Phylogenetic tree construction was performed with PAUP* version 4.0b10 [34] by using a maximum likelihood (ML) approach, under an optimum model of evolution as determined by MODELTEST v3.7 [35] . Due to the large dataset and to reduce the time required for computation, optimal trees were searched for by using a nearest neighbor interchange (NNI) heuristic search strategy. Bootstrapping was performed within PAUP* and displayed using exported PDF files created using Figtree v1.0 (previously available from: http://tree.bio.ed.ac.uk/ software/figtree/), which were subsequently annotated using Adobe Acrobat Professional 6.0.