When disease transmission does not occur at the hotspot, it can still occur at a certain distance from the hotspot. This phenomenon has received little attention so far. Specifically, infective contacts may be observed when infectious individuals travel to the hotspot and cross the territory of susceptible individuals and, reversely, when susceptible individuals cross the territory of infectious individuals. This second type of transmission may be prominent when the disease reduces the mobility of sick individuals (i.e., sickness behavior [16, 17, 18] ). For example, in humans, sick individuals often stay home, which alters disease dynamics [19, 20] . Sick wild animals also commonly reduce their rate of search for food or water [21] . Such transmission may particularly apply to parasites that can survive in the environment (e.g., gastrointestinal parasites) for which the spatial overlap of the home ranges of sympatric hosts favors transmission [22] .