**Table 2**. *Enzymological patterning of L-phenylalanine biosynthesis
in pseudomonad species representing*

*Groups I through V*

+------+---+-----+----------+------+----------+----+---------+---------+
| **rR |   | **  | **S      | **D  | *        | ** |         | **Tyr   |
| NA** |   | ATC | pecies** | NA** | *Phenyla | Sp |         | osine** |
|      |   | C** |          |      | lanine** | ec |         |         |
| **ho |   |     |          | **ho |          | if |         | **activ |
| molo |   | *   |          | molo | **patt   | ic |         | ation** |
| gy** |   | *No |          | gy** | erning** | A  |         |         |
|      |   | .** |          |      |          | ct |         | **fact  |
| **gr |   |     |          | **gr | **gr     | iv |         | or^e^** |
| oup^ |   |     |          | oup^ | oup^c^** | it |         |         |
| a^** |   |     |          | b^** |          | y^ |         |         |
|      |   |     |          |      |          | d^ |         |         |
|      |   |     |          |      |          | ** |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          | *  | >       |         |
|      |   |     |          |      |          | *A | **PPA** |         |
|      |   |     |          |      |          | GN |         |         |
|      |   |     |          |      |          | ** |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| I \* | 1 |     | *P.      | 1    | Ia       | 0  | 27.1    | 0.7     |
|      | 7 |     | s        |      |          |    |         |         |
|      | 5 |     | tutzeri* |      |          |    |         |         |
|      | 8 |     |          |      |          |    |         |         |
|      | 8 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 35.5    | 0.7     |
|      | 7 |     | s        |      |          |    |         |         |
|      | 5 |     | tutzeri* |      |          |    |         |         |
|      | 9 |     |          |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 6.6     | 1.1     |
|      | 4 |     | alca     |      |          |    |         |         |
|      | 9 |     | ligenes* |      |          |    |         |         |
|      | 0 |     |          |      |          |    |         |         |
|      | 9 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| \*   | 2 |     | *P.      |      |          | 0  | 13.1    | 1.0     |
|      | 5 |     | me       |      |          |    |         |         |
|      | 4 |     | ndocina* |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     | *P.      |      | Ib       | 0  | 9.8     | 0.8     |
|      |   |     | aerugi   |      |          | .3 |         |         |
|      |   |     | nosa*^f^ |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| \*   | 1 |     | *P.      |      |          | 1  | 11.9    | 1.3     |
|      | 2 |     | putida*  |      |          | .2 |         |         |
|      | 6 |     |          |      |          |    |         |         |
|      | 3 |     |          |      |          |    |         |         |
|      | 3 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| \*   | 1 |     | *P.      |      |          | 0  | 16.3    | 1.1     |
|      | 3 |     | fluo     |      |          | .3 |         |         |
|      | 5 |     | rescens* |      |          |    |         |         |
|      | 2 |     |          |      |          |    |         |         |
|      | 5 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 2.6     | 1.0     |
|      | 0 |     | s        |      |          | .1 |         |         |
|      | 8 |     | yringae* |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
|      | 2 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 5.1     | 1.0     |
|      | 9 |     | mori*    |      |          | .8 |         |         |
|      | 8 |     |          |      |          |    |         |         |
|      | 7 |     |          |      |          |    |         |         |
|      | 2 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 12.1    | 1.0     |
|      | 0 |     | aptata*  |      |          | .2 |         |         |
|      | 2 |     |          |      |          |    |         |         |
|      | 0 |     |          |      |          |    |         |         |
|      | 5 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 1  | 16.0    | 1.1     |
|      | 0 |     | c        |      |          | .5 |         |         |
|      | 8 |     | ichorii* |      |          |    |         |         |
|      | 5 |     |          |      |          |    |         |         |
|      | 7 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| II   | 1 |     | *P.      | 2a   | IIa      | 0  | 7.1     | 3.0     |
|      | 0 |     | cepacia* |      |          | .2 |         |         |
|      | 8 |     |          |      |          |    |         |         |
|      | 5 |     |          |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 1  | 10.8    | 1.7     |
|      | 7 |     | cepacia* |      |          | .4 |         |         |
|      | 4 |     |          |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
|      | 0 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| \*   | 1 |     | *P.      |      |          | 0  | 3.2     | 2.1     |
|      | 0 |     | ma       |      |          | .2 |         |         |
|      | 2 |     | rginata* |      |          |    |         |         |
|      | 4 |     |          |      |          |    |         |         |
|      | 8 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 4.3     | 3.8     |
|      | 0 |     | ma       |      |          | .2 |         |         |
|      | 8 |     | rginata* |      |          |    |         |         |
|      | 5 |     |          |      |          |    |         |         |
|      | 4 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 5.9     | 2.5     |
|      | 9 |     | ma       |      |          | .2 |         |         |
|      | 3 |     | rginata* |      |          |    |         |         |
|      | 0 |     |          |      |          |    |         |         |
|      | 2 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 2 |     | *P.      |      |          | 0  | 3.1     | 2.0     |
|      | 5 |     | cary     |      |          | .1 |         |         |
|      | 4 |     | ophylli* |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
|      | 8 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| \*   | 2 |     | *P.      | 2b   | IIb      | 0  | 9.5     | 1.0     |
|      | 7 |     | pi       |      |          | .1 |         |         |
|      | 5 |     | ckettii* |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 7  | 15.6    | 1.1     |
|      | 0 |     | solan    |      |          | .1 |         |         |
|      | 6 |     | acearum* |      |          |    |         |         |
|      | 9 |     |          |      |          |    |         |         |
|      | 2 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| III  | 1 |     | *P.      | 3a   | IIIa     | 0  | 2.2     | 6.3     |
|      | 7 |     | acid     |      |          |    |         |         |
|      | 4 |     | ovorans* |      |          |    |         |         |
|      | 7 |     |          |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 5.9     | 11.5    |
|      | 7 |     | testo    |      |          |    |         |         |
|      | 5 |     | steroni* |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
|      | 0 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| \*   | 1 |     | *P.      | 3b   | IIIb     | 0  | 1.9     | 3.4     |
|      | 7 |     | dela     |      |          | .9 |         |         |
|      | 5 |     | fieldii* |      |          |    |         |         |
|      | 0 |     |          |      |          |    |         |         |
|      | 5 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 1.6     | 4.3     |
|      | 1 |     | facilis* |      |          | .2 |         |         |
|      | 2 |     |          |      |          |    |         |         |
|      | 2 |     |          |      |          |    |         |         |
|      | 8 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| \*   | 1 |     | *P.      | 3c   | IIIc     | 0  | 0.9     | 19.7    |
|      | 5 |     | saccha   |      |          | .9 |         |         |
|      | 9 |     | rophila* |      |          |    |         |         |
|      | 4 |     |          |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| IV   | 1 |     | *P.      | 4    | IV       | 0  | 0       | 1.0     |
| \*   | 1 |     | d        |      |          | .5 |         |         |
|      | 5 |     | iminuta* |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
|      | 8 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 0       |         |
|      | 3 |     | d        |      |          | .9 |         |         |
|      | 1 |     | iminuta* |      |          |    |         |         |
|      | 8 |     |          |      |          |    |         |         |
|      | 4 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 0       |         |
|      | 1 |     | vesi     |      |          | .6 |         |         |
|      | 4 |     | cularis* |      |          |    |         |         |
|      | 2 |     |          |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
| V \* | 1 |     | *P.      | 5    | V        | 0  | 0.4     | 1.0     |
|      | 3 |     | malt     |      |          | .7 |         |         |
|      | 6 |     | ophilia* |      |          |    |         |         |
|      | 3 |     |          |      |          |    |         |         |
|      | 7 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 1 |     | *P.      |      |          | 0  | 0.8     | 1.3     |
|      | 7 |     | malt     |      |          | .3 |         |         |
|      | 8 |     | ophilia* |      |          |    |         |         |
|      | 0 |     |          |      |          |    |         |         |
|      | 6 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      | 6 |     | *X.      |      |          | 0  | 5.9     | 1.0     |
|      | 7 |     | cam      |      |          | .1 |         |         |
|      | 2 |     | pestris* |      |          |    |         |         |
|      | 1 |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+
|      |   |     |          |      |          |    |         |         |
+------+---+-----+----------+------+----------+----+---------+---------+

**Updated taxonomic designations:** Organisms listed in **DNA homology
group 1** are members of the contemporary *Pseudomonas* genus. Members
of **group 2a** are designated as species of *Burkholderia*. Members of
**group 2b** are designated as species of *Ralstonia*. Members of
**group 3a** are named as species of *Comomonas*. Members of **group
3b** are named as species of *Acidivorax*. *P. saccharophila* in **group
3c** is designated *Pelomonas saccharophila*. Members of **group 4** are
designated as species of *Brevundimonas*. P. maltophilia in **group V**
is designated *Stenotrophomonas maltophilia.*

The rRNA homology groups I and V populate the Gamma division of
Proteobacteria (also known previously as Superfamily B). rRNA homology
groups II and III populate the Beta division of Proteobacteria (also
previously known as Superfamily A). rRNA homology group IV populates the
Alpha division of Proteobacteria (also known previously as Superfamily
C).

**Key taxonomy reference:** Anzai Y, H Kim, J Park, H Wakabayashi, and H
Oyaizu. 2000. Phylogenetic affiliation of the pseudomonads based on 16S
rRNA sequence. Int J Sys Evol Microbiol 50:1563-1589.