Recombination analyses. Similarity values based on Jukes-Cantor corrected nucleotide distances between full-length sequences were calculated using the program SequenceDist in the Simmonic2005 v1.6 Sequence Editor Package [35] . To assess similarity across the genomes, sequence scans were performed using a fragment length of 300 bases and an increment of 9 bases between fragments. For sequence comparisons with HBoV1 and HBoV2, a mean pair-wise distance was computed using a set of 14 HBoV1 and 7 HBoV2A sequences.