Orb - weaver spider - wikipedia Orb - weaver spider Orb - weaver spider Argiope catenulata Scientific classification Kingdom : Animalia Phylum : Arthropoda Subphylum : Chelicerata Class : Arachnida Order : Araneae Infraorder : Araneomorphae Superfamily : Araneoidea Family : Araneidae Clerck, 1757 Genera See List of Araneidae genera. Diversity 172 genera, 3122 species Orb - weaver spiders or araneids are members of the spider family Araneidae. They are the most common group of builders of spiral wheel - shaped webs often found in gardens, fields and forests. `` Orb '' was previously used in English to mean `` circular '', hence the English name of the group. Araneids have eight similar eyes, hairy or spiny legs, and no stridulating organs. The family is cosmopolitan, including many well - known large or brightly colored garden spiders. With 3122 species in 172 genera worldwide, Araneidae is the third - largest family of spiders ( behind Salticidae and Linyphiidae ). Araneid webs are constructed in a stereotyped fashion. A framework of nonsticky silk is built up before the spider adds a final spiral of silk covered in sticky droplets. Orb - webs are also produced by members of other spider families. The long - jawed orb weavers ( Tetragnathidae ) were formerly included in the Araneidae ; they are closely related, being part of the superfamily Araneoidea. The family Arkyidae has been split off from the Araneidae. The cribellate or hackled orb - weavers ( Uloboridae ) belong to a different group of spiders. Their webs are strikingly similar, but use a different kind of sticky silk. Contents 1 Web 2 Natural history 3 Reproduction 3.1 Sexual Size Dimorphism 4 Systematics 5 See also 6 References 7 Further reading 8 External links Web Argiope sp. sitting on the stabilimentum at the center of the web Spiderlings in the web near where they hatched Close - up of the cephalothorax on Eriophora sp. ( possibly E. heroine or E. pustuosa ) Generally, orb - weaving spiders are three - clawed builders of flat webs with sticky spiral capture silk. The building of a web is an engineering feat, begun when the spider floats a line on the wind to another surface. The spider secures the line and then drops another line from the center, making a `` Y ''. The rest of the scaffolding follows with many radii of nonsticky silk being constructed before a final spiral of sticky capture silk. The third claw is used to walk on the nonsticky part of the web. Characteristically, the prey insect that blunders into the sticky lines is stunned by a quick bite, and then wrapped in silk. If the prey is a venomous insect, such as a wasp, wrapping may precede biting and / or stinging. Many orb - weavers build a new web each day. Most orb - weavers tend to be active during the evening hours ; they hide for most of the day. Generally, towards evening, the spider will consume the old web, rest for approximately an hour, then spin a new web in the same general location. Thus, the webs of orb - weavers are generally free of the accumulation of detritus common to other species, such as black widow spiders. Some orb - weavers do not build webs at all. Members of the genera Mastophora in the Americas, Cladomelea in Africa, and Ordgarius in Australia produce sticky globules, which contain a pheromone analog. The globule is hung from a silken thread dangled by the spider from its front legs. The pheromone analog attracts male moths of only a few species. These get stuck on the globule and are reeled in to be eaten. Interestingly, both types of bolas spiders are highly camouflaged and difficult to locate. The spiny orb - weaving spiders in the genera Gasteracantha and Micrathena look like plant seeds or thorns hanging in their orb - webs. Some species of Gasteracantha have very long, horn - like spines protruding from their abdomens. One feature of the webs of some orb - weavers is the stabilimentum, a crisscross band of silk through the center of the web. It is found in a number of genera, but Argiope, the yellow and banded garden spiders of North America, is a prime example. The band has been hypothesized to be a lure for prey, a marker to warn birds away from the web, and a camouflage for the spider when it sits in the center of the web. However, recent research suggests the stabilimentum actually decreases the visibility of the silk to insects, thus making it harder for prey to avoid the web. The orb - web consists of a frame and supporting radii overlaid with a sticky capture spiral, and the silks used by orb - weaver spiders have exceptional mechanical properties to withstand the impact of flying prey. During the Cretaceous, a radiation of angiosperm plants and their insect pollinators occurred. Fossil evidence shows that the orb web was in existence at this time, which permitted a concurrent radiation of the spider predators along with their insect prey. The capacity of orb -- webs to absorb the impact of flying prey led orbicularian spiders to become the dominant predators of aerial insects in many ecosystems. Insects and spiders have comparable rates of diversification, suggesting they co-radiated, and the peak of this radiation occurred 100 Mya before the origin of angiosperms. Vollrath and Selden ( 2007 ) make the bold proposition that insect evolution was driven less by flowering plants than by spider predation -- particularly through orb webs -- as a major selective force. Most arachnid webs are vertical and the spiders usually hang with their head downward. A few webs, such as those of orb - weavers in the genus Metepeira have the orb hidden within a tangled space of web. Some Metepiera are semisocial and live in communal webs. In Mexico, such communal webs have been cut out of trees or bushes and used for living fly paper. In 2009, workers at a Baltimore Wastewater Treatment Plant called for help to deal with over 100 million orb - weaver spiders, living in a community that managed to spin a phenomenal web that covered some 4 acres of a building with spider densities in some areas reaching 35,176 spiders per cubic meter. Natural History Argiope lobata in Southern Spain The oldest known true orb - weaver is Mesozygiella dunlopi, from the Lower Cretaceous. Several fossils provide direct evidence that the three major orb - weaving families, namely Araneidae, Tetragnathidae and Uloboridae, had evolved by this time, about 140 million years ago. They probably originated during the Jurassic ( 200 to 140 million years ago ). Based on new molecular evidence in silk genes, all three families are likely to have a common origin. The two families, Deinopoidea and Araneoidea, have similar behavioral sequences and spinning apparatuses to produce architecturally similar webs. The Araneidae weave true viscid silk with an aqueous glue property, and the Deinopoidea use dry fibrils and sticky silk. The Deinopoidea ( including the Uloboridae ), have a cribellum -- a flat, complex spinning plate from which the cribellate silk is released. They also have a calamistrum -- an apparatus of bristles used to comb the cribellate silk from the cribellum. The Araneoidea, or the `` ecribellate '' spiders, do not have these two structures. The two families of orb - weaving spiders are morphologically very distinct, yet there is much similarity between their web form and web construction behavior. The cribellates retained the ancestral character, yet the cribellum was lost in the escribellates. The lack of a functional cribellum in araneoids is most likely synapomorphic. If the orb - weaver spiders are a monophyletic group, the fact that only some species in the group lost a feature adds to the controversy. The cribellates are split off as a separate taxon that retained the primitive feature, which makes the lineage paraphyletic and not synonymous with any real evolutionary lineage. The morphological and behavioral evidence surrounding orb webs led to the disagreement over a single origin or a dual origin. However, molecular analysis provides more support for a monophyletic origin. Reproduction Araneids species either mate at the central hub of the web, where the male slowly traverses the web, trying not to get eaten, and when reaching the hub, mounts the female ; or the male constructs a mating thread inside or outside the web to attract the female via vibratory courtship, and if successful, mating occurs on the thread. In the cannibalistic and polyandrous orb - web spider Argiope bruennichi, the much smaller males are attacked during their first copulation and are cannibalized in up to 80 % of the cases. All surviving males die after their second copulation, a pattern observed on other Argiope species. Whether a male survives his first copulation depends on the duration of the genital contact : males that jump off early ( before 5 seconds ) have a chance of surviving, while males that copulate longer ( greater than 10 seconds ) invariably die. Prolonged copulation, although associated with cannibalism, enhances sperm transfer and relative paternity. It was observed that when males mated with a non-sibling female, the duration of their copulation was prolonged, and consequently the males were cannibalized more frequently. When males mated with a sibling female, they copulated briefly, and thus were more likely to escape cannibalism. By escaping, their chance of mating again with a non-kin female likely would be increased. These observations suggest that males can adaptively adjust their investment based on the degree of genetic relatedness of the female in order to avoid inbreeding depression. Sexual size dimorphism Sexual dimorphism refers to physical differences between males and females of the same species. One such difference can be in size. Araneids often exhibit size dimorphism typically known as extreme sexual size dimorphism, due to the extent of differences in size. The size difference among species of Araneidae ranges greatly. Some females, such as those of the Nephila pilipes, can be at least 9 times larger than the male, while others are only slightly larger than the male. The larger size female is typically thought to be selected through fecundity selection, the idea that bigger females can produce more eggs, thus more offspring. Although a great deal of evidence points towards the greatest selection pressure on larger female size, there is some evidence that selection can favor small male size as well. Araneids also exhibit a phenomenon called sexual cannibalism, which is commonly found throughout Araneidae. Evidence suggests a negative correlation between sexual size dimorphism and incidences of sexual cannibalism. Other evidence, however, has shown that differences in cannibalistic events among araneids when having smaller or slightly larger males is advantageous. Some evidence has shown that extreme dimorphism may be the result of males avoiding detection by the females. For males of these species, being smaller in size may be advantageous in moving the central hub of a web so female spiders may be less likely to detect the male, or even if detected as prey to be eaten, the small size may indicate little nutritional value. Larger -- bodied male araneids may be advantageous when mating on a mating thread because the thread is constructed from the edge of the web orb to structural threads or to nearby vegetation. Here larger males may be less likely to be cannibalized, as the males are able to copulate while the female is hanging, which may make them safer from cannibalism. In one subfamily of Araneid that uses a mating thread, Gasteracanthinae, sexual cannibalism is apparently absent despite extreme size dimorphism. Systematics. Main article : List of Araneidae genera Eriophora transmarina from Australia Argiope lobata from Ukraine Eriovixia laglaizei As of August 2017, the World Spider Catalog placed 172 genera ( comprising over 3,100 species ) in the family Araneidae. See also Arthropods portal List of Araneidae species Spider families References Jump up ^ `` Family : Araneidae Clerck, 1757 ''. World Spider Catalog. Natural History Museum Bern. Retrieved 2016 - 10 - 01. ^ Jump up to : `` Currently valid spider genera and species ''. World Spider Catalog. Natural History Museum, Bern. Retrieved 16 August 2017. Jump up ^ `` orb ''. Merriam - Webster Dictionary. Retrieved 5 December 2015. Jump up ^ Dimitrov, Dimitar ; Benavides, Ligia R. ; Arnedo, Miquel A. ; Giribet, Gonzalo ; Griswold, Charles E. ; Scharff, Nikolaj & Hormiga, Gustavo ( 2016 ). `` Rounding up the usual suspects : a standard target - gene approach for resolving the interfamilial phylogenetic relationships of ecribellate orb - weaving spiders with a new family - rank classification ( Araneae, Araneoidea ) '' ( PDF ). Cladistics. doi : 10.1111 / cla. 12165. Retrieved 2016 - 10 - 18. Jump up ^ Blackledge, Todd A. & Wenzel, John W. ( 2000 ). `` The evolution of cryptic spider silk : a behavioral test ''. Behavioral Ecology. 11 ( 2 ) : 142 -- 145. doi : 10.1093 / beheco / 11.2. 142. ^ Jump up to : Garb, Jessica E. ; DiMauro, Teresa ; Vo, Victoria & Hayashi, Cheryl Y. ( 2006 ). `` Silk genes support the single origin of orb webs ''. Science. 312 : 1762. PMID 16794073. doi : 10.1126 / science. 1127946. Jump up ^ `` Detangling History ''. Smithsonian : 24. September 2006. `` Two pieces of Spanish amber contain the oldest known spider web and orb -- weaving spider ; both specimens are at least 110 million years old. The new findings, along with an analysis of the proteins in spider silk, indicate that orb -- weaving spiders date as far back as 144 million years. '' ^ Jump up to : Penney, David & Ortuño, Vicente M. ( 2006 ). `` Oldest true orb - weaving spider ( Araneae : Araneidae ) ''. Biology Letters. 2 : 447 -- 450. PMC 1686203. PMID 17148427. doi : 10.1098 / rsbl. 2006.0506. ^ Jump up to : Blackledge, Todd A. ; Scharff, Nikolaj ; Coddington, Jonathan A. ; Szüts, Tamas ; Wenzel, John W. ; Hayashi, Cheryl Y. & Agnarsson, Ingi ( 2009 ). `` Reconstructing web evolution and spider diversification in the molecular era ''. Proceedings of the National Academy of Sciences. 106 : 5229 -- 5234. PMC 2656561. PMID 19289848. doi : 10.1073 / pnas. 0901377106. Jump up ^ Vollrath, Fritz & Selden, Paul ( 2007 ). `` The role of behavior in the evolution of spiders, silks, and webs ''. Annual Review of Ecology, Evolution, and Systematics. 38 : 819 -- 846. doi : 10.1146 / annurev. ecolsys. 37.091305. 110221. Jump up ^ Alford, Justine ( 2 November 2014 ). `` Orb - Weaver Spiders Stuff A Treatment Plant With A 4 - Acre Web ''. IFLScience. Retrieved 6 April 2015. Jump up ^ Peñalver, Enrique ; Grimaldi, David A. & Delclòs, Xavier ( 2006 ). `` Early Cretaceous spider web with its prey ''. Science. 312 ( 5781 ) : 1761. PMID 16794072. doi : 10.1126 / science. 1126628. Jump up ^ Shear, William A. ( 1986 ). `` The evolution of web - building behavior in spiders : a third generation of hypotheses ''. In Shear, William A. Spiders : webs, behavior, and evolution. Stanford CA : Stanford University Press. pp. 364 -- 400. ISBN 978 - 0 - 8047 - 1203 - 3. ^ Jump up to : Coddington, Jonathan A. ( 1986 ). `` The monophyletic origin of the orb web ''. In Shear, William A. Spiders : webs, behavior, and evolution. Stanford CA : Stanford University Press. pp. 319 -- 363. ISBN 978 - 0 - 8047 - 1203 - 3. ^ Jump up to : Elgar, Mark ( 1991 ). `` Sexual Cannibalism, Size Dimorphism, and Courtship Behavior in Orb - Weaving Spiders ( Araneidae ) ''. Evolution. 45 : 444 -- 448. JSTOR 2409679. doi : 10.2307 / 2409679. ^ Jump up to : Schneider, J.M. ; Gilberg, S. ; Fromhage, L. & Uhl, G. ( 2006 ). `` Sexual conflict over copulation duration in a cannibalistic spider ''. Animal Behaviour. 71 : 781 -- 788. doi : 10.1016 / j. anbehav. 2005.05. 012. Jump up ^ Welke, K.W. & Schneider, J.M. ( 2010 ). `` Males of the orb - web spider Argiope bruennichi sacrifice themselves to unrelated females ''. Biol. Lett. 6 ( 5 ) : 585 -- 588. PMC 2936157. PMID 20410027. doi : 10.1098 / rsbl. 2010.0214. Jump up ^ Gustavo, H. ; Scharff, N. & Coddington, J. ( 2000 ). `` The Phylogenetic Basis of Sexual Size Dimorphism in Orb - Weaving Spiders ( Araneae, Orbiculariae ) '' ( PDF ). Systematic Biology. 49 ( 3 ) : 435 -- 462. doi : 10.1080 / 10635159950127330. ^ Jump up to : Legrand, R.S. ; Morse, D.H. ( 2000 ). `` Factors driving extreme sexual size dimorphism of a sit - and - wait predator under low density ''. Biological Journal of the Linnean Society. 71 : 643 -- 664. doi : 10.1111 / j. 1095 - 8312.2000. tb01283. x. Jump up ^ Elgar, M.A. ( 1990 ). `` Sexual dimorphism in leg - length among orb - weaving spiders : a possible role for sexual cannibalism. ''. Journal of Zoology ( London ). 220 : 455 -- 470. Further reading Wikimedia Commons has media related to Araneidae. Wikispecies has information related to : Araneidae Crompton, John ( 1950 ). The Life of the Spider. New York : Mentor. OCLC 610423670. Dondale, C.D. ; Redner, J.H. ; Paquin, P. ; Levi, H.W. ( 2003 ). The Orb - Weaving Spiders of Canada and Alaska. Araneae : Uloboridae, Tetragnathidae, Araneidae, Theridiosomatidae. Insects and Arachnids of Canada. 23. Ottawa : NRC Research Press. ISBN 978 - 0 - 660 - 18898 - 0. Kaston, B.J. ( 1953 ). How to Know the Spiders. Pictured key nature series ( 1st ed. ). Dubuque, IA : W.C. Brown Co. OCLC 681432632. Main, Barbara York ( 1976 ). Spiders. Australian naturalist library ( 2nd ed. ). Sydney : Collins. OCLC 849736139. Foelix, Rainer F. ( 1996 ). Biology of Spiders ( 2nd ed. ). New York : Oxford University Press. OCLC 300192823. Levi, H.W. ( 1993 ). `` The new orb - weaver genus Lewisepeira ( Araneae : Araneidae ) '' ( PDF ). Psyche. 100 : 127 -- 136. doi : 10.1155 / 1993 / 97657. External links Spiders of Australia Spiders of northwestern Europe Araneae, Arachnology Home Pages World Spider Catalog Orb weavers of Kentucky, University of Kentucky Pictures of Mangora species Gasteracantha cancriformis, spinybacked orbweaver on the University of Florida / Institute of Food and Agricultural Sciences Featured Creatures website Neoscona crucifera and N. domiciliorum on the University of Florida / Institute of Food and Agricultural Sciences Featured Creatures website HOE Taxon identifiers Wd : Q5883 ADW : Araneidae BugGuide : 1972 EoL : 8819 Fossilworks : 57485 GBIF : 7359 ITIS : 82737 NCBI : 6913 WSC : urn : lsid : nmbe.ch : spiderfam : 0102 World Spider Catalog : 0102 Extant Araneae families Kingdom : Animalia Phylum : Arthropoda Subphylum : Chelicerata Class : Arachnida Suborder Mesothelae Liphistiidae ( segmented spiders ) Suborder Opisthothelae y g l o m o r p h Actinopodidae ( mouse spiders and relatives ) Antrodiaetidae ( folding trapdoor spiders ) Atypidae ( atypical tarantulas or purseweb spiders ) Barychelidae ( brushed trapdoor spiders ) Ctenizidae ( cork - lid trapdoor spiders ) Cyrtaucheniidae ( wafer trapdoor spiders ) Dipluridae ( funnel - web tarantulas ) Euctenizidae Hexathelidae ( funnel - webs or venomous funnel - web tarantulas ) Idiopidae Mecicobothriidae ( dwarf tarantulas ) Microstigmatidae Migidae ( tree trapdoor spiders ) Nemesiidae ( funnel - web tarantulas ) Paratropididae ( bald - legged spiders ) Theraphosidae ( true tarantulas ) r n o m o r p h o n - n l y n s Austrochilidae Caponiidae Diguetidae ( coneweb spiders ) Drymusidae ( false violin spiders ) Dysderidae ( woodlouse hunters ) Filistatidae ( crevice weaver spiders ) Gradungulidae ( large - clawed spiders ) Hypochilidae ( lampshade spiders ) Leptonetidae ( leptonetid spiders ) Ochyroceratidae ( midget ground weavers ) Oonopidae ( goblin spiders ) Orsolobidae Periegopidae Pholcidae ( cellar spiders ) Plectreuridae ( plectreurid spiders ) Scytodidae ( spitting spiders ) Segestriidae ( tube - dwelling spiders ) Sicariidae ( violin spiders, assassin spiders ) Telemidae ( long - legged cave spiders ) Tetrablemmidae ( armored spiders ) Trogloraptoridae ( Trogloraptor marchingtoni ) n l g y n Agelenidae ( araneomorph funnel weavers ) Amaurobiidae ( tangled nest spiders ) Ammoxenidae ( termite hunters ) Amphinectidae ( including Neolanidae ) Anapidae Anyphaenidae ( anyphaenid sac spiders ) Araneidae ( orb - weaver spiders ) Archaeidae ( pelican spiders ) Chummidae Cithaeronidae Clubionidae ( sac spiders ) Corinnidae ( dark sac spiders ) Ctenidae ( wandering spiders or tropical wolf spiders ) Cyatholipidae Cybaeidae Cycloctenidae Deinopidae ( net - casting spiders ) Desidae ( intertidal spiders ) Dictynidae ( dictynid spiders ) Eresidae ( velvet spiders ) Eutichuridae Gallieniellidae Gnaphosidae ( flat - bellied ground spiders ) Hahniidae ( dwarf sheet spiders ) Hersiliidae ( tree trunk spiders ) Holarchaeidae Homalonychidae Huttoniidae Lamponidae ( white - tailed spiders ) Linyphiidae ( sheet weavers or money spiders ) Liocranidae ( liocranid sac spiders ) Lycosidae ( wolf spiders ) Malkaridae ( shield spiders ) Mecysmaucheniidae Mimetidae ( pirate spiders ) Miturgidae ( long - legged sac spiders ) Mysmenidae ( spurred orb - weavers ) Nesticidae ( cave cobweb spiders ) Nicodamidae ( red and black spiders ) Oecobiidae ( disc web spiders ) Oxyopidae ( lynx spiders ) Palpimanidae ( palp - footed spiders ) Pararchaeidae Penestomidae Philodromidae ( running crab spiders ) Phrurolithidae Phyxelididae Pimoidae Pisauridae ( nursery web spiders ) ( including Halidae ) Prodidomidae ( long - spinneret ground spiders ) Psechridae Salticidae ( jumping spiders ) Selenopidae ( wall crab spiders ) Senoculidae ( bark hunters ) Sinopimoidae ( member of Linyphiidae ? ) Sparassidae ( huntsman spiders ) Stenochilidae Stiphidiidae Symphytognathidae ( dwarf orb - weavers ) Synaphridae Synotaxidae Tetragnathidae ( long jawed orb - weavers ) Theridiidae ( tangle - web spiders ) Theridiosomatidae ( ray spiders ) Thomisidae ( crab spiders ) Titanoecidae Trachelidae Trechaleidae Trochanteriidae Udubidae Uloboridae ( cribellate orb weavers ) Zodariidae ( ant spiders ) Zoropsidae ( zoropsid spiders ) List of families of spiders Spider taxonomy List of spider common names Bold are families with more than 1000 species Retrieved from `` https://en.wikipedia.org/w/index.php?title=Orb-weaver_spider&oldid=797219721 '' Categories : Araneidae Hidden categories : Articles using diversity taxobox Articles with ' species ' microformats All articles with unsourced statements Articles with unsourced statements from August 2007 Articles containing potentially dated statements from August 2017 All articles containing potentially dated statements Talk Contents About Wikipedia Wikispecies Asturianu Català Cebuano Čeština Dansk Deutsch Eesti Español Esperanto Euskara فارسی Français Gaeilge Bahasa Indonesia Íslenska Italiano עברית Lietuvių Magyar Nederlands 日本 語 Norsk Plattdüütsch Polski Português Română Runa Simi Русский Suomi Svenska Türkçe Українська Tiếng Việt Winaray 中文 Edit links This page was last edited on 25 August 2017, at 17 : 33. 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Question: where do orb weavers go during the day
Answer: they hide