Bootscan analysis on the genome sequences of PARV4-related viruses highlighted some genomic regions that may have undergone past recombination events between ovine parvovirus and PARV4 (Fig. S1 ). GARD analysis on the DataMonkey server (http://www.datamonkey.org/) also revealed the presence of 11 potential recombination breakpoints (data not shown), suggesting a multitude of possible past recombination events. However, incongruence testing with the Kishino-Hasegawa (KH) test failed to confirm the presence of phylogenetic incongruence in the partitions proposed by the bootscan and GARD analyses. Hence, it is more likely that the potential recombination signals were falsepositives resulting from local differences in evolutionary rates among different genes.