Interestingly, MAA/MAL-1 did not bind to the surface of goblet cells, suggesting that certain SAa2-3Gal glycans that show high affinity for MAA/MAL-I lectins, such as SAa2-3Galb1-4GlcNAcb and SAa2-3Galb1-4(6OSO3)GlcNAcb [4, 5] , are not present to a significant extent on the cell membranes of goblet cells. However, one-third of the goblet cells showed strong MAL-II staining, suggesting that the MAL-II specific glycans, such as SAa2-3Galb1-3(SAa2-6)GalNAca and SAa2-3Galb1-3(6OSO3)-GalNAca are highly expressed on goblet cells [4, 5, 6, 23] .