Darwin 's finches - wikipedia Darwin 's finches Darwin 's finches Large ground finch, Medium ground finch Small tree finch, Green warbler - finch Scientific classification Kingdom : Animalia Phylum : Chordata Class : Aves Order : Passeriformes Family : Thraupidae Genera Geospiza Camarhynchus / Platyspiza Certhidea Pinaroloxias Darwin 's finches ( also known as the Galápagos finches ) are a group of about fifteen species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or tribe Geospizini. They belong to the tanager family and are not closely related to the true finches. The closest known relative of the Galápagos finches is Tiaris obscurus. They were first collected by Charles Darwin on the Galápagos Islands during the second voyage of the Beagle. Apart from the Cocos finch, which is from Cocos Island, the others are found only on the Galápagos Islands. The term `` Darwin 's finches '' was first applied by Percy Lowe in 1936, and popularised in 1947 by David Lack in his book Darwin 's Finches. David Lack based his analysis on the large collection of museum specimens collected by the 1905 -- 06 Galápagos expedition of the California Academy of Sciences, to whom Lack dedicated his 1947 book. The birds vary in size from 10 to 20 cm and weigh between 8 and 38 grams. The smallest are the warbler - finches and the largest is the vegetarian finch. The most important differences between species are in the size and shape of their beaks, which are highly adapted to different food sources. The birds are all dull - coloured. A long - term study carried out for more than 40 years by the Princeton University researchers Peter and Rosemary Grant has documented micro-evolutionary changes in beak size affected by El Nino / La Nina cycles in the Pacific. Contents 1 Darwin 's theory 1.1 Text from The Voyage of the Beagle 1.2 Text from On the Origin of Species 2 Polymorphism in Darwin 's finches 3 Taxonomy 3.1 Family 3.2 Species 4 Molecular basis of beak evolution 5 See also 6 Notes 7 References 8 External links Darwin 's theory During the survey voyage of HMS Beagle, Darwin was unaware of the significance of the birds of the Galápagos. He had learned how to preserve bird specimens while at the University of Edinburgh and had been keen on shooting, but he had no expertise in ornithology and by this stage of the voyage concentrated mainly on geology. In Galápagos he mostly left bird shooting to his servant Syms Covington. Nonetheless, these birds were to play an important part in the inception of Darwin 's theory of evolution by natural selection. On the Galápagos Islands and afterward, Darwin thought in terms of `` centres of creation '' and rejected ideas concerning the transmutation of species. From Henslow 's teaching, he was interested in the geographical distribution of species, particularly links between species on oceanic islands and on nearby continents. On Chatham Island, he recorded that a mockingbird was similar to those he had seen in Chile, and after finding a different one on Charles Island he carefully noted where mockingbirds had been caught. In contrast, he paid little attention to the finches. When examining his specimens on the way to Tahiti, Darwin noted that all of the mockingbirds on Charles Island were of one species, those from Albemarle of another, and those from James and Chatham Islands of a third. As they sailed home about nine months later, this, together with other facts, including what he had heard about Galápagos tortoises, made him wonder about the stability of species. Following his return from the voyage, Darwin presented the finches to the Zoological Society of London on 4 January 1837, along with other mammal and bird specimens that he had collected. The bird specimens, including the finches, were given to John Gould, the famous English ornithologist, for identification. Gould set aside his paying work and at the next meeting, on 10 January, reported that the birds from the Galápagos Islands that Darwin had thought were blackbirds, `` gross - beaks '' and finches were actually `` a series of ground Finches which are so peculiar ( as to form ) an entirely new group, containing 12 species ''. This story made the newspapers. Darwin had been in Cambridge at that time. In early March, he met Gould again and for the first time got a full report on the findings, including the point that his Galápagos `` wren '' was another closely allied species of finch. The mockingbirds that Darwin had labelled by island were separate species rather than just varieties. Gould found more species than Darwin had expected, and concluded that 25 of the 26 land birds were new and distinct forms, found nowhere else in the world but closely allied to those found on the South American continent. Darwin now saw that, if the finch species were confined to individual islands, like the mockingbirds, this would help to account for the number of species on the islands, and he sought information from others on the expedition. Specimens had also been collected by Captain Robert FitzRoy, FitzRoy 's steward Harry Fuller and Darwin 's servant Covington, who had labelled them by island. From these, Darwin tried to reconstruct the locations from where he had collected his own specimens. The conclusions supported his idea of the transmutation of species. Seen here is adapted radiation of finch A. ( Geospiza magnirostris ) into three other species of finches found on the Galapagos Islands. Due to the absence of other species of birds, the finches adapted to new niches. The finches beaks and bodies changed allowing them to eat certain types of foods such as nuts, fruits, and insects. Geospiza magnirostris Geospiza parvula Certhidea olivacea Geospiza fortis Text from the voyage of the Beagle At the time that he rewrote his diary for publication as Journal and Remarks ( later The Voyage of the Beagle ), he described Gould 's findings on the number of birds, noting that `` Although the species are thus peculiar to the archipelago, yet nearly all in their general structure, habits, colour of feathers, and even tone of voice, are strictly American ''. In the first edition of The Voyage of the Beagle, Darwin said that `` It is very remarkable that a nearly perfect gradation of structure in this one group can be traced in the form of the beak, from one exceeding in dimensions that of the largest gros - beak, to another differing but little from that of a warbler ''. By the time the first edition was published, the development of Darwin 's theory of natural selection was in progress. For the 1845 second edition of The Voyage ( now titled Journal of Researches ), Darwin added more detail about the beaks of the birds, and two closing sentences which reflected his changed ideas : `` Seeing this gradation and diversity of structure in one small, intimately related group of birds, one might really fancy that from an original paucity of birds in this archipelago, one species had been taken and modified for different ends. '' The remaining land - birds form a most singular group of finches, related to each other in the structure of their beaks, short tails, form of body and plumage : there are thirteen species, which Mr. Gould has divided into four subgroups. All these species are peculiar to this archipelago ; and so is the whole group, with the exception of one species of the sub-group Cactornis, lately brought from Bow Island, in the Low Archipelago. Of Cactornis, the two species may be often seen climbing about the flowers of the great cactus - trees ; but all the other species of this group of finches, mingled together in flocks, feed on the dry and sterile ground of the lower districts. The males of all, or certainly of the greater number, are jet black ; and the females ( with perhaps one or two exceptions ) are brown. The most curious fact is the perfect gradation in the size of the beaks in the different species of Geospiza, from one as large as that of a hawfinch to that of a chaffinch, and ( if Mr. Gould is right in including his sub-group, Certhidea, in the main group ) even to that of a warbler. The largest beak in the genus Geospiza is shown in Fig. 1, and the smallest in Fig. 3 ; but instead of there being only one intermediate species, with a beak of the size shown in Fig. 2, there are no less than six species with insensibly graduated beaks. The beak of the sub-group Certhidea, is shown in Fig. 4. The beak of Cactornis is somewhat like that of a starling, and that of the fourth subgroup, Camarhynchus, is slightly parrot - shaped. Seeing this gradation and diversity of structure in one small, intimately related group of birds, one might really fancy that from an original paucity of birds in this archipelago, one species had been taken and modified for different ends. In a like manner it might be fancied that a bird originally a buzzard, had been induced here to undertake the office of the carrion - feeding Polybori of the American continent. Text from on the Origin of species Darwin discussed the divergence of species of birds in the Galápagos more explicitly in his chapter on geographical distribution in On the Origin of Species : The most striking and important fact for us in regard to the inhabitants of islands, is their affinity to those of the nearest mainland, without being actually the same species. ( In ) the Galapagos Archipelago... almost every product of the land and water bears the unmistakable stamp of the American continent. There are twenty - six land birds, and twenty - five of these are ranked by Mr. Gould as distinct species, supposed to have been created here ; yet the close affinity of most of these birds to American species in every character, in their habits, gestures, and tones of voice, was manifest... The naturalist, looking at the inhabitants of these volcanic islands in the Pacific, distant several hundred miles from the continent, yet feels that he is standing on American land. Why should this be so ? why should the species which are supposed to have been created in the Galapagos Archipelago, and nowhere else, bear so plain a stamp of affinity to those created in America ? There is nothing in the conditions of life, in the geological nature of the islands, in their height or climate, or in the proportions in which the several classes are associated together, which resembles closely the conditions of the South American coast : in fact there is a considerable dissimilarity in all these respects. On the other hand, there is a considerable degree of resemblance in the volcanic nature of the soil, in climate, height, and size of the islands, between the Galapagos and Cape de Verde Archipelagos : but what an entire and absolute difference in their inhabitants ! The inhabitants of the Cape de Verde Islands are related to those of Africa, like those of the Galapagos to America. I believe this grand fact can receive no sort of explanation on the ordinary view of independent creation ; whereas on the view here maintained, it is obvious that the Galapagos Islands would be likely to receive colonists, whether by occasional means of transport or by formerly continuous land, from America ; and the Cape de Verde Islands from Africa ; and that such colonists would be liable to modification ; -- the principle of inheritance still betraying their original birthplace. Polymorphism in Darwin 's finches Whereas Darwin spent just five weeks in the Galápagos, and David Lack spent three months, Peter and Rosemary Grant and their colleagues have made research trips to the Galápagos for about thirty years, particularly studying Darwin 's finches. Here, we look briefly at the case of the large cactus finch Geospiza conirostris on Isla Genovesa ( formerly Tower Island ), which is formed from a shield volcano, and is home to a variety of birds. These birds, like all well - studied groups, show various kinds of morphism. Males are dimorphic in song type : songs A and B are quite distinct. Also, males with song A have shorter bills than B males. This is also a clear difference. With these beaks, males are able to feed differently on their favourite cactus, the prickly pear Opuntia. Those with long beaks are able to punch holes in the cactus fruit and eat the fleshy aril pulp, which surrounds the seeds, whereas those with shorter beaks tear apart the cactus base and eat the pulp and any insect larvae and pupae ( both groups eat flowers and buds ). This dimorphism clearly maximises their feeding opportunities during the non-breeding season when food is scarce. If the population is panmixic, then Geospiza conirostris exhibits a balanced genetic polymorphism and not, as originally supposed, a case of nascent sympatric speciation. The selection maintaining the polymorphism maximises the species ' niche by expanding its feeding opportunity. The genetics of this situation can not be clarified in the absence of a detailed breeding program, but two loci with linkage disequilibrium is a possibility. Another interesting dimorphism is for the bills of young finches, which are either ' pink ' or ' yellow '. All species of Darwin 's finches exhibit this morphism, which lasts for two months. No interpretation of this phenomenon is known. Taxonomy Family For some decades, taxonomists have placed these birds in the family Emberizidae along with the New World sparrows and Old World buntings ( Sulloway 1982 ). However, the Sibley -- Ahlquist taxonomy puts Darwin 's finches with the tanagers ( Monroe and Sibley 1993 ), and at least one recent work follows that example ( Burns and Skutch 2003 ). The American Ornithologists ' Union, in its North American check - list, places the Cocos finch in the Emberizidae but with an asterisk indicating that the placement is probably wrong ( AOU 1998 -- 2006 ) ; in its tentative South American check - list, the Galápagos species are incertae sedis, of uncertain place ( Remsen et al. 2007 ). Species Genus Geospiza Large cactus finch ( Geospiza conirostris ) Sharp - beaked ground finch ( Geospiza difficilis ) Vampire finch ( Geospiza difficilis septentrionalis ) Medium ground finch ( Geospiza fortis ) Small ground finch ( Geospiza fuliginosa ) Large ground finch ( Geospiza magnirostris ) Common cactus finch ( Geospiza scandens ) Genus Camarhynchus Large tree finch ( Camarhynchus psittacula ) Medium tree finch ( Camarhynchus pauper ) Small tree finch ( Camarhynchus parvulus ) Woodpecker finch ( Camarhynchus pallidus ) -- sometimes separated in Cactospiza Mangrove finch ( Camarhynchus heliobates ) Genus Certhidea Green warbler - finch ( Certhidea olivacea ) Grey warbler - finch ( Certhidea fusca ) Genus Pinaroloxias Cocos finch ( Pinaroloxias inornata ) Genus Platyspiza Vegetarian finch ( Platyspiza crassirostris ) Molecular basis of beak evolution Developmental research in 2004 found that bone morphogenetic protein 4 ( BMP4 ), and its differential expression during development, resulted in variation of beak size and shape among finches. BMP4 acts in the developing embryo to lay down skeletal features, including the beak. The same group showed that the development of the different beak shapes in Darwin 's finches are also influenced by slightly different timing and spatial expressions of a gene called calmodulin ( CaM ). Calmodulin acts in a similar way to BMP4, affecting some of the features of beak growth. The authors suggest that changes in the temporal and spatial expression of these two factors are possible developmental controls of beak morphology. In a recent study genome sequencing revealed a 240 kilobase haplotype encompassing the ALX1 gene that encodes a transcription factor affecting craniofacial development is strongly associated with beak shape diversity. Moreover, these changes in the beak size have also altered vocalization in Darwin 's finches. See also Species flock Adaptive radiation Notes Jump up ^ http://www.nature.com/news/darwin-s-iconic-finches-join-genome-club-1.16896 Jump up ^ http://www.uu.se/en/media/news/article/?id=4166. Jump up ^ http://rstb.royalsocietypublishing.org/content/365/1543/1093 Jump up ^ Grant & Grant 2008, p. 3 ^ Jump up to : PODOS, JEFFREY ; NOWICKI, STEPHEN. `` Beaks, Adaptation, and Vocal Evolution in Darwin 's Finches ''. BioScience. 54 ( 6 ). doi : 10.1641 / 0006 - 3568 ( 2004 ) 054 ( 0501 : baavei ) 2.0.co ; 2. Jump up ^ Sato, Akie ( 2001 ). `` On the Origin of Darwin 's Finches '' ( PDF ). Oxford. Jump up ^ Lack, David. 1947. Darwin 's Finches. Cambridge University Press ( reissued in 1961 by Harper, New York, with a new preface by Lack ; reissued in 1983 by Cambridge University Press with an introduction and notes by Laurene M. Ratcliffe and Peter T. Boag ). ISBN 0 - 521 - 25243 - 1 Jump up ^ Steinheimer 2004, p. 300 Jump up ^ Lewitt, Dan. `` Galapagos Finch Evolution -- Dan Lewitt -- HHMI ( 2013 ) ''. Retrieved 6 June 2017. ^ Jump up to : Grant, K. Thalia and Estes, Gregory B. 2009 `` Darwin in Galapagos : Footsteps to a New World '' Princeton University Press, Princeton. Jump up ^ Steinheimer 2004, pp. 301 -- 303 Jump up ^ Keynes 2000, p. xix. Eldredge 2006 Jump up ^ Chancellor, Gordon ; Keynes, Randal ( October 2006 ), Darwin 's field notes on the Galapagos : ' A little world within itself ', Darwin Online Jump up ^ Eldredge 2006 Jump up ^ Desmond & Moore 1991, pp. 208 -- 209 ^ Jump up to : Sulloway 1982, pp. 57 -- 58 Jump up ^ Desmond & Moore 1991, p. 248 Jump up ^ Sulloway 2006 Jump up ^ Darwin 1839, pp. 461 -- 462 Jump up ^ Darwin 1839, p. 462. Jump up ^ Darwin 1845, pp. 379 -- 380 Jump up ^ Darwin 1887 Jump up ^ Darwin 1845, p. 380. Jump up ^ Darwin 1859, pp. 397 -- 398. Jump up ^ Huxley J. 1955. Morphism in birds. 11th Int Ornith Cong ( Basel 1954, p309 - 328 ) touches on this theme. Jump up ^ B. Rosemary Grant and Peter R. Grant 1989. Evolutionary dynamics of a natural population : the large cactus finch of the Galápagos. Chicago, p241 first para. Jump up ^ Grant, Peter R. 1999. Ecology and evolution of Darwin 's finches. Princeton NJ, p428 in Afterword. Jump up ^ Maynard Smith J. 1998. Evolutionary genetics. 2nd ed, Chapter 5, Oxford. Jump up ^ Grant, Peter R. 1999. Ecology and evolution of Darwin 's finches. Princeton NJ. ( see plate 7 ) Jump up ^ Abzhanov, Arhat ; Protas, Meredith ; Grant, B. Rosemary ; Grant, Peter R. ; Tabin, Clifford J. ( September 3, 2004 ), `` Bmp4 and Morphological Variation of Beaks in Darwin 's Finches '', Science, 305 ( 5689 ) : 1462 -- 1465, Bibcode : 2004Sci... 305.1462 A, ISSN 0036 - 8075, OCLC 1644869, PMID 15353802, doi : 10.1126 / science. 1098095 Jump up ^ Abzhanov, Arhat ; Kuo, Winston P. ; Hartmann, Christine ; Grant, B. Rosemary ; Grant, Peter R. ; Tabin, Clifford J. ( August 3, 2006 ), `` The calmodulin pathway and evolution of elongated beak morphology in Darwin 's finches '', Nature, 442 ( 7102 ) : 563 -- 567, Bibcode : 2006Natur. 442... 563A, ISSN 0028 - 0836, OCLC 1586310, PMID 16885984, doi : 10.1038 / nature04843 Jump up ^ Andersson, Leif ; Lamichhaney, Sangeet ; Berglund, Jonas ; Almén, Markus Sällman ; Maqbool, Khurram ; Grabherr, Manfred ; Martinez - Barrio, Alvaro ; Promerová, Marta ; Rubin, Carl - Johan ; Wang, Chao ; Zamani, Neda ; Grant, B. Rosemary ; Grant, Peter R. ; Webster, Matthew T. ( February 11, 2015 ), `` Evolution of Darwin 's finches and their beaks revealed by genome sequencing '', Nature, 518 ( 7102 ) : 563 -- 7, Bibcode : 2006Natur. 442... 563A, ISSN 0028 - 0836, OCLC 1586310, PMID 25686609, doi : 10.1038 / nature14181 Jump up ^ 200 years after Darwin, this is how the iconic Galapagos finches are still evolving 2016 April 22 Jump up ^ Podos & Nowicki, Jeffrey & Stephen ( 2004 ). `` Beaks, Adaptation, and Vocal Evolution in Darwin 's Finches ''. Bioscience. 54 : 501 -- 510 -- via Oxford academy. References Darwin, Charles ( 1839 ), Narrative of the surveying voyages of His Majesty 's Ships Adventure and Beagle between the years 1826 and 1836, describing their examination of the southern shores of South America, and the Beagle 's circumnavigation of the globe. Journal and remarks. 1832 -- 1836, III, London : Henry Colburn Darwin, Charles ( 1845 ), Journal of researches into the natural history and geology of the countries visited during the voyage of H.M.S. Beagle round the world, under the Command of Capt. Fitz Roy, R.N ( 2nd. ed. ), London : John Murray Darwin, Charles ( 1859 ), On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life ( 1st ed. ), London : John Murray Darwin, Francis ( 1887 ), `` Chapter 1, The Foundations of the ' Origin of Species ' '', in Darwin, Francis, The life and letters of Charles Darwin, including an autobiographical chapter, 2, London : John Murray Desmond, Adrian ; Moore, James ( 1991 ), Darwin, London : Michael Joseph, Penguin Group, ISBN 0 - 7181 - 3430 - 3, OCLC 185764721 Eldredge, Niles ( 2006 ), `` Confessions of a Darwinist '', The Virginia Quarterly Review ( Spring 2006 ), pp. 32 -- 53, retrieved 2008 - 11 - 04 Grant, Peter R. ; Grant, B. Rosemary ( 2008 ), How and Why Species Multiply : The Radiation of Darwin 's Finches, Princeton University Press, ISBN 978 - 0 - 691 - 13360 - 7 Lack, David ( 1940 ), `` Evolution of the Galapagos Finches '', Nature ( published 7 September 1940 ), 146 ( 3697 ), pp. 324 -- 327, Bibcode : 1940Natur. 146... 324L, doi : 10.1038 / 146324a0, retrieved 2013 - 12 - 06 Check - list of North American Birds, American Ornithologists ' Union, 1998 -- 2006, archived from the original on April 4, 2007, retrieved 2007 - 04 - 09 Burns, Kevin J. ; Skutch, Alexander F. ( 2003 ), `` Tanagers and Tanager - Finches '', in Christopher Perrins, ed., The Firefly Encyclopedia of Birds, Firefly Books, pp. 629 -- 631, ISBN 1 - 55297 - 777 - 3, retrieved 2007 - 04 - 09 CS1 maint : Extra text : editors list ( link ) It is not clear whether this placement was made by Burns and Skutch or by Perrins. Keynes, Richard ( 2000 ), Charles Darwin 's zoology notes & specimen lists from H.M.S. Beagle, Cambridge University Press, retrieved 2008 - 12 - 08 Sibley, Charles G. ( 1993 ), A World Checklist of Birds, New Haven, Conn. : Yale University Press, ISBN 0 - 300 - 07083 - 7, retrieved 2013 - 12 - 06 Monroe and Sibley consider the tanagers to be a tribe ( Thraupini ) of a big family Fringillidae rather than a family of their own ( Thraupidae ). Zimmer, J. ( 2007 - 04 - 05 ), A classification of the bird species of South America, American Ornithologists ' Union, archived from the original on 2009 - 03 - 02, retrieved 2007 - 04 - 09 Steinheimer, F.D. ( 2004 ), `` Charles Darwin 's bird collection and ornithological knowledge during the voyage of H.M.S. Beagle, 1831 -- 1836 '', Journal of Ornithology, 145 ( 4 ), pp. 300 -- 320, doi : 10.1007 / s10336 - 004 - 0043 - 8, retrieved 2008 - 12 - 08 Sulloway, Frank J. ( 1982 ), `` The Beagle collections of Darwin 's finches ( Geospizinae ) '', Bulletin of the British Museum ( Natural History ), Historical Series, 43 ( 2 ), pp. 49 -- 94, retrieved 2008 - 12 - 08 Sulloway, Frank J. ( Spring 1982 ), `` Darwin and His Finches : The Evolution of a Legend '' ( PDF ), Journal of the History of Biology, 15 ( 1 ), pp. 1 -- 53, doi : 10.1007 / BF00132004, retrieved 2008 - 12 - 09 Sulloway, Frank J. ( 2006 ), `` Why Darwin Rejected Intelligent Design '' ( PDF ), in Brockman, John, Intelligent Thought : Science versus the Intelligent Design Movement, New York : Vintage, pp. 107 -- 126 External links Wikimedia Commons has media related to Darwin 's finches. Grant, K.T. and Estes, G.B. 2009. `` Darwin in Galapagos : Footsteps to a New World '' Princeton University Press, Princeton. ( 1 ) Sulloway, F.J. ( 1982 ) : `` Darwin and his finches : the evolution of a legend ''. J. Hist. Biol. 15 : p. 1 -- 53 Different bills and song melodies Genetics and the Origin of Birds Species, Grant and Grant in PNAS Sato et al. Phylogeny of Darwin 's finches as revealed by mtDNA sequences in PNAS Sato A, Tichy H, O'hUigin C, Grant PR, Grant BR, Klein J ( March 2001 ), `` On the origin of Darwin 's finches '', Mol. Biol. Evol., 18 ( 3 ) : 299 -- 311, PMID 11230531 Galápagos Online. Darwin 's Finches. Galapagos Online. List of birds of the Galapagos Islands. Darwin 's Finches Evolve Before Scientists ' Eyes : new developments reported 13 July 2006 Fink F.A.Q. Darwin 's finches inspired the naming of the Fink project, a collaborative initiative for porting open source software to the Darwin platform to enable its use and evolution in the Apple Mac OS X environment. `` Fink '' is the German name for `` finch. '' Aug 2006 Nature Article that shows how modulation of a certain gene during development can account for the differences seen in beak shape. Speciation Kimball 's Biology Pages Birds ( class : Aves ) Anatomy Bird anatomy Flight Eggs Feathers Plumage Beak Vision Dactyly Preen gland Behaviour Singing Intelligence Migration Sexual selection Lek mating Seabird breeding Incubation Brood parasites Nesting Hybrids Evolution Origin of birds Origin of flight Evolution of birds Darwin 's finches Seabirds Fossil birds Archaeopteryx Omnivoropterygiformes Confuciusornithiformes Enantiornithes Chaoyangiiformes Patagopterygiformes Ambiortiformes Songlingornithiformes Apsaraviformes Gansuiformes Ichthyornithiformes Hesperornithes Lithornithiformes Dinornithiformes Aepyornithiformes Gastornithiformes Human interaction Ringing Ornithology Bird collections Birdwatching Bird feeding Conservation Aviculture Waterfowl hunting Cockfighting Pigeon racing Falconry Pheasantry Egg collecting Ornithomancy Lists Families and orders Genera Glossary of bird terms List by population Lists by region Recently extinct birds Late Quaternary prehistoric birds Sibley -- Ahlquist taxonomy Historical & Fictional Neornithes Palaeognathae Struthioniformes ( ostriches ) Rheiformes ( rheas ) Tinamiformes ( tinamous ) Apterygiformes ( kiwis ) Casuariiformes ( emus and cassowaries ) Neognathae Galloanserae ( fowls ) Anseriformes ( waterfowls ) Anatidae Anatinae Anserinae swans true geese Aythyinae Dendrocygninae Merginae Oxyurinae Plectropterinae Stictonettinae Tadorninae Thalassorninae Anhimidae Anhima Chauna Anseranatidae Anatalavis Anseranas Galliformes ( landfowls - gamebirds ) Cracidae Cracinae Oreophasinae Penelopinae Megapodidae Aepypodius Alectura Eulipoa Leipoa Macrocephalon Megapodius Talegalla Numididae Acryllium Agelastes Guttera Numida Odontophoridae Callipepla Colinus Cyrtonyx Dactylortyx Dendrortyx Odontophorus Oreortyx Philortyx Rhynchortyx Phasianidae Meleagridinae Perdicinae Phasianinae ( pheasants and relatives ) Tetraoninae Neoaves Columbea Columbimorphae Columbiformes ( doves and pigeons ) Mesitornithiformes ( mesites ) Pteroclidiformes ( sandgrouses ) Mirandornithes Phoenicopteriformes ( flamingos ) Podicipediformes ( grebes ) Passerea Otidimorphae Cuculiformes ( cuckoos ) Musophagiformes ( turacos ) Otidiformes ( bustards ) Strisores Caprimulgiformes ( nightjars and relatives ) Steatornithiformes Podargiformes Apodiformes ( swifts and hummingbirds ) Opisthocomiformes Opisthocomiformes ( hoatzin ) Cursorimorphae Charadriiformes ( gulls and relatives ) Gruiformes ( cranes and relatives ) Phaethontimorphae Phaethontiformes ( tropicbirds ) Eurypygiformes ( kagu and sunbittern ) Aequornithes Gaviiformes ( loons or divers ) Sphenisciformes ( penguins ) Procellariiformes ( albatrosses and petrels ) Ciconiiformes ( storks ) Suliformes ( cormorants and relatives ) Pelecaniformes ( pelicans and relatives ) Australaves Cariamiformes ( seriemas and relatives ) Falconiformes ( falcons and relatives ) Psittaciformes ( parrots ) Passeriformes ( perching birds ) Afroaves Cathartiformes ( New World vultures and condors ) Accipitriformes ( eagles and hawks ) Strigiformes ( owls ) Coliiformes ( mousebirds ) Trogoniformes ( trogons and quetzals ) Leptosomatiformes ( cuckoo roller ) Bucerotiformes ( hornbills and hoopoes ) Coraciiformes ( kingfishers and rollers ) Piciformes ( woodpeckers and relatives ) Portal Outline Retrieved from `` https://en.wikipedia.org/w/index.php?title=Darwin%27s_finches&oldid=807869375 '' Categories : Charles Darwin Endemic birds of the Galápagos Islands Evolution of birds Finches Geospizini History of evolutionary biology Polymorphism ( biology ) Hidden categories : Articles with ' species ' microformats CS1 maint : Extra text : editors list Talk Contents About Wikipedia Български Català Čeština Deutsch Español فارسی Français Gaeilge Galego 한국어 Italiano עברית Magyar Nederlands 日本 語 Norsk Norsk nynorsk Polski Português Русский Simple English Suomi Svenska Türkçe Українська Tiếng Việt 中文 Edit links This page was last edited on 30 October 2017, at 15 : 34. Text is available under the Creative Commons Attribution - ShareAlike License ; additional terms may apply. By using this site, you agree to the Terms of Use and Privacy Policy. Wikipedia ® is a registered trademark of the Wikimedia Foundation, Inc., a non-profit organization. About Wikipedia

Question: how many types of finch species are there
Answer: about fifteen