The recombination parameters are the recombination rates between two adjacent loci, e.g., r 1 for the recombination rate between markers a and b, and the interference coefficients, e.g., i 12 for interference between recombination events in the segments between markers a and b and b and c. To define these parameters we followed Christiansen [48] , and in particular the recombination distributions for two, three, and four loci (respectively, Tables 2.7, 2.8, and 2.9 of [48] ). It is important to realize that given the definitions of these parameters, the estimator of the recombination rate between two loci is not affected by the number of loci considered. In other words, we obtain the same estimation of the recombination rate between markers a and b whether we consider genotypic frequencies at just these two loci, or the frequencies at these two loci plus a third locus, or the complete information to which we have access, the fourmarker genotypes. Information on additional loci only affects the estimates of the interference coefficients.