Analysis of the genes surrounding the DpAV4 ORF-90-91-93 cluster confirms that this virus has an ascovirus origin since this region contains ORFs that are close homologs of genes in iridovirus and ascovirus genomes. Upstream from the ORF-90-91-93 cluster, an ORF encoding the DNA-dependent RNA polymerase 1 subunit C is present, which is an ortholog of the iridoviral CIV ORF176R and the ascoviral SfAV1a ORF008. Downstream from this cluster, there are two genes, absent in known ascoviral genomes, but similar to the iridoviral CIV ORF115L and CIV ORF132L. These two genes encode, respectively, a chromosomal replication initiation protein and zinc finger protein. In between them, a gene encoding a small protein is present that is similar to that encoded by the ORF069L of the iridovirus CIV, and which corresponds to the ALI-like protein also found in entomopoxviruses [30] .