Wasp - wikipedia Wasp This is the latest accepted revision, reviewed on 24 May 2018. For other uses, see Wasp ( disambiguation ). Wasp Temporal range : Jurassic -- Present PreЄ Є Pg A social wasp, Vespula germanica Scientific classification Kingdom : Animalia Phylum : Arthropoda Class : Insecta Order : Hymenoptera Suborder : Apocrita ( except clade Anthophila and family Formicidae ) A wasp is any insect of the order Hymenoptera and suborder Apocrita that is neither a bee nor an ant. The Apocrita have a common evolutionary ancestor and form a clade ; wasps as a group do not form a clade, but are paraphyletic with respect to bees and ants. The most commonly known wasps, such as yellowjackets and hornets, are in the family Vespidae and are eusocial, living together in a nest with an egg - laying queen and non-reproducing workers. Eusociality is favoured by the unusual haplodiploid system of sex determination in Hymenoptera, as it makes sisters exceptionally closely related to each other. However, the majority of wasp species are solitary, with each adult female living and breeding independently. Wasps play many ecological roles. Some are predators or pollinators, whether to feed themselves or to provision their nests. Many, notably the cuckoo wasps, are kleptoparasites, laying eggs in the nests of other wasps. Many of the solitary wasps are parasitoidal, meaning they lay eggs on or in other insects ( any life stage from egg to adult ) and often provision their own nests with such hosts. Unlike true parasites, the wasp larvae eventually kill their hosts. Solitary wasps parasitize almost every pest insect, making wasps valuable in horticulture for biological pest control of species such as whitefly in tomatoes and other crops. Wasps first appeared in the fossil record in the Jurassic, and diversified into many surviving superfamilies by the Cretaceous. They are a successful and diverse group of insects with tens of thousands of described species ; wasps have spread to all parts of the world except for the polar regions. The largest social wasp is the Asian giant hornet, at up to 5 centimetres ( 2.0 in ) in length ; among the largest solitary wasps is a group of species known as tarantula hawks, along with the giant scoliid of Indonesia ( Megascolia procer ). The smallest wasps are solitary chalcid wasps in the family Mymaridae, including the world 's smallest known insect, with a body length of only 0.139 mm ( 0.0055 in ), and the smallest known flying insect, only 0.15 mm ( 0.0059 in ) long. Wasps have appeared in literature from Classical times, as the eponymous chorus of old men in Aristophanes ' 422 BC comedy Σφῆκες ( Sphēkes ), The Wasps, and in science fiction from H.G. Wells 's 1904 novel The Food of the Gods and How It Came to Earth, featuring giant wasps with three - inch - long stings. The name `` Wasp '' has been used for many warships and other military equipment. Contents 1 Taxonomy and phylogeny 1.1 Paraphyletic grouping 1.2 Fossils 1.3 Diversity 2 Social wasps 3 Solitary wasps 4 Biology 4.1 Anatomy 4.2 Diet 4.3 Sex determination 4.4 Inbreeding avoidance 5 Ecology 5.1 As pollinators 5.2 As parasitoids 5.3 As parasites 5.4 As predators 5.5 As models for mimics 5.6 As prey 6 Relationship with humans 6.1 As pests 6.2 In horticulture 6.3 In sport 6.4 In fashion 6.5 In literature 6.6 In military names 7 See also 8 Notes 9 References 10 Sources 11 External links Taxonomy and phylogeny Wasps are paraphyletic, consisting of the clade Apocrita without ants and bees, which are not usually considered to be wasps. The Hymenoptera also contain the somewhat wasplike Symphyta, the sawflies. The familiar common wasps and yellowjackets belong to one family, the Vespidae. Palaeovespa florissantia, a fossil wasp ( Vespinae ) from the Eocene rocks of the Florissant fossil beds of Colorado, c. 34 mya Paraphyletic grouping The wasps are a cosmopolitan paraphyletic grouping of hundreds of thousands of species, consisting of the narrow - waisted Apocrita without the ants and bees. The Hymenoptera also contain the somewhat wasplike but unwaisted Symphyta, the sawflies. The term wasp is sometimes used more narrowly for the Vespidae, which includes the common wasp or yellow jacket genera Vespula and Dolichovespula and the hornets, Vespa ; or simply for the common wasp and close lookalikes. Fossils Male Electrostephanus petiolatus fossil from the Middle Eocene, preserved in Baltic amber Hymenoptera in the form of Symphyta ( Xyelidae ) first appeared in the fossil record in the Lower Triassic. Apocrita, wasps in the broad sense, appeared in the Jurassic, and had diversified into many of the extant superfamilies by the Cretaceous ; they appear to have evolved from the Symphyta. Fig wasps with modern anatomical features first appeared in the Lower Cretaceous of the Crato Formation in Brazil, some 65 million years before the first fig trees. The Vespidae include the extinct genus Palaeovespa, seven species of which are known from the Eocene rocks of the Florissant fossil beds of Colorado and from fossilised Baltic amber in Europe. Also found in Baltic amber are crown wasps of the genus Electrostephanus. Diversity Megascolia procer, a giant solitary species from Java in the Scoliidae. This specimen 's length is 7.7 cm and its wingspan is 11.5 cm. Wasps are a diverse group, estimated at over a hundred thousand described species around the world, and a great many more as yet undescribed. For example, there are over 800 species of fig trees, mostly in the tropics, and almost all of these has its own specific fig wasp ( Chalcidoidea ) to effect pollination. Megarhyssa macrurus, a parasitoid. The body of a female is c. 2 inches ( 51 mm ) long, with an ovipositor c. 4 inches ( 100 mm ) long Many wasp species are parasitoids ; the females deposit eggs on or in a host arthropod on which the larvae then feed. Some larvae start off as parasitoids, but convert at a later stage to consuming the plant tissues that their host is feeding on. In other species, the eggs are laid directly into plant tissues and form galls, which protect the developing larvae from predators but not necessarily from other parasitic wasps. In some species, the larvae are predatory themselves ; the wasp eggs are deposited in clusters of eggs laid by other insects, and these are then consumed by the developing wasp larvae. Tarantula hawk wasp dragging an orange - kneed tarantula to her burrow ; this species has the most painful sting of any wasp. The largest social wasp is the Asian giant hornet, at up to 5 centimetres ( 2.0 in ) in length. The various tarantula hawk wasps are of a similar size and can overpower a spider many times its own weight, and move it to its burrow, with a sting that is excruciatingly painful to humans. The solitary giant scoliid, Megascolia procer, with a wingspan of 11.5 cm, has subspecies in Sumatra and Java ; it is a parasitoid of the Atlas beetle Chalcosoma atlas. The female giant ichneumon wasp Megarhyssa macrurus is 12.5 centimetres ( 5 in ) long including its very long but slender ovipositor which is used for boring into wood and inserting eggs. The smallest wasps are solitary chalcid wasps in the family Mymaridae, including the world 's smallest known insect, Dicopomorpha echmepterygis ( 139 micrometres long ) and Kikiki huna with a body length of only 158 micrometres, the smallest known flying insect. There are estimated to be 100,000 species of ichneumonoid wasps in the families Braconidae and Ichneumonidae. These are almost exclusively parasitoids, mostly utilising other insects as hosts. Another family, the Pompilidae, is a specialist parasitoid of spiders. Some wasps are even parasitoids of parasitoids ; the eggs of Euceros are laid beside lepidopteran larvae and the wasp larvae feed temporarily on their haemolymph, but if a parasitoid emerges from the host, the hyperparasites continue their life cycle inside the parasitoid. Parasitoids maintain their extreme diversity through narrow specialism. In Peru, 18 wasp species were found living on 14 fly species in only two species of Gurania climbing squash. Social wasps Social wasps constructing a paper nest Of the dozens of extant wasp families, only the family Vespidae contains social species, primarily in the subfamilies Vespinae and Polistinae. With their powerful stings and conspicuous warning coloration, often in black and yellow, social wasps are frequent models for Batesian mimicry by non-stinging insects, and are themselves involved in mutually beneficial Müllerian mimicry of other distasteful insects including bees and other wasps. All species of social wasps construct their nests using some form of plant fiber ( mostly wood pulp ) as the primary material, though this can be supplemented with mud, plant secretions ( e.g., resin ), and secretions from the wasps themselves ; multiple fibrous brood cells are constructed, arranged in a honeycombed pattern, and often surrounded by a larger protective envelope. Wood fibers are gathered from weathered wood, softened by chewing and mixing with saliva. The placement of nests varies from group to group ; yellow jackets such as Dolichovespula media and D. sylvestris prefer to nest in trees and shrubs ; Protopolybia exigua attaches its nests on the underside of leaves and branches ; Polistes erythrocephalus chooses sites close to a water source. Play media Social wasps on their nest Other wasps, like Agelaia multipicta and Vespula germanica, like to nest in cavities that include holes in the ground, spaces under homes, wall cavities or in lofts. While most species of wasps have nests with multiple combs, some species, such as Apoica flavissima, only have one comb. The length of the reproductive cycle depends on latitude ; Polistes erythrocephalus, for example, has a much longer ( up to 3 months longer ) cycle in temperate regions. Solitary wasps Potter wasp building mud nest, France. The latest ring of mud is still wet. The vast majority of wasp species are solitary insects. Having mated, the adult female forages alone and if it builds a nest, does so for the benefit of its own offspring. Some solitary wasps nest in small groups alongside others of their species, but each is involved in caring for its own offspring ( except for such actions as stealing other wasps ' prey or laying in other wasp 's nests ). There are some species of solitary wasp that build communal nests, each insect having its own cell and providing food for its own offspring, but these wasps do not adopt the division of labour and the complex behavioural patterns adopted by eusocial species. Adult solitary wasps spend most of their time in preparing their nests and foraging for food for their young, mostly insects or spiders. Their nesting habits are more diverse than those of social wasps. Many species dig burrows in the ground. Mud daubers and pollen wasps construct mud cells in sheltered places. Potter wasps similarly build vase - like nests from mud, often with multiple cells, attached to the twigs of trees or against walls. Predatory wasp species normally subdue their prey by stinging it, and then either lay their eggs on it, leaving it in place, or carry it back to their nest where an egg may be laid on the prey item and the nest sealed, or several smaller prey items may be deposited to feed a single developing larva. Apart from providing food for their offspring, no further maternal care is given. Members of the family Chrysididae, the cuckoo wasps, are kleptoparasites and lay their eggs in the nests of unrelated host species. Biology Anatomy European hornet, Vespa crabro Like all insects, wasps have a hard exoskeleton which protects their three main body parts, the head, the mesosoma ( including the thorax and the first segment of the abdomen ) and the metasoma. There is a narrow waist, the petiole, joining the first and second segments of the abdomen. The two pairs of membranous wings are held together by small hooks and the forewings are larger than the hind ones ; in some species, the females have no wings. In females there is usually a rigid ovipositor which may be modified for injecting venom, piercing or sawing. It either extends freely or can be retracted, and may be developed into a stinger for both defence and for paralysing prey. In addition to their large compound eyes, wasps have several simple eyes known as ocelli, which are typically arranged in a triangle just forward of the vertex of the head. Wasps possess mandibles adapted for biting and cutting, like those of many other insects, such as grasshoppers, but their other mouthparts are formed into a suctorial proboscis, which enables them to drink nectar. The larvae of wasps resemble maggots, and are adapted for life in a protected environment ; this may be the body of a host organism or a cell in a nest, where the larva either eats the provisions left for it or, in social species, is fed by the adults. Such larvae have soft bodies with no limbs, and have a blind gut ( presumably so that they do not foul their cell ). Sand wasp Bembix oculata ( Crabronidae ) feeding on a fly after paralysing it with its sting Diet Black wasp Sphex pensylvanicus ( Sphecidae ) feeding on nectar Adult solitary wasps mainly feed on nectar, but the majority of their time is taken up by foraging for food for their carnivorous young, mostly insects or spiders. Apart from providing food for their larval offspring, no maternal care is given. Some wasp species provide food for the young repeatedly during their development ( progressive provisioning ). Others, such as potter wasps ( Eumeninae ) and sand wasps ( Ammophila, Sphecidae ), repeatedly build nests which they stock with a supply of immobilised prey such as one large caterpillar, laying a single egg in or on its body, and then sealing up the entrance ( mass provisioning ). Spider wasp ( Pompilidae ) dragging a jumping spider ( Salticidae ) to provision a nest Predatory and parasitoidal wasps subdue their prey by stinging it. They hunt a wide variety of prey, mainly other insects ( including other Hymenoptera ), both larvae and adults. The Pompilidae specialize in catching spiders to provision their nests. Some social wasps are omnivorous, feeding on fallen fruit, nectar, and carrion such as dead insects. Adult male wasps sometimes visit flowers to obtain nectar. Some wasps, such as Polistes fuscatus, commonly return to locations where they previously found prey to forage. In many social species, the larvae exude copious amounts of salivary secretions that are avidly consumed by the adults. These include both sugars and amino acids, and may provide essential protein - building nutrients that are otherwise unavailable to the adults ( who can not digest proteins ). Sex determination In wasps, as in other Hymenoptera, sex is determined by a haplodiploid system, which means that females are unusually closely related to their sisters, enabling kin selection to favour the evolution of eusocial behaviour. Females are diploid, meaning that they have 2n chromosomes and develop from fertilized eggs. Males, called drones, have a haploid ( n ) number of chromosomes and develop from an unfertilized egg. Wasps store sperm inside their body and control its release for each individual egg as it is laid ; if a female wishes to produce a male egg, she simply lays the egg without fertilizing it. Therefore, under most conditions in most species, wasps have complete voluntary control over the sex of their offspring. Experimental infection of Muscidifurax uniraptor with the bacterium Wolbachia induced thelytokous reproduction and an inability to produce fertile, viable male offspring. Inbreeding avoidance Females of the solitary wasp parasitoid Venturia canescens can avoid mating with their brothers through kin recognition. In experimental comparisons, the probability that a female will mate with an unrelated male was about twice as high as the chance of her mating with brothers. Female wasps appear to recognize siblings on the basis of a chemical signature carried or emitted by males. Sibling - mating avoidance reduces inbreeding depression that is largely due to the expression of homozygous deleterious recessive mutations. Ecology As pollinators Minute pollinating fig wasps, Pleistodontes : the trees and wasps have coevolved and are mutualistic. While the vast majority of wasps play no role in pollination, a few species can effectively transport pollen and pollinate several plant species. Since wasps generally do not have a fur - like covering of soft hairs and a special body part for pollen storage ( Pollen basket ) as some bees do, pollen does not stick to them well. However it has been shown that even without hairs, several wasp species are able to effectively transport pollen, therefore contributing for potential pollination of several plant species. Pollen wasps in the subfamily Masarinae gather nectar and pollen in a crop inside their bodies, rather than on body hairs like bees, and pollinate flowers of Penstemon and the water leaf family, Hydrophyllaceae. The Agaonidae ( fig wasps ) are the only pollinators of nearly 1000 species of figs, and thus are crucial to the survival of their host plants. Since the wasps are equally dependent on their fig trees for survival, the coevolved relationship is fully mutualistic. As parasitoids The parasitoidal ichneumon wasp Dolichomitus imperator ovipositing through wood, using its immensely long ovipositor to lay its eggs inside hidden larvae, detected by vibration Latina rugosa planidia ( arrows, magnified ) attached to an ant larva ; the Eucharitidae are among the few parasitoids able to overcome the strong defences of ants. Main article : Parasitoid wasp Most solitary wasps are parasitoids. As adults, those that do feed typically only take nectar from flowers. Parasitoid wasps are extremely diverse in habits, many laying their eggs in inert stages of their host ( egg or pupa ), sometimes paralysing their prey by injecting it with venom through their ovipositor. They then insert one or more eggs into the host or deposit them upon the outside of the host. The host remains alive until the parasitoid larvae pupate or emerge as adults. The Ichneumonidae are specialized parasitoids, often of Lepidoptera larvae deeply buried in plant tissues, which may be woody. For this purpose, they have exceptionally long ovipositors ; they detect their hosts by smell and vibration. Some of the largest species, including Rhyssa persuasoria and Megarhyssa macrurus, parasitise horntails, large sawflies whose adult females also have impressively long ovipositors. Some parasitic species have a mutualistic relationship with a polydnavirus that weakens the host 's immune system and replicates in the oviduct of the female wasp. One family of chalcid wasps, the Eucharitidae, has specialized as parasitoids of ants, most species hosted by one genus of ant. Eucharitids are among the few parasitoids that have been able to overcome ants ' effective defences against parasitoids. The Chrysididae, such as this Hedychrum rutilans, are known as cuckoo or jewel wasps for their parasitic behaviour and metallic iridescence. As parasites Further information : kleptoparasitism and cuckoo wasp Many species of wasp, including especially the cuckoo or jewel wasps ( Chrysididae ), are kleptoparasites, laying their eggs in the nests of other wasp species to exploit their parental care. Most such species attack hosts that provide provisions for their immature stages ( such as paralyzed prey items ), and they either consume the provisions intended for the host larva, or wait for the host to develop and then consume it before it reaches adulthood. An example of a true brood parasite is the paper wasp Polistes sulcifer, which lays its eggs in the nests of other paper wasps ( specifically Polistes dominula ), and whose larvae are then fed directly by the host. Sand wasps Ammophila often save time and energy by parasitising the nests of other females of their own species, either kleptoparasitically stealing prey, or as brood parasites, removing the other female 's egg from the prey and laying their own in its place. According to Emery 's rule, social parasites, especially among insects, tend to parasitise species or genera to which they are closely related. For example, the social wasp Dolichovespula adulterina parasitises other members of its genus such as D. norwegica and D. arenaria. As predators European beewolf Philanthus triangulum provisioning her nest with a honeybee Many wasp lineages, including those in the families Vespidae, Crabronidae, Sphecidae, and Pompilidae, attack and sting prey items that they use as food for their larvae ; while Vespidae usually macerate their prey and feed the resulting bits directly to their brood, most predatory wasps paralyze their prey and lay eggs directly upon the bodies, and the wasp larvae consume them. Apart from collecting prey items to provision their young, many wasps are also opportunistic feeders, and will suck the body fluids of their prey. Although vespid mandibles are adapted for chewing and they appear to be feeding on the organism, they are often merely macerating it into submission. The impact of the predation of wasps on economic pests is difficult to establish. The roughly 140 species of beewolf ( Philanthinae ) hunt bees, including honeybees, to provision their nests ; the adults feed on nectar and pollen. As models for mimics The wasp beetle Clytus arietis is one of many Batesian mimics of wasps. Further information : mimicry and aposematism With their powerful stings and conspicuous warning coloration, social wasps are the models for many species of mimic. Two common cases are Batesian mimicry, where the mimic is harmless and is essentially bluffing, and Müllerian mimicry, where the mimic is also distasteful, and the mimicry can be considered mutual. Batesian mimics of wasps include many species of hoverfly and the wasp beetle. Many species of wasp are involved in Müllerian mimicry, as are many species of bee. Bee - eaters such as Merops apiaster specialise in feeding on bees and wasps. As prey While wasp stings deter many potential predators, bee - eaters ( in the bird family Meropidae ) specialise in eating stinging insects, making aerial sallies from a perch to catch them, and removing the venom from the stinger by repeatedly brushing the prey firmly against a hard object, such as a twig. The honey buzzard attacks the nests of social hymenopterans, eating wasp larvae ; it is the only known predator of the dangerous Asian giant hornet or `` yak - killer '' ( Vespa mandarinia ). Likewise, Roadrunners are the only real predators of tarantula hawk wasps. Relationship with humans Encarsia formosa, a parasitoid, is sold commercially for biological control of whitefly, an insect pest of tomato and other horticultural crops. Tomato leaf covered with nymphs of whitefly parasitised by Encarsia formosa As pests Social wasps are considered pests when they become excessively common, or nest close to buildings. People are most often stung in late summer, when wasp colonies stop breeding new workers ; the existing workers search for sugary foods and are more likely to come into contact with humans ; if people then respond aggressively, the wasps sting. Wasp nests made in or near houses, such as in roof spaces, can present a danger as the wasps may sting if people come close to them. Stings are usually painful rather than dangerous, but in rare cases, people may suffer life - threatening anaphylactic shock. In horticulture Further information : biological pest control Some species of parasitic wasp, especially in the Trichogrammatidae, are exploited commercially to provide biological control of insect pests. For example, in Brazil, farmers control sugarcane borers with the parasitic wasp Trichogramma galloi. One of the first species to be used was Encarsia formosa, a parasitoid of a range of species of whitefly. It entered commercial use in the 1920s in Europe, was overtaken by chemical pesticides in the 1940s, and again received interest from the 1970s. Encarsia is used especially in greenhouses to control whitefly pests of tomato and cucumber, and to a lesser extent of aubergine ( eggplant ), flowers such as marigold, and strawberry. Several species of parasitic wasp are natural predators of aphids and can help to control them. For instance, Aphidius matricariae is used to control the peach - potato aphid. In sport Wasps RFC is an English professional rugby union team originally based in London but now playing in Coventry ; the name dates from 1867 at a time when names of insects were fashionable for clubs. The club 's first kit is black with yellow stripes. The club has an amateur side called Wasps FC. Among the other clubs bearing the name are a basketball club in Wantirna, Australia, and Alloa Athletic F.C., a football club in Scotland. In fashion Wasp waist, c. 1900, demonstrated by Polaire, a French actress famous for this silhouette Wasps have been modelled in jewellery since at least the nineteenth century, when diamond and emerald wasp brooches were made in gold and silver settings. A fashion for wasp waisted female silhouettes with sharply cinched waistlines emphasizing the wearer 's hips and bust arose repeatedly in the nineteenth and twentieth centuries. In literature The Ancient Greek playwright Aristophanes wrote the comedy play Σφῆκες ( Sphēkes ), The Wasps, first put on in 422 BC. The `` wasps '' are the chorus of old jurors. H.G. Wells made use of giant wasps in his novel The Food of the Gods and How It Came to Earth ( 1904 ) : It flew, he is convinced, within a yard of him, struck the ground, rose again, came down again perhaps thirty yards away, and rolled over with its body wriggling and its sting stabbing out and back in its last agony. He emptied both barrels into it before he ventured to go near. When he came to measure the thing, he found it was twenty - seven and a half inches across its open wings, and its sting was three inches long... The day after, a cyclist riding, feet up, down the hill between Sevenoaks and Tonbridge, very narrowly missed running over a second of these giants that was crawling across the roadway. Wasp ( 1957 ) is a science fiction book by the English writer Eric Frank Russell ; it is generally considered Russell 's best novel. In Stieg Larsson 's book The Girl Who Played with Fire ( 2006 ) and its film adaptation, Lisbeth Salander has adopted her kickboxing ringname, `` The Wasp '', as her hacker handle and has a wasp tattoo on her neck, indicating her high status among hackers, unlike her real world situation, and that like a small but painfully stinging wasp, she could be dangerous. Parasitoidal wasps played an indirect role in the nineteenth - century evolution debate. The Ichneumonidae contributed to Charles Darwin 's doubts about the nature and existence of a well - meaning and all - powerful Creator. In an 1860 letter to the American naturalist Asa Gray, Darwin wrote : I own that I can not see as plainly as others do, and as I should wish to do, evidence of design and beneficence on all sides of us. There seems to me too much misery in the world. I can not persuade myself that a beneficent and omnipotent God would have designedly created the Ichneumonidae with the express intention of their feeding within the living bodies of caterpillars, or that a cat should play with mice. In military names HMS Wasp ( 1880 ), one of nine Royal Navy warships to bear the name With its powerful sting and familiar appearance, the wasp has given its name to many ships, aircraft and military vehicles. Nine ships and one shore establishment of the Royal Navy have been named HMS Wasp, the first an 8 - gun sloop launched in 1749. Eleven ships of the United States Navy have similarly borne the name USS Wasp, the first a merchant schooner acquired by the Continental Navy in 1775. The eighth of these, an aircraft carrier, gained two Second World War battle stars, prompting Winston Churchill to remark `` Who said a Wasp could n't sting twice ? '' In the Second World War, a German self - propelled howitzer was named Wespe, while the British developed the Wasp flamethrower from the Bren Gun Carrier. In aerospace, the Westland Wasp was a military helicopter developed in England in 1958 and used by the Royal Navy and other navies. The AeroVironment Wasp III is a miniature UAV developed for United States Air Force special operations. See also Bee and wasp stings Characteristics of common wasps and bees Schmidt sting pain index Notes Jump up ^ Specimen measured from photograph. Jump up ^ Methods to estimate species diversity include extrapolating the rate of species descriptions by subfamily ( as in the Braconidae ) until zero is reached ; and extrapolating geographically from the species distribution of well - studied taxa to the group of interest ( say, the Braconidae ). 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The Guardian. 19 August 2009. Retrieved 10 June 2015. Jump up ^ `` Wasps ''. British Pest Control Association. Retrieved 5 August 2015. Jump up ^ `` Allergy to Wasp and Bee Stings ''. Allergy UK. Retrieved 5 August 2015. Jump up ^ `` New wasp parasite being studied ''. The Royal Society of New Zealand. 20 April 2000. Retrieved 15 July 2013. Jump up ^ Moskvitch, Katia ( 22 January 2013 ). `` Wasps v moths : Biocontrol uses nature against crop pests ''. BBC. Retrieved 10 June 2015. Jump up ^ Hoddle, Mark. `` Encarsia formosa ''. Cornell University. Retrieved 12 June 2015. Jump up ^ `` Aphid Predators ''. Royal Horticultural Society. Retrieved 5 August 2015. Jump up ^ C.R. Adams ; K.M. Bamford ; M.P. Early ( 22 October 2013 ). Principles of Horticulture. Elsevier. pp. 71 -- 72. ISBN 978 - 1 - 4831 - 4184 - 8. Jump up ^ `` History 1867 -- 1930 London Wasps ''. Wasps.co.uk. Archived from the original on 22 July 2014. Jump up ^ `` Wasps Football Club ''. Pitcher. Retrieved 5 August 2015. Jump up ^ `` Wantirna Wasps Basketball Club ''. Retrieved 5 August 2015. Jump up ^ `` Alloa Athletic Football Club ''. Scottish Professional Football League. Retrieved 10 September 2015. Jump up ^ `` Diamond and emerald wasp brooch by Fontanna ''. A La Vieille Russie. Archived from the original on 10 June 2015. Retrieved 10 June 2015. Crown rose diamond and emerald wasp brooch set in silver and gold. By Fontana French, ca. 1875. Width : 3 inches. $46,500 Jump up ^ Kunzle, David. `` Fashion and Fetishism ''. Archived from the original on 21 November 2014. Retrieved 17 June 2015. Jump up ^ Klingerman, Katherine Marie ( May 2006 ). Binding Femininity : The Effects of Tightlacing on the Female Pelvis ( PDF ). University of Vermont ( MA Thesis ). Jump up ^ `` Ancient Greece -- Aristophanes -- The Wasps ''. Retrieved 10 June 2015. ^ Jump up to : Wells, H.G. ( 1904 ). Food of the Gods. Macmillan. Jump up ^ Russell, Eric Frank ( 1957 ). Wasp. Gollancz Science Fiction. ISBN 0 - 575 - 07095 - 1. Jump up ^ Rosenberg, Robin S. ; O'Neill, Shannon ; McDonald - Smith, Lynne ( 2013 ). The Psychology of the Girl with the Dragon Tattoo : Understanding Lisbeth Salander and Stieg Larsson 's Millennium Trilogy. BenBella Books. p. 34. ISBN 978 - 1 - 936661 - 35 - 0. Jump up ^ `` Letter 2814 -- Darwin, C.R. to Gray, Asa, ''. 22 May 1860. Retrieved 4 May 2011. ^ Jump up to : `` USS Wasp. History ''. United States Navy. Retrieved 1 July 2016. Jump up ^ Colledge, J.J. ; Warlow, Ben ( 2006 ) ( 1969 ). Ships of the Royal Navy : The Complete Record of all Fighting Ships of the Royal Navy ( Rev. ed. ). London : Chatham Publishing. ISBN 978 - 1 - 86176 - 281 - 8. OCLC 67375475. Jump up ^ USS Wasp Veterans ( 15 June 1999 ). USS Wasp. Turner Publishing. pp. 8 -- 11. ISBN 978 - 1 - 56311 - 404 - 5. Jump up ^ Jentz, Thomas L. ; Doyle, Hilary Louis ( 2011 ). Panzer Tracts No. 23 : Panzer Production From 1933 to 1945. Panzer Tracts. Jump up ^ Bishop, Chris ( 2002 ). The Encyclopedia of Weapons of World War II. Sterling Publishing. p. 272. ISBN 9781586637620. Jump up ^ James, Derek N. ( 1991 ). Westland Aircraft since 1915. Putnam. p. 365. ISBN 0 - 85177 - 847 - X. Jump up ^ `` US Air Force Wasp III Fact Sheet ''. Archived from the original on 23 September 2015. Retrieved 12 June 2015. Sources Ross, Kenneth G. ( 1991 ). The Social Biology of Wasps. Cornell Press. ISBN 978 - 0 - 801 - 49906 - 7. External links Wikimedia Commons has media related to Wasp. Wikimedia Commons has media related to Wasp nests. Wikisource has the text of the 1911 Encyclopædia Britannica article Wasp. Differences between Bees and Wasps A pictorial life cycle of organ pipe wasps N.I.H. Medline Encyclopedia -- Insect bites and stings Waspweb DermNet arthropods / bites Extant Hymenopteran families Kingdom : Animalia Phylum : Arthropoda Class : Insecta Subclass : Pterygota Infraclass : Neoptera Superorder : Endopterygota y m p h y Xyeloidea Xyelidae Pamphilioidea Megalodontesidae Pamphiliidae ( web - spinning sawflies ) Tenthredinoidea Argidae ( argid sawflies ) Blasticotomidae ( fern sawflies ) Cimbicidae ( cimbicid sawflies ) Diprionidae ( conifer sawflies ) Pergidae ( pergid sawflies ) Tenthredinidae ( common sawflies ) U n i c. Cephoidea Cephidae ( stem sawflies ) Siricoidea Anaxyelidae ( cedar wood wasps ) Siricidae ( horntails ) Xiphydrioidea Xiphydriidae ( wood wasps ) Orussoidea Orussidae ( parasitic wood wasps ) p o r i U n i l r i d r s i i Ichneumonoidea Braconidae ( braconids ) Ichneumonidae ( ichneumon wasps ) Ceraphronoidea Ceraphronidae Megaspilidae Proctotrupomorpha Platygastroidea Platygastridae Scelionidae Cynipoidea Austrocynipidae Cynipidae ( gall wasps ) Figitidae Ibaliidae Liopteridae Proctotrupoidea ( s. str. ) Proctorenyxidae Roproniidae Heloridae Pelecinidae Peradeniidae Proctotrupidae Vanhorniidae Diaprioidea Austroniidae Diapriidae Maamingidae Monomachidae Mymarommatoidea Mymarommatidae Chalcidoidea ( chalcid wasps ) Agaonidae ( fig wasps ) Aphelinidae Chalcididae Encyrtidae Eucharitidae Eulophidae Eupelmidae Eurytomidae Leucospidae Mymaridae ( fairyflies ) Ormyridae Perilampidae Pteromalidae Rotoitidae Signiphoridae Tanaostigmatidae Tetracampidae Torymidae Trichogrammatidae Evanioidea Aulacidae Evaniidae ( ensign wasps ) Gasteruptiidae Stephanoidea Stephanidae Megalyroidea Megalyridae Trigonaloidea Trigonalidae u l Chrysidoidea Bethylidae Chrysididae ( cuckoo wasps ) Dryinidae Embolemidae Plumariidae Sclerogibbidae Scolebythidae Vespoidea Rhopalosomatidae ( rhopalosomatid wasps ) Vespidae ( paper wasps, potter wasps, pollen wasps, yellowjackets, hornets ) Tiphioidea Bradynobaenidae Tiphiidae ( tiphiid wasps ) Thynnoidea Chyphotidae Thynnidae ( flower wasps ) Sierolomorphidae ( sierolomorphid wasps ) Pompiloidea Mutillidae ( velvet ants ) Myrmosidae Pompilidae ( spider wasps ) Sapygidae ( sapygid, or club - horned wasps ) Scolioidea Scoliidae ( scoliid wasps ) Formicoidea Formicidae ( ants ) Apoidea Spheciformes ( sphecoid wasps ) Ampulicidae ( cockroach wasps ) Crabronidae ( sand wasps, bee wolves ) Heterogynaidae Sphecidae ( thread - waisted wasps ) Anthophila ( bees ) Andrenidae ( mason bees ) Apidae ( honey bees, bumblebees, cuckoo bees, carpenter bees, orchid bees, stingless bees ) Colletidae ( plasterer bees ) Halictidae ( sweat bees ) Megachilidae ( mason bees, leafcutter bees ) Melittidae Stenotritidae Italic are paraphyletic groups Based on Malm and Nyman ( 2015 ) and Peters et al ( 2017 ) Eusociality Topics Evolution of eusociality Presociality Social insects Gamergate Group selection Haplodiploidy Identity in social insects Kin recognition Kin selection Sexual selection in social insects Thelytoky Worker policing Groups Hymenoptera Ant Apidae Crabronidae Halictidae Honey bee Vespidae Mammalia Blesmol Dwarf mongoose Meerkat Crustacea Synalpheus Thysanoptera Kladothrips Hemiptera Aphididae Coleoptera Austroplatypus incompertus Isoptera In culture Bee ( mythology ) Pioneers, works Karl von Frisch The Dancing Bees 1927 Charles Duncan Michener The Bees of the World 2000 E.O. Wilson The Ants 1990 Sociobiology : The New Synthesis 1975 Insects in culture Aspects of insects in culture In the arts Insects in art Beetlewing John Hampson Insects in film Insects in literature Insects in music List of insect - inspired songs Insects on stamps In fishing Fishing bait Fly fishing Artificial fly Fly tying Maggot Mayfly Mealworm In medicine Apitherapy Apitoxin Melittin Maggot Spanish fly Cantharidin In mythology Bee Butterfly Cicada Dragonfly Praying mantis Scarab Entomophagy ( as food ) Adults Ant Cicada Cricket Grasshopper Termite Larvae Bamboo worm Mealworm Mopani worm Palm weevil Rhinoceros beetle Silkworm Waxworm Witchetty grub Other aspects Biomimicry Cricket fighting Entomological warfare Flea circus Insects in religion Jingzhe Economic entomology Beneficial insects Pest control Encarsia formosa Ladybird Parasitoid wasp Pollination Bees crops pollinated Bumblebee Honey bee western Beetles Flies Lepidoptera Products Beekeeping Bee pollen Beeswax Honey Propolis Royal jelly Other insects Carmine / Cochineal Polish Chitin Kermes Sericulture Silk Lac / Shellac Model organism Drosophila melanogaster Harmful insects Crop pests Aphid Boll weevil Colorado potato beetle Cottony cushion scale Japanese beetle Locust Phylloxera Western corn rootworm Livestock pests Botfly Horn fly Horse - fly Screwworm fly Tsetse fly Warble fly Biting / stinging Insect bites and stings Insect sting allergy Bed bug Bee sting Flea Horse - fly Louse Mosquito Wasp Wood - eating Deathwatch beetle Furniture beetle House longhorn beetle Termite Woodworm Other pests Home - stored product entomology Clothes moth Cockroach Housefly Pioneers Jan Swammerdam Alfred Russel Wallace Jean - Henri Fabre Hans Zinsser ( Rats, Lice and History ) Lafcadio Hearn ( Insect Literature ) Related Living things in culture Arthropods Birds Fish Fungi Mammals Microbes Molluscs Reptiles Plants Zoomusicology BNF : cb12122999w ( data ) NDL : 00563011 Retrieved from `` https://en.wikipedia.org/w/index.php?title=Wasp&oldid=842806556 '' Categories : Wasps Pest insects Biological pest control wasps Extant Jurassic first appearances Paraphyletic groups Insects in culture Hidden categories : CS1 Spanish - language sources ( es ) CS1 Portuguese - language sources ( pt ) CS1 : Julian -- Gregorian uncertainty Good articles Use dmy dates from January 2015 Articles with ' species ' microformats Commons category with local link different than on Wikidata Wikipedia articles with BNF identifiers Use dmy dates from May 2011 Talk Contents About Wikipedia ܐܪܡܝܐ Avañe'ẽ বাংলা Bahasa Banjar Bân - lâm - gú Башҡортса Беларуская Català Чӑвашла ChiShona Eesti Ελληνικά Español Esperanto Euskara فارسی Français Gaeilge Galego 客家 語 / Hak - kâ - ngî Հայերեն हिन्दी Ido Bahasa Indonesia Iñupiak Íslenska עברית Kapampangan ქართული Kaszëbsczi Kiswahili Коми Kreyòl ayisyen Latina Latviešu Lietuvių Lingála Livvinkarjala മലയാളം Bahasa Melayu Mìng - dĕ̤ng - ngṳ̄ नेपाली 日本 語 Nouormand Occitan Олык марий ਪੰਜਾਬੀ پنجابی Перем Коми Picard Português Română Runa Simi Русский Scots Simple English Српски / srpski Srpskohrvatski / српскохрватски Basa Sunda தமிழ் తెలుగు ไทย Tsetsêhestâhese Türkçe Українська اردو Vahcuengh Vepsän kel ' Tiếng Việt Walon Winaray Wolof 粵語 Žemaitėška 中文 67 more Edit links This page was last edited on 24 May 2018, at 20 : 19. 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Question: what do you call a group of wasps
Answer: paraphyletic