In wild animal populations, high quality feeding spots (e.g., fruit trees), breeding sites, waterholes or sleeping sites can exacerbate direct physical contacts. Empirical and theoretical studies on the epidemiological importance of habitat hotspots have mainly focused on how the spatial aggregation of animals favors disease transmission at the hotspot itself [8, 9] . For example, the aggregation of wild boar at watering sites significantly increases the transmission of tuberculosis-like lesions [8] . However, interindividual contacts may not always significantly increase at the hotspot itself. This is for example the case of habitat hotspots that some animal species only visited occasionally, such as some mineral licks [10, 11] . Also, animals present at the same time at a particularly large hotspot may not be close enough to each other to transmit infectious diseases. This is the case of large forest clearings [12, 13] or large waterholes. Finally, species such as primates and ungulates might avoid defecating in hotspots of high food resources, limiting the transmission of fecal-oral parasites at hotspots [14, 15] .