A scan of sequence divergence between complete genome sequences further supported the hypothesis of past recombi-nation between HBoV1 and 2 in the generation of HBoV3 and recombination of HBoV2 and HBoV3 in the generation of HBoV4, with both recombination points near the NP1 and VP1 junction (Figure 3 ). When different HBoV2 variants were similarly analyzed for recombination, intraspecies HBoV2 recombinants were also detected (data not shown) [7] .