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| **BIOGRAPHICAL SKETCH**         |   |       |         |                |
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| Provide the following           |   |       |         |                |
| information for the key         |   |       |         |                |
| personnel in the order listed   |   |       |         |                |
| for Form Page 2. Photocopy this |   |       |         |                |
| page or follow this format for  |   |       |         |                |
| each person.                    |   |       |         |                |
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| NAME                            |   | POS   |         |                |
|                                 |   | ITION |         |                |
| **R. Scott Hawley**             |   | TITLE |         |                |
|                                 |   |       |         |                |
|                                 |   | *     |         |                |
|                                 |   | *Inve |         |                |
|                                 |   | stiga |         |                |
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| EDUCATION/TRAINING              |   |       |         |                |
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| INSTITUTION AND LOCATION        | D |       | YEAR(s) | FIELD OF STUDY |
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|                                 | R |       |         |                |
|                                 | E |       |         |                |
|                                 | E |       |         |                |
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| University of California at     | B |       | 1975    | Biology        |
| Riverside                       | . |       |         |                |
|                                 | S |       |         |                |
|                                 | . |       |         |                |
+---------------------------------+---+-------+---------+----------------+
| University of Washington,       | P |       | 1979    | Genetics       |
| Seattle                         | h |       |         |                |
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|                                 | D |       |         |                |
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**A. Positions and Honors.**

[Professional Experience]{.underline}

1975-1979 Graduate Student, University of Washington, Seattle. Sponsor:
Dr. L.M. Sandler

1979-1982 Helen Hay Whitney Post-doctoral Fellow, Inst. for Cancer
Research, Philadelphia, PA

1982-1991 Assist./Assoc. Prof., Dept. of Genetics and Mol. Biol., Albert
Einstein Coll. of Med, NY

2001. Professor, Section of Molecular and Cellular Biology, University
      of California at Davis

2001-present Investigator, Stowers Institute of Medical Research, Kansas
City, MO

2006-present Professor of Molecular Biosciences with Tenure, University
of Kansas

2006-present Professor of Physiology with Tenure, University of Kansas
Medical Center

2006-present Adjunct Professor of Biology, University of Missouri at
Kansas City

[Honors and Awards]{.underline}

Helen Hay Whitney Postdoctoral Fellowship (1979-1982)

Searle Scholarship (1984-1987)

American Cancer Society Faculty Research Awards (1984-1993)

Chair, Gordon Research Conference on Meiosis (1994)

Editorial Board of *Genetics, PLoS, and PloS Genetics* (current)

Board of Directors of the Genetics Society of America (1996-1999)

Fellow of the AAAS (elected 2001)

Co-chair, Cold Spring Harbor Advanced *Drosophila* Genetics Course
(1994-2004)

American Cancer Society Research Professor (2005-2010)

Member of the American Academy of Arts and Sciences (2006)

Appointed to the Biological Sciences Advisory Committee of the National
Science Foundation (2007-2009)

**B. Selected peer-reviewed publications (94 Total)**

Hawley, R. S. 1980. Chromosomal sites necessary for normal levels of
meiotic recombination in Drosophila melanogaster. 1. Evidence for and
mapping of the sites. Genetics 94:625-646.

Zhang, P., B. Knowles, L. S. B. Goldstein, and R. S. Hawley. 1990. A
kinesin-like protein required for distributive chromosome segregation in
Drosophila. Cell 62:1053-1062.

Theurkauf, W. E. and R. S. Hawley. 1992. Meiotic spindle assembly in
*Drosophila* females: behavior of nonexchange chromosomes and the
effects of mutations in the *nod* kinesin-like protein. [J. Cell
Biol]{.underline}. 116: 1167-1180.

Hawley, R.S. H. Irick, A.E. Zitron, D.A. Haddox, A. Lohe, C. New, M.
Whitley, T. Arbel, J. Jang, K. McKim, and G.C. Childs. 1993. There are
two mechanisms of achiasmate segregation in *Drosophila* females, one of
which requires heterochromatic homology. [Developmental
Genetics]{.underline} 13: 440-467.

McKim, K., J.K. Jang, W. Theurkauf, and R.S. Hawley. 1993. Mechanical
basis of meiotic metaphase arrest. [Nature]{.underline} 362: 364-366.

Afshar, K., N. Barton, R.S. Hawley and L.S.B. Goldstein 1994. DNA
binding and meiotic chromosomal localization of the *Drosophila* NOD
kinesin-like protein [Cell]{.underline} 81:129-138.

Jang, J. K., L. Messina, T.A. Arbel, and R.S. Hawley. 1995. Induction of
metaphase arrest in *Drosophila* oocytes by chiasma-based kinetochore
tension. [Science]{.underline} 268: 1917-1919.

McKim, K.S. and R.S. Hawley 1995. Chromosomal Control of Meiotic Cell
Division. [Science]{.underline} 270:1595-1601.

Dernburg, A.F., Sedat, J.W. and R.S. Hawley. 1996. Direct evidence of a
role for heterochromatin in meiotic chromosome segregation.
[Cell]{.underline} 86:135-146.

Koehler, K.E., C.L. Boulton, H.E. Collins, R.L. French, K.C. Herman,
L.D. Lacefield, S.M. Madden, C.D. Scheutz, and R.S. Hawley. 1996.
Spontaneous *X* chromosome nondisjunction events occurring at MI and MII
have different recombinational histories. [Nature Genetics]{.underline}
14:406-414.

McKim, K.S., J.J. Sekelsky, G. Chin, C. Steinberg, R. Khodosh, and R.S.
Hawley. 1998. Meiotic synapsis in the absence of recombination.
[Science]{.underline} 279:876-878.

Matthies, H.J.G., Baskin, R.J., and R.S. Hawley. 2001. The orphan
kinesin NOD lacks motile properties but does possess a microtubule
stimulated ATPase activity. [Molecular Biology of the Cell]{.underline}
12(12): 4000-4012.

Page, S.L. and R.S. Hawley. 2001. *c(3)G* encodes a *Drosophila*
synaptonemal complex protein. [Genes &]{.underline}
[Development]{.underline} 15:3130-3143.

Kramer, J.J. and R.S. Hawley. 2003. Mutants in the spindle-associated
transmembrane protein Axs impair meiotic spindle assembly in
*Drosophila* oocytes. [Nature Cell Biology]{.underline} 5:261-263.

Harris, D., Orme, C., Kramer, J.J., Namba, R. Champion, M.D., Palladino,
M.J., Natzle, J.E., and., and R.S.

Hawley. 2003. A deficiency screen of the major autosomes identifies a
single gene(*matrimony*) that is haplo-insufficient for achiasmate
segregation in *Drosophila* oocytes. [Genetics]{.underline}165:
637-652*.*

Page, S.L. and R.S. Hawley 2004. The genetics and molecular biology of
the synaptonemal complex. [Ann. Rev. Cell Dev. Biology]{.underline},
*Vol. 20: 525-558*

Page, S.L. and R.S. Hawley. 2003. Chromosome Choreography - the Meiotic
Ballet. [Science]{.underline} 301(5634):785-789)

Gilliland, W.D., Wayson, S.M., and R.S. Hawley 2005. The meiotic defects
of mutants in the *Drosophila* *mps1* gene reveal a critical role of
Mps1 in the segregation of achiasmate homologs. [Current
Biology]{.underline} 15(7): 672-677.

Anderson, L. K., Royer, S., Page, S.L., McKim, K., Lai, A., Lilly, M.Y,
and R. S. Hawley. 2005. Juxtaposition of *C(2)M* and the Transverse
Filament Protein *C(3)G* within the Central Region of *Drosophila*
Synaptonemal Complex. [PNAS]{.underline} 102(12): 4482-4487.

Page, S. L. and R. S. Hawley. 2005. The *Drosophila* meiotic mutant
*mei-352* is an allele of *klp3A* and reveals a role for a kinesin-like
protein in crossover distribution. [Genetics]{.underline} 170:
1797-1807*.*

Cui, W., Sproul, L.R., Gustafson, S.M., Matthies, H.J., Gilbert, S.P.,
and R.S. Hawley 2005. *Drosophila* nod protein binds preferentially to
the ends of microtubules and promotes microtubule polymerization *in
vitro.* [Molecular Biology of the Cell]{.underline} 16(11):5400-5409*.*

Cui, W. and R.S. Hawley 2005. The HhH(2)/NDD domain of the Drosophila
Nod chromokinesins-like protein is required for binding to chromosomes
in the oocyte nucleus.. [Genetics]{.underline} 171(4):1823-1835

Gong, W., McKim, K.S., and R.S. Hawley 2005. All paired up with no place
to go: pairing, synapsis, and DSB formation in a balancer heterozygote.
[PLOS-Genetics 1(5):589-602]{.underline}.

Gilliland, W.D. and R.S. Hawley 2005. Cohesin and the maternal age
effect. [Cell]{.underline} 123(3):371-3.

Xiang, Y. and R.S. Hawley 2006. Centromere co-orientation and the
mechanism of secondary nondisjunction in *Drosophila* females. 
[Genetics]{.underline} 174: 67-68.

Hawley, R.S. and W.D. Gilliland 2006 Sometimes the result is not the
answer: the truths and the lies that come from using the complementation
test. [Genetics]{.underline} 174: 5-15.

**[BOOKS]{.underline}**

*The Human Genome: A User's Guide*. 1998. by R. Scott Hawley and
Catherine Mori. Academic Press.

*Drosophila Protocols*. 1999. by William S. Sullivan, Michael Ashburner,
and R. Scott Hawley, eds. Cold Spring Harbor Press, New York.

*Advanced Genetic Analysis: Finding Meaning in the Genome*. 2002. by R.
Scott Hawley and Michelle Walker.

Blackwell Science.

*The Human Genome: A User's Guide*. *2nd Edition*. 2004. by Julia
Richards and R. Scott Hawley. Academic Press.

*Drosophila: A Laboratory Handbook, 2^nd^ Edition*. 2004 by Michael
Ashburner, R. Scott Hawley, and Kent Golic. Cold Spring Harbor Press.

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