The synthesis and decay of genomic nucleocapsids are described in the same way as for those of the anti-genomic nucleocapsids except that the polymerases on the anti-genomic templates are not released at intergenic regions: dpol r;p;j dt ¼ k e;pol ðencap pol trc l trc À n sec l À l trc À l lec pol r;p; j Þ j ¼ 1 ð19Þ