Geminiviruses (family Geminiviridae) are single-stranded DNA (ssDNA) viruses that cause severe disease in major crop plants worldwide. Most geminiviruses belong to the genus Begomovirus, which are transmitted exclusively by the whitefly Bemisia tabaci (Harrison and Robinson, 1999) . Many begomoviruses have bipartite genomes known as DNA A and DNA B. DNA A contains the AV1 (coat protein) and AV2 ORFs (open reading frames) in the virus strand and, on the complementary strand, four ORFs: AC1 (replication initiation protein), AC2 (transcriptional activator protein), AC3 (replication enhancer protein) and AC4. The virus and complementary strands of DNA B contain two ORFs: BV1 (nuclear shuttle protein) and BC1 (movement protein). Some begomoviruses have a monopartite genome and lack DNA B. Phylogenetic analysis shows that begomoviruses can be generally divided into two groups, the Old World begomoviruses (eastern hemisphere, Asia, Africa, Europe, the Mediterranean areas) and the New World begomoviruses (western hemisphere, the Americas). All the New World begomoviruses are evolved to bipartite with lack of AV2 ORF in DNA A, whereas both bipartite and monopartite begomoviruses in the Old World encode AV2 ORF (Harrison and Robinson, 1999) .