MOUV formed a uniquely divergent lineage, distant from all other hantaviruses identified to date, except for NVAV (Figure 2 ), in phylogenetic trees based on L-segment sequences, generated by the maximum-likelihood and Bayesian methods, implemented in PAUP* (Phylogenetic Analysis Using Parsimony, 4.0b10) [23] , RAxML Blackbox webserver [24] and MrBayes 3.1 [25] , under the best-fit GTR+I+Γ model of evolution established using jModeltest 0.1.1 [26] . Topologies were well supported by bootstrap analysis of 100 iterations, and posterior node probabilities based on two runs each of 2 million generations sampled every 100 generations with burn-in of 25%.