

text: The current coelacanth range is primarily along the eastern African coast, although "Latimeria menadoensis" was discovered off Indonesia. Coelacanths have been found in the waters of Kenya, Tanzania, Mozambique, South Africa, Madagascar, Comoros and Indonesia. Most "Latimeria chalumnae" specimens that have been caught have been captured around the islands of Grande Comore and Anjouan in the Comoros Archipelago (Indian Ocean). Though there are cases of "L. chalumnae" caught elsewhere, amino acid sequencing has shown no big difference between these exceptions and those found around Comore and Anjouan. Even though these few may be considered strays, there are several reports of coelacanths being caught off the coast of Madagascar. This leads scientists to believe that the endemic range of "Latimeria chalumnae" coelacanths stretches along the eastern coast of Africa from the Comoros Islands, past the western coast of Madagascar to the South African coastline. Mitochondrial DNA sequencing of coelacanths caught off the coast of southern Tanzania suggests a divergence of the two populations some 200,000 years ago. This could refute the theory that the Comoros population is the main population while others represent recent offshoots. A live specimen was seen and recorded on video in November 2019 at 69 m off the village of Umzumbe on the South Coast of KwaZulu-Natal, about 325 km south of the iSimangaliso Wetland Park. This is the farthest south since the original discovery, and the second shallowest record after 54 m in the Diepgat Canyon. These sightings suggest that they may live shallower than previously thought, at least at the southern end of their range, where colder, better-oxygenated water is available at shallower depths.


title: Coelacanth

text: The geographical range of the Indonesia coelacanth, "Latimeria menadoensis", is believed to be off the coast of Manado Tua Island, Sulawesi, Indonesia in the Celebes Sea. Key components confining coelacanths to these areas are food and temperature restrictions, as well as ecological requirements such as caves and crevices that are well-suited for drift feeding. Teams of researchers using submersibles have recorded live sightings of the fish in the Sulawesi Sea as well as in the waters of Biak in Papua.


title: Coelacanth

text: Anjouan Island and the Grande Comore provide ideal underwater cave habitats for coelacanths. The islands' underwater volcanic slopes, steeply eroded and covered in sand, house a system of caves and crevices which allow coelacanths resting places during the daylight hours. These islands support a large benthic fish population that helps to sustain coelacanth populations.


title: Coelacanth

text: During the daytime, coelacanths rest in caves anywhere from 100 to 500 meters deep. Others migrate to deeper waters. The cooler waters (below 120 meters) reduce the coelacanths' metabolic costs. Drifting toward reefs and night feeding saves vital energy. Resting in caves during the day also saves energy that otherwise would be expended to fight currents.


title: Coelacanth

text: Coelacanth locomotion is unique. To move around they most commonly take advantage of up- or down-wellings of current and drift. Their paired fins stabilize movement through the water. While on the ocean floor, they do not use the paired fins for any kind of movement. Coelacanths generate thrust with their caudal fins for quick starts. Due to the abundance of its fins, the coelacanth has high maneuverability and can orient its body in almost any direction in the water. They have been seen doing headstands as well as swimming belly up. It is thought that the rostral organ helps give the coelacanth electroreception, which aids in movement around obstacles.


title: Coelacanth

text: Coelacanths are fairly peaceful when encountering others of their kind, remaining calm even in a crowded cave. They do avoid body contact, however, withdrawing immediately if contact occurs. When approached by foreign potential predators (e.g. a submersible), they show panic flight reactions, suggesting that coelacanths are most likely prey to large deepwater predators. Shark bite marks have been seen on coelacanths; sharks are common in areas inhabited by coelacanths. Electrophoresis testing of 14 coelacanth enzymes shows little genetic diversity between coelacanth populations. Among the fish that have been caught were about equal numbers of males and females. Population estimates range from 210 individuals per population to 500 per population. Because coelacanths have individual color markings, scientists think that they recognize other coelacanths via electric communication.


title: Coelacanth

text: Coelacanths are nocturnal piscivores that feed mainly on benthic smaller fish and various cephalopods. They are "passive drift feeders", slowly drifting along currents with only minimal self-propulsion, eating whatever prey they encounter. Coelacanths also use their rostral organ for its electroreption to be able to detect nearby prey in low light settings.


title: Coelacanth

text: Coelacanths are ovoviviparous, meaning that the female retains the fertilized eggs within her body while the embryos develop during a gestation period of five years. Typically, females are larger than the males; their scales and the skin folds around the cloaca differ. The male coelacanth has no distinct copulatory organs, just a cloaca, which has a urogenital papilla surrounded by erectile caruncles. It is hypothesized that the cloaca everts to serve as a copulatory organ.


title: Coelacanth

text: Coelacanth eggs are large, with only a thin layer of membrane to protect them. Embryos hatch within the female and eventually are born alive, which is a rarity in fish. This was only discovered when the American Museum of Natural History dissected its first coelacanth specimen in 1975 and found it pregnant with five embryos. Young coelacanths resemble the adult, the main differences being an external yolk sac, larger eyes relative to body size and a more pronounced downward slope of the body. The juvenile coelacanth's broad yolk sac hangs below the pelvic fins. The scales and fins of the juvenile are completely matured; however, it does lack odontodes, which it gains during maturation.


title: Coelacanth

text: A study that assessed the paternity of the embryos inside two coelacanth females indicated that each clutch was sired by a single male. This could mean that females mate monandrously, i.e. with one male only. Polyandry, female mating with multiple males, is common in both plants and animals and can be advantageous (e.g. insurance against mating with an infertile or incompatible mate), but also confers costs (increased risk of infection, danger of falling prey to predators, increased energy input when searching for new males).


title: Coelacanth

text: Because little is known about the coelacanth, the conservation status is difficult to characterize. According to Fricke et al. (1995), it is important to conserve the species. From 1988 to 1994, Fricke counted some 60 individuals of "L. chalumnae" on each dive. In 1995 that number dropped to 40. Even though this could be a result of natural population fluctuation, it also could be a result of overfishing. The IUCN currently classifies "L. chalumnae" as "critically endangered", with a total population size of 500 or fewer individuals. "L. menadoensis" is considered Vulnerable, with a significantly larger population size (fewer than 10,000 individuals).


title: Coelacanth

text: The major threat towards the coelacanth is the accidental capture by fishing operations, especially commercial deep-sea trawling. Coelacanths usually are caught when local fishermen are fishing for oilfish. Fishermen sometimes snag a coelacanth instead of an oilfish because they traditionally fish at night, when oilfish (and coelacanths) feed.


title: Coelacanth

text: Before scientists became interested in coelacanths, they were thrown back into the water if caught. Now that they are recognized as important, fishermen trade them to scientists or other officials. Before the 1980s, this was a problem for coelacanth populations. In the 1980s, international aid gave fiberglass boats to the local fishermen, which moved fishing beyond the coelacanth territories into more productive waters. Since then, most of the motors on the boats failed, forcing the fishermen back into coelacanth territory and putting the species at risk again.


title: Coelacanth

text: Methods to minimize the number of coelacanths caught include moving fishers away from the shore, using different laxatives and malarial salves to reduce the demand for oilfish, using coelacanth models to simulate live specimens, and increasing awareness of the need for conservation. In 1987 the Coelacanth Conservation Council advocated the conservation of coelacanths. The CCC has branches located in Comoros, South Africa, Canada, the United Kingdom, the U.S., Japan, and Germany. The agencies were established to help protect and encourage population growth of coelacanths.


title: Coelacanth

text: A "deep release kit" was developed in 2014 and distributed by private initiative, consisting of a weighted hook assembly that allows a fisherman to return an accidentally caught coelacanth to deep waters where the hook can be detached once it hits the seafloor. Conclusive reports about the effectiveness of this method are still pending.


title: Coelacanth

text: In 2002, the South African Coelacanth Conservation and Genome Resource Programme was launched to help further the studies and conservation of the coelacanth. This program focuses on biodiversity conservation, evolutionary biology, capacity building, and public understanding. The South African government committed to spending R10 million on the program. In 2011, a plan was made for a Tanga Coelacanth Marine Park to conserve biodiversity for marine animals including the coelacanth. The park was designed to reduce habitat destruction and improve prey availability for endangered species.


title: Coelacanth

text: Coelacanths are considered a poor source of food for humans and likely most other fish-eating animals. Coelacanth flesh has large amounts of oil, urea, wax esters, and other compounds that give the flesh a distinctly unpleasant flavor, make it difficult to digest, and can cause diarrhea. Their scales themselves secrete mucus, which combined with the excessive oil their bodies produce, make coelacanths a slimy food. Where the coelacanth is more common, local fishermen avoid it because of its potential to sicken consumers. As a result, the coelacanth has no real commercial value apart from being coveted by museums and private collectors.


title: Coelacanth

text: Because of the surprising nature of the coelacanth's discovery, they have been a frequent source of inspiration in modern artwork, craftsmanship, and literature. At least 22 countries have depicted them on their postage stamps, particularly the Comoros, which has issued 12 different sets of coelacanth stamps. The coelacanth is also depicted on the 1000 Comorian franc banknote, as well as the 5 CF coin.


title: Alfred Nobel

text: Alfred Bernhard Nobel ( , ; 21 October 1833 – 10 December 1896) was a Swedish chemist, engineer, inventor, businessman, and philanthropist. He is best known for having bequeathed his fortune to establish the Nobel Prize, though he also made several important contributions to science, holding 355 patents in his lifetime. Nobel's most famous invention was dynamite, a safer and easier means of harnessing the explosive power of nitroglycerin; it was patented in 1867.


title: Alfred Nobel

text: Nobel displayed an early aptitude for science and learning, particularly in chemistry and languages; he became fluent in six languages and filed his first patent at age 24. He embarked on many business ventures with his family, most notably owning Bofors, an iron and steel producer that he developed into a major manufacturer of cannons and other armaments.


title: Alfred Nobel

text: Nobel was later inspired to donate his fortune to the Nobel Prize institution, which would annually recognize those who "conferred the greatest benefit to humankind". The synthetic element nobelium was named after him, and his name and legacy also survives in companies such as Dynamit Nobel and AkzoNobel, which descend from mergers with companies he founded.


title: Alfred Nobel

text: Nobel was elected a member of the Royal Swedish Academy of Sciences, which, pursuant to his will, would be responsible for choosing the Nobel laureates in physics and in chemistry.


title: Alfred Nobel

text: Alfred Nobel was born in Stockholm, United Kingdoms of Sweden and Norway on 21 October 1833. He was the third son of Immanuel Nobel (1801–1872), an inventor and engineer, and Karolina Andriette Nobel (née Ahlsell 1805–1889). The couple married in 1827 and had eight children. The family was impoverished and only Alfred and his three brothers survived beyond childhood. Through his father, Alfred Nobel was a descendant of the Swedish scientist Olaus Rudbeck (1630–1702), and in his turn, the boy was interested in engineering, particularly explosives, learning the basic principles from his father at a young age. Alfred Nobel's interest in technology was inherited from his father, an alumnus of Royal Institute of Technology in Stockholm.Following various business failures, Nobel's father moved to Saint Petersburg, Russia and grew successful there as a manufacturer of machine tools and explosives. He invented the veneer lathe (which made possible the production of modern plywood) and started work on the torpedo. In 1842, the family joined him in the city. Now prosperous, his parents were able to send Nobel to private tutors and the boy excelled in his studies, particularly in chemistry and languages, achieving fluency in English, French, German and Russian. For 18 months, from 1841 to 1842, Nobel went to the only school he ever attended as a child, in Stockholm.


title: Alfred Nobel

text: Nobel gained proficiency in Swedish, French, Russian, English, German, and Italian. He also developed sufficient literary skill to write poetry in English. His "Nemesis" is a prose tragedy in four acts about the Italian noblewoman Beatrice Cenci. It was printed while he was dying, but the entire stock was destroyed immediately after his death except for three copies, being regarded as scandalous and blasphemous. It was published in Sweden in 2003 and has been translated into Slovenian, French, Italian, and Spanish.


title: Alfred Nobel

text: Nobel was Lutheran and regularly attended the Church of Sweden Abroad during his Paris years, led by pastor Nathan Söderblom who received the Nobel Peace Prize in 1930. He became an agnostic in youth and was an atheist later in life, though still donated generously to the Church.


title: Alfred Nobel

text: Nobel travelled for much of his business life, maintaining companies in Europe and America while keeping a home in Paris from 1873 to 1891. He remained a solitary character, given to periods of depression. He remained unmarried, although his biographers note that he had at least three loves, the first in Russia with a girl named Alexandra who rejected his proposal. In 1876, Austro-Bohemian Countess Bertha Kinsky became his secretary, but she left him after a brief stay to marry her previous lover Baron Arthur Gundaccar von Suttner. Her contact with Nobel was brief, yet she corresponded with him until his death in 1896, and probably influenced his decision to include a peace prize in his will. She was awarded the 1905 Nobel Peace prize "for her sincere peace activities". Nobel's longest-lasting relationship was with Sofija Hess from Celje whom he met in 1876. The liaison lasted for 18 years.


title: Alfred Nobel

text: In the years of 1865 to 1873, Alfred Nobel had his home in Krümmel, Hamburg, he afterward moved to a house in the Avenue Malakoff in Paris that same year.


title: Alfred Nobel

text: In 1894, when he acquired Bofors-Gullspång, the Björkborn Manor was included, he stayed at his manor house in Sweden during the summers. The manor house became his very last residence in Sweden and has after his death functioned as a museum.


title: Alfred Nobel

text: Alfred Nobel died on 10 December 1896, in Sanremo, Italy, at his very last residence, Villa Nobel, overlooking the Mediterranean Sea.


title: Alfred Nobel

text: As a young man, Nobel studied with chemist Nikolai Zinin; then, in 1850, went to Paris to further the work. There he met Ascanio Sobrero, who had invented nitroglycerin three years before. Sobrero strongly opposed the use of nitroglycerin because it was unpredictable, exploding when subjected to variable heat or pressure. But Nobel became interested in finding a way to control and use nitroglycerin as a commercially usable explosive; it had much more power than gunpowder. In 1851 at age 18, he went to the United States for one year to study, working for a short period under Swedish-American inventor John Ericsson, who designed the American Civil War ironclad, USS "Monitor". Nobel filed his first patent, an English patent for a gas meter, in 1857, while his first Swedish patent, which he received in 1863, was on "ways to prepare gunpowder".The family factory produced armaments for the Crimean War (1853–1856), but had difficulty switching back to regular domestic production when the fighting ended and they filed for bankruptcy. In 1859, Nobel's father left his factory in the care of the second son, Ludvig Nobel (1831–1888), who greatly improved the business. Nobel and his parents returned to Sweden from Russia and Nobel devoted himself to the study of explosives, and especially to the safe manufacture and use of nitroglycerin. Nobel invented a detonator in 1863, and in 1865 designed the blasting cap.


title: Alfred Nobel

text: On 3 September 1864, a shed used for preparation of nitroglycerin exploded at the factory in Heleneborg, Stockholm, Sweden, killing five people, including Nobel's younger brother Emil. Fazed by the accident, Nobel founded the company Nitroglycerin Aktiebolaget AB in Vinterviken so that he could continue to work in a more isolated area. Nobel invented dynamite in 1867, a substance easier and safer to handle than the more unstable nitroglycerin. Dynamite was patented in the US and the UK and was used extensively in mining and the building of transport networks internationally. In 1875, Nobel invented gelignite, more stable and powerful than dynamite, and in 1887, patented ballistite, a predecessor of cordite.


title: Alfred Nobel

text: Nobel was elected a member of the Royal Swedish Academy of Sciences in 1884, the same institution that would later select laureates for two of the Nobel prizes, and he received an honorary doctorate from Uppsala University in 1893.


title: Alfred Nobel

text: Nobel's brothers Ludvig and Robert founded the oil company Branobel and became hugely rich in their own right. Nobel invested in these and amassed great wealth through the development of these new oil regions. During his life, Nobel was issued 355 patents internationally, and by his death, his business had established more than 90 armaments factories, despite his apparently pacifist character.


title: Alfred Nobel

text: Nobel found that when nitroglycerin was incorporated in an absorbent inert substance like "kieselguhr" (diatomaceous earth) it became safer and more convenient to handle, and this mixture he patented in 1867 as "dynamite". Nobel demonstrated his explosive for the first time that year, at a quarry in Redhill, Surrey, England. In order to help reestablish his name and improve the image of his business from the earlier controversies associated with dangerous explosives, Nobel had also considered naming the highly powerful substance "Nobel's Safety Powder", but settled with Dynamite instead, referring to the Greek word for "power" ().


title: Alfred Nobel

text: Nobel later combined nitroglycerin with various nitrocellulose compounds, similar to collodion, but settled on a more efficient recipe combining another nitrate explosive, and obtained a transparent, jelly-like substance, which was a more powerful explosive than dynamite. Gelignite, or blasting gelatine, as it was named, was patented in 1876; and was followed by a host of similar combinations, modified by the addition of potassium nitrate and various other substances. Gelignite was more stable, transportable and conveniently formed to fit into bored holes, like those used in drilling and mining, than the previously used compounds. It was adopted as the standard technology for mining in the "Age of Engineering", bringing Nobel a great amount of financial success, though at a cost to his health. An offshoot of this research resulted in Nobel's invention of ballistite, the precursor of many modern smokeless powder explosives and still used as a rocket propellant.


title: Alfred Nobel

text: There is a well known story about the origin of the Nobel Prize. In 1888, the death of his brother Ludvig supposedly caused several newspapers to publish obituaries of Alfred in error. One French newspaper condemned him for his invention of military explosives—in many versions of the story, dynamite is quoted, although this was mainly used for civilian applications—and this is said to have brought about his decision to leave a better legacy after his death. The obituary stated, "" ("The merchant of death is dead"), and went on to say, "Dr. Alfred Nobel, who became rich by finding ways to kill more people faster than ever before, died yesterday." Nobel read the obituary and was appalled at the idea that he would be remembered in this way. His decision to posthumously donate the majority of his wealth to found the Nobel Prize has been credited to him wanting to leave behind a better legacy. However, it has been questioned whether or not the obituary in question actually existed.


title: Alfred Nobel

text: On 27 November 1895, at the Swedish-Norwegian Club in Paris, Nobel signed his last will and testament and set aside the bulk of his estate to establish the Nobel Prizes, to be awarded annually without distinction of nationality. After taxes and bequests to individuals, Nobel's will allocated 94% of his total assets, 31,225,000 Swedish kronor, to establish the five Nobel Prizes. This converted to £1,687,837 (GBP) at the time. In 2012, the capital was worth around SEK 3.1 billion (US$472 million, EUR 337 million), which is almost twice the amount of the initial capital, taking inflation into account.


title: Alfred Nobel

text: The first three of these prizes are awarded for eminence in physical science, in chemistry and in medical science or physiology; the fourth is for literary work "in an ideal direction" and the fifth prize is to be given to the person or society that renders the greatest service to the cause of international fraternity, in the suppression or reduction of standing armies, or in the establishment or furtherance of peace congresses.


title: Alfred Nobel

text: The formulation for the literary prize being given for a work "in an ideal direction" (' in Swedish), is cryptic and has caused much confusion. For many years, the Swedish Academy interpreted "ideal" as "idealistic" (') and used it as a reason not to give the prize to important but less romantic authors, such as Henrik Ibsen and Leo Tolstoy. This interpretation has since been revised, and the prize has been awarded to, for example, Dario Fo and José Saramago, who do not belong to the camp of literary idealism.


title: Alfred Nobel

text: There was room for interpretation by the bodies he had named for deciding on the physical sciences and chemistry prizes, given that he had not consulted them before making the will. In his one-page testament, he stipulated that the money go to discoveries or inventions in the physical sciences and to discoveries or improvements in chemistry. He had opened the door to technological awards, but had not left instructions on how to deal with the distinction between science and technology. Since the deciding bodies he had chosen were more concerned with the former, the prizes went to scientists more often than engineers, technicians or other inventors.


title: Alfred Nobel

text: Sweden's central bank Sveriges Riksbank celebrated its 300th anniversary in 1968 by donating a large sum of money to the Nobel Foundation to be used to set up a sixth prize in the field of economics in honour of Alfred Nobel. In 2001, Alfred Nobel's great-great-nephew, Peter Nobel (born 1931), asked the Bank of Sweden to differentiate its award to economists given "in Alfred Nobel's memory" from the five other awards. This request added to the controversy over whether the Bank of Sweden Prize in Economic Sciences in Memory of Alfred Nobel is actually a legitimate "Nobel Prize".


title: Alfred Nobel

text: Nobel was accused of high treason against France for selling Ballistite to Italy, so he moved from Paris to Sanremo, Italy in 1891. On 10 December 1896, he suffered a stroke and died. He had left most of his wealth in trust, unbeknownst to his family, in order to fund the Nobel Prize awards. He is buried in Norra begravningsplatsen in Stockholm.


title: Alfred Nobel

text: The "Monument to Alfred Nobel" (, ) in Saint Petersburg is located along the Bolshaya Nevka River on Petrogradskaya Embankment. It was dedicated in 1991 to mark the 90th anniversary of the first Nobel Prize presentation. Diplomat Thomas Bertelman and Professor Arkady Melua were initiators of the creation of the monument (1989). Professor A. Melua has provided funds for the establishment of the monument (J.S.Co. "Humanistica", 1990–1991). The abstract metal sculpture was designed by local artists Sergey Alipov and Pavel Shevchenko, and appears to be an explosion or branches of a tree. Petrogradskaya Embankment is the street where Nobel's family lived until 1859.


title: Alfred Nobel

text: Criticism of Nobel focuses on his leading role in weapons manufacturing and sales, and some question his motives in creating his prizes, suggesting they are intended to improve his reputation.


title: Quadrilateral Security Dialogue

text: The Quadrilateral Security Dialogue (QSD), commonly known as the Quad, is a strategic security dialogue between Australia, India, Japan, and the United States that is maintained by talks between member countries. The dialogue was initiated in 2007 by Japanese Prime Minister Shinzo Abe, with the support of Australian Prime Minister John Howard, Indian Prime Minister Manmohan Singh, and U.S. Vice President Dick Cheney. The dialogue was paralleled by joint military exercises of an unprecedented scale, titled Exercise Malabar. The diplomatic and military arrangement was widely viewed as a response to increased Chinese economic and military power, and the Chinese government responded to the Quadrilateral dialogue by issuing formal diplomatic protests to its members, calling it "Asian NATO", Indian Foreign Minister S. Jaishankar denies China's allegations and claimed India never had ‘NATO mentality’.


title: Quadrilateral Security Dialogue

text: The Quad ceased following the withdrawal of Australia during Kevin Rudd's tenure as prime minister, reflecting ambivalence in Australian policy over the growing tension between the United States and China in the Asia-Pacific. Following Rudd's replacement by Julia Gillard in 2010, enhanced military cooperation between the United States and Australia was resumed, leading to the placement of U.S. Marines near Darwin, Australia, overlooking the Timor Sea and Lombok Strait. Meanwhile, India, Japan, and the United States continued to hold joint naval exercises under Malabar.


title: Quadrilateral Security Dialogue

text: During the 2017 ASEAN Summits in Manila, all four former members led by Abe, Australian Prime Minister Malcolm Turnbull, Indian Prime Minister Narendra Modi, and US President Donald Trump agreed to revive the quadrilateral alliance in order to counter China militarily and diplomatically in the "Indo-Pacific" region, particularly in the South China Sea. Tensions between Quad members and China have led to fears of what was dubbed by some commentators "a new Cold War" in the region.


title: Quadrilateral Security Dialogue

text: In a joint statement in March 2021, "The Spirit of the Quad," the Quad members described "a shared vision for a Free and Open Indo-Pacific," and a "rules-based maritime order in the East and South China seas," which the Quad members state are needed to counter Chinese maritime claims. The Quad pledged to respond to COVID-19, and held a first Quad Plus meeting that included representatives from South Korea, New Zealand, and Vietnam to work on its response to it. As of 2022, Quad countries together have a combined GDP nominal of US$34.5 trillion (33.9% of the gross world product) and a combined GDP PPP of US$44.4 (27.5% of the world's GDP).


title: Quadrilateral Security Dialogue

text: In the early twenty-first century, the strategic preoccupation of the United States with Iraq and Afghanistan served as a distraction from major power shifts in the Asia-Pacific, brought about by increased Chinese economic power, which undermined America's traditional role in the region. In the long term the United States has sought a policy of "soft containment" of China by organizing strategic partnerships with democracies at its periphery. While US alliances with Japan, Australia and India now form the bulwark of this policy, the development of closer US military ties to India has been a complex process since the collapse of the Soviet Union. Australian commentaries showed mixed attitudes to a Quadrilateral security arrangement isolating China.


title: Quadrilateral Security Dialogue

text: Active US-Indian military cooperation expanded in 1991 following the economic liberalization of India when American Lt. General Claude C. Kicklighter, then commander of the United States Army Pacific, proposed army-to-army cooperation. This cooperation further expanded in the mid-1990s under an early Indian centre-right coalition, and in 2001 India offered the United States military facilities within its territory for offensive operations in Afghanistan. US Defense Secretary Donald Rumsfeld and his Indian counterpart Pranab Mukherjee signed a "New Framework for India-US Defense" in 2005 under the Indian United Progressive Alliance government, increasing cooperation regarding military relations, defence industry and technology sharing, and the establishment of a "Framework on maritime security cooperation." India and the United States conducted dozens of joint military exercises in the ensuing years before the development of the Quadrilateral dialogue, interpreted as an effort to "contain" China. Indian political commentator Brahma Chellaney referred to the emerging Quadrilateral Security Dialogue between the United States, Japan, Australia and India as part of a new "Great Game" in Asia, and Indian diplomat M. K. Rasgotra has maintained that American efforts to shape security pacts in Asia will result not in an "Asian Century," but rather in an "American Century in Asia."


title: Quadrilateral Security Dialogue

text: Some, like US Lt. General Jeffrey B. Kohler, viewed US-India defence agreements as potentially lucrative for American defence industries and oversaw the subsequent sale of American military systems to India. Nevertheless, some Indian commentators opposed increased American military cooperation with India, citing the American presence in Iraq, hostility to Iran and "attempts at encircling China" as fundamentally destabilizing to Asian peace, and objecting to the presence of American warships with nuclear capabilities off the coast of southern India, or to American calls for the permanent hosting of American naval vessels in Goa or Kochi.


title: Quadrilateral Security Dialogue

text: The Trilateral Strategic Dialogue (TSD) was a series of trilateral meetings between the United States, Japan, and Australia. The TSD originally convened at senior officials level in 2002, then was upgraded to ministerial level in 2005. The United States expected regional allies to help facilitate evolving US global strategy to fight against terrorism and nuclear proliferation. In return, Japan and Australia expected benefits including continued US strategic involvement and the maintenance of strategic guarantees in the region.


title: Quadrilateral Security Dialogue

text: In 2021, some commentators wrote that an ad-hoc Tsunami Core Group in response to the 2004 Indian Ocean earthquake and tsunami may have been an early precursor to the Quad.


title: Quadrilateral Security Dialogue

text: The "nine-dash line" refers to the ill-defined demarcation line used by the People's Republic of China (China) and the Republic of China (Taiwan), for their claims of the major part of the South China Sea. The contested area in the South China Sea includes the Paracel Islands, the Spratly Islands, and various other areas including Pratas Island and the Vereker Banks, the Macclesfield Bank and the Scarborough Shoal. Despite having made the vague claim public in 1947, neither the PRC nor the ROC has () filed a formal and specifically defined claim to the area. An early map showing a U-shaped eleven-dash line was published in the then-Republic of China on 1 December 1947. Two of the dashes in the Gulf of Tonkin were later removed at the behest of Chinese Premier Zhou Enlai, reducing the total to nine. Chinese scholars asserted at the time that the version of the map with nine dashes represented the maximum extent of historical claims to the South China Sea. Subsequent editions added a tenth dash to the east of Taiwan island in 2013, extending it into the East China Sea.


title: Quadrilateral Security Dialogue

text: In early 2007, Prime Minister Abe proposed the Quadrilateral Security Dialogue, or "Quadrilateral Initiative", under which India would join a formal multilateral dialogue with Japan, the United States and Australia.


title: Quadrilateral Security Dialogue

text: The initiation of an American, Japanese, Australian and Indian defence arrangement, modelled on the concept of a Democratic Peace, was credited to Japanese Prime Minister Shinzo Abe. The Quadrilateral was supposed to establish an "Asian Arc of Democracy", envisioned to ultimately include countries in Central Asia, Mongolia, the Korean Peninsula, and other countries in Southeast Asia: "virtually all the countries on China’s periphery, except for China itself." This led some critics, such as former U.S. State Department official Morton Abramowitz, to call the project "an anti-Chinese move", while others have called it a "democratic challenge" to the projected Chinese century, mounted by Asian powers in coordination with the United States. While China has traditionally favoured the Shanghai Cooperation Organisation, the Quadrilateral was viewed as an "Asian NATO;" Daniel Twining of the German Marshall Fund of the United States has written that the arrangement "could lead to military conflict," or could instead "lay an enduring foundation for peace" if China becomes a democratic leader in Asia.


title: Quadrilateral Security Dialogue

text: China sent diplomatic protests to all four members of the Quadrilateral before any formal convention of its members. In May 2007 in Manila, Australian Prime Minister John Howard participated with other members in the inaugural meeting of the Quadrilateral at Cheney's urging, one month after joint naval exercises near Tokyo by India, Japan and the United States. In September 2007 further naval exercises were held in the Bay of Bengal, including Australia. These were followed in October by a further security agreement between Japan and India, ratified during a visit by Indian Prime Minister Manmohan Singh to Tokyo, to promote sea lane safety and defence collaboration; Japan had previously established such an agreement only with Australia.


title: Quadrilateral Security Dialogue

text: Though the Quadrilateral initiative of the Bush Administration improved relationships with New Delhi, it gave the impression of "encircling" China. The security agreement between Japan and India furthermore made China conspicuous as absent on the list of Japan's strategic partners in Asia. These moves appeared to "institutionally alienate" China, the Association of South-East Asian Nations (ASEAN), and promote a "Washington-centric" ring of alliances in Asia.


title: Quadrilateral Security Dialogue

text: The Japanese Prime Minister succeeding Abe, Taro Aso, downplayed the importance of China in the Japan-India pact signed following the creation of the Quadrilateral, stating, "There was mention of China – and we do not have any assumption of a third country as a target such as China." Indian Foreign Secretary Shiv Shankar Menon similarly argued that the defence agreement was long overdue because of Indian freight trade with Japan, and did not specifically target China. On the cusp of visits to China and meetings with Prime Minister Wen Jiabao and President Hu Jintao in January 2008, the Indian prime minister, Manmohan Singh, declared that "India is not part of any so-called contain China effort," after being asked about the Quadrilateral.