A comparison of contemporaneous cluster 1 and non-cluster animals was not possible (insufficient individuals occurred in the same time intervals) but cluster 2 individuals had significantly greater body condition (BW resid ) (LM, P = 0.036) and grouped macroparasite infection (PC1 p ) (LM, P = 0.048) than non-cluster individuals from the same sampling points ( Figure 6 ). For individual macroparasite species, there were significantly greater infections in cluster 2 animals for the louse, P. serrata (GLM, Poisson errors, P = 0.048) and the catenotaeniid cestode, S. lobata (GLM, negative binomial errors, P = 0.035).