Multiple tissue northern (MTN) blots (Clontech) were hybridized with the 32 P-labeled probes. The probe corresponding to 1378-1634 bp of H-Lse was used for hybridization. After stringent wash, the blot was placed on the storage phosphor screen (Packard, USA) and exposed for 3 hours in the dark. The signal was detected at the Cyclone storage phosphor system (Packard).