查尔斯·克拉克 (植物学家)

查尔斯·克拉克（Charles M. Clarke）是专业从事于猪笼草属食虫植物的植物学家与分类学家。克拉克出生于澳大利亚墨尔本。他获得了墨尔本莫纳什大学（Monash University）的生物学荣誉学位，同时还在新南威尔士阿米代尔的新英格兰大学（University of New England）取得了生态系统管理博士学位。
克拉克第一次到婆罗洲研究猪笼草是在1987年。1989年至1990年，他生活在文莱以研究猪笼草属植物的生态问题。在这两次旅行间，克拉克还曾在昆士兰州的詹姆斯·库克大学（James Cook University）任教生态学和生物统计学，同时还在香港担任园艺顾问。
克拉克现就任于莫纳什大学马来西亚分校。克拉克已完成了五本关于猪笼草属的著作，其中最重要的是《婆罗洲的猪笼草属植物》（Nepenthes of Borneo）和《苏门答腊岛与西马来西亚的猪笼草》（Nepenthes of Sumatra and Peninsular Malaysia）。
克拉克描述了5个猪笼草属物种，分别为本斯通猪笼草（Nepenthes benstonei）、陈氏猪笼草（Nepenthes chaniana）、泉氏猪笼草（Nepenthes izumiae）、贾桂琳猪笼草（Nepenthes jacquelineae）和坚韧猪笼草（Nepenthes tenax）。

**著作**
 Clarke, C.M. 1992. The ecology of metazoan communities in Nepenthes pitcher plants in Borneo, with special reference to the community of Nepenthes bicalcarata Hook.f. Ph.D. thesis, University of New England, Armidale, N.S.W. 269 pp.- Clarke, C.M. 1993. PDFlink| The Possible Functions of the Thorns of Nepenthes bicalcarata (Hook.f.) Pitchers.|194 KiB Carnivorous Plant Newsletter 22(1–2): 27–28.- Clarke, C.M. & R.L. Kitching 1993. The Metazoan Food Webs from Six Bornean Nepenthes Species. Ecological Entomology 18: 7–16.- Clarke, C.M. & J.A. Moran 1994. A further record of aerial pitchers in Nepenthes ampullaria Jack. Malayan Nature Journal 47: 321–323.- Clarke, C.M. & R.L. Kitching 1995. Swimming Ants and Pitcher Plants: a Unique Ant-Plant Interaction from Borneo. Journal of Tropical Ecology 11(4): 589–602.- Clarke, C.M. 1997. Nepenthes of Borneo. Natural History Publications (Borneo), Kota Kinabalu. xi + 207 pp.- Clarke, C.M. 1997.  Another nice trip to Sumatra date=20150923200719 . Carnivorous Plant Newsletter 26(1): 4–10.- Clarke, C.M. 1997. The effects of pitcher dimorphism on the metazoan community of the carnivorous pitcher plant Nepenthes bicalcarata Hook.f. Malayan Nature Journal 50: 149–157.- Clarke, C.M. 1998. Initial colonisation and prey capture in Nepenthes bicalcarata (Nepenthaceae) pitchers in Brunei. Sandakania 12: 27–36.- Clarke, C.M. 1998. The aquatic arthropod community of the pitcher plant, Nepenthes bicalcarata (Nepenthaceae) in Brunei. Sandakania 11: 55–60.- Clarke, C.M. 1998. A re-examination of geographical variation in Nepenthes food webs. Ecography 21: 430–436.- Clarke, C.M. 1999. Nepenthes benstonei (Nepenthaceae), a new pitcher plant from Peninsular Malaysia. Sandakania 13: 79–87.- Clarke, C.M. 2001. Ecology & Conservation of Montane Nepenthes (Nepenthaceae) in Sumatra. Fourth International Flora Malesiana Symposium. 20-24 July 1998, Kuala Lumpur.- Clarke, C.M. 2001. Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu. ix + 325 pp.- Clarke, C.M. 2001. A Guide to the Pitcher Plants of Sabah. Natural History Publications (Borneo), Kota Kinabalu. iv + 40 pp.- Moran, J.A., M.A. Merbach, N.J. Livingston, C.M. Clarke & W.E. Booth 2001.  Termite Prey Specialization in the Pitcher Plant Nepenthes albomarginata - Evidence from Stable Isotope Analysis. Annals of Botany 88: 307–311.- Clarke, C.M. 2002. A Guide to the Pitcher Plants of Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu. iv + 32 pp.
*Clarke, C.M., T. Davis & R. Tamin 2003. Nepenthes izumiae (Nepenthaceae): a new species from Sumatra. Blumea 48: 179–182.
*Moran, J.A., C.M. Clarke & B.J. Hawkins 2003. From Carnivore to Detritivore? Isotopic Evidence for Leaf Litter Utilization by the Tropical Pitcher Plant Nepenthes ampullaria. International Journal of Plant Sciences 164: 635–639.
*Dong, T.T.X., Q.M. Xiao, C.M. Clarke, H.S. Zhong, N.J. Zhao, K.L. Chun & K.W.K. Tsim 2003. Phylogeny of Astragalus in China: Molecular evidence from the DNA sequences of 5S rRNA spacer, ITS, and 18S rRNA. Journal of Agricultural and Food Chemistry 51(23): 6709–6714.
*Clarke, C.M. & C.C. Lee 2004. A Pocket Guide: Pitcher Plants of Sarawak. Natural History Publications (Borneo), Kota Kinabalu. vi + 81 pp.
*Cantley, R., C.M. Clarke, J. Cokendolpher, B. Rice & A. Wistuba 2004.  Nepenthes clipeata Survival Project. International Carnivorous Plant Society.
*Clarke, C.M., C.C. Lee & S. McPherson 2006. Nepenthes chaniana (Nepenthaceae), a new species from north-western Borneo. Sabah Parks Journal 7: 53–66.
*Clarke, C.M & R. Kruger 2006. Nepenthes tenax C.Clarke and R.Kruger (Nepenthaceae), a new species from Cape York Peninsula, Queensland. Austrobaileya 7(2): 319–324.
*Clarke, C.M. 2006. Introduction. In: Danser, B.H. The Nepenthaceae of the Netherlands Indies. Natural History Publications (Borneo), Kota Kinabalu. pp. 1–15
*Clarke, C.M., U. Bauer, C.C. Lee, A.A. Tuen, K. Rembold & J.A. Moran 2009.  Tree shrew lavatories: a novel nitrogen sequestration strategy in a tropical pitcher plant. Biology Letters 5(5): 632–635. 10.1098/rsbl.2009.0311
*Chin, L., J.A.  Moran & C. Clarke 2010. Trap geometry in three giant montane pitcher plant species from Borneo is a function of tree shrew body size. New Phytologist, published online January 22, 2010. 10.1111/j.1469-8137.2009.03166.x

**参考文献**

**扩展阅读**
 Clarke, C.M. 1997. Nepenthes of Borneo.  Natural History Publications (Borneo), Kota Kinabalu. xi + 207 pp. ISBN 983-812-057-X