The filoviruses are members of the order Mononegavirales and contain a single-stranded negative-sense RNA genome, which is encapsidated by the nucleoprotein (NP). The MARV genome encodes seven structural proteins [9, 10] : the polymerase (L), VP35 and VP30 associate with NP to generate the helical nucleocapsid (NC) [11, 12, 13] . The viral glycoprotein (GP), which is inserted in the viral envelope, mediates cell entry [14, 15] . The major matrix protein VP40 plays a key role in virus assembly, and VP24, the second matrix protein, is suggested to support the template function of the NC [12, 16, 17, 18] .