From the passage identify the symbols of Demeter. Display the results in a comma separated format.

Context: Demeter Goddess of grain, agriculture, harvest, growth, and nourishment. Demeter, whose Roman counterpart is Ceres, is a daughter of Cronus and Rhea, and was swallowed and then regurgitated by her father. She is a sister of Zeus, by whom she bore Persephone, who is also known as Kore, i.e. "the girl." One of the central myths associated with Demeter involves Hades' abduction of Persephone and Demeter's lengthy search for her. Demeter is one of the main deities of the Eleusinian Mysteries, in which the rites seemed to center around Demeter's search for and reunion with her daughter, which symbolized both the rebirth of crops in spring and the rebirth of the initiates after death. She is depicted as a mature woman, often crowned and holding sheafs of wheat and a torch. Her symbols are the cornucopia, wheat-ears, the winged serpent, and the lotus staff. Her sacred animals include pigs and snakes.

Answer: cornucopia, wheat-ears, the winged serpent, the lotus staff