Dates for the principal nodes are indicated in Table 4 . Within the cluster concerned with the recombination (Figure 5b tMRCAs estimated from ALN2-2 and E_161 are consistent, suggesting that the appropriate priors have successfully counterbalanced the influence of a lower substitution rate in the E-gene. Inference 2 placed tRE at 76 (HPD: 45-160) years before origin, which localizes the recombination within the Neudoerfl lineage and after the split with the Scharl lineage. The upper (older) bound for the tRE corresponds to the youngest of the parental divergence times in the tree Figure 3 . As the estimate for M is much older than time point m, the latter can be considered as the theoretical upper bound for the observed recombination event. The lower bound leaves open the possibility that recombination occurred after the LIV 369/T2 -LI/G divergence, whereas the upper bound sets it within a clade comprising LI/G, LIV 69/T2, LI/ 261, LI/K, LI/A, LI/NOR and LI/917. Based on previous phylogenetic dispersal reconstructions [39] , the first bound places the event in Scotland, whereas the second allows a wide range of locations within the UK, after the initial virus emergence in Ireland.