Number range CHAPTER**JEFFREY J. HARD**

National Marine Fisheries Service

Northwest Fisheries Science Center

Conservation Biology Division

2725 Montlake Boulevard East

Seattle, Washington 98112

Tel: (206) 860-3275

Fax: (206) 860-3335

E-mail: [[jeff.hard@noaa.gov]{.underline}](mailto:jeff.hard@noaa.gov)

Website : [http://www.nwfsc.noaa.gov](http://www.nwfsc.noaa.gov/)

**RESEARCH INTERESTS**

Evolution of life histories

Quantitative genetics

Conservation biology

Genetic and ecological consequences of harvest and propagation

Experimental design and statistical analysis

**EDUCATION**

1991 Ph.D. Ecology and Evolutionary Biology, University of Oregon,
Eugene

1984 M.S. Fisheries, University of Alaska, Juneau

1979 B.S. Biology, Oregon State University, Corvallis (cum. GPA, 3.73;
Chemistry minor)

**EXPERIENCE**

1997- Population Biology Program Manager, Conservation Biology Division,
Northwest Fisheries Science Center, Seattle, Washington

2002 Supervisory Research Fishery Biologist and Acting Director,
Conservation Biology Division, Northwest Fisheries Science Center,
Seattle, Washington

1992- Research Fishery Biologist, U.S. Department of Commerce, NMFS,
Northwest Fisheries Science Center, Seattle, Washington

1986-91 Graduate Research Fellow, Department of Biology, University of
Oregon, Eugene

1982-86 Fishery Biologist, U.S. Department of Commerce, NMFS, Auke Bay
Laboratory and Little Port Walter Field Station, Alaska

1977-82 Fishery Technician, U.S. Department of Commerce, NMFS, Auke Bay
Laboratory and Little Port Walter Field Station, Alaska

1975-76 Research Assistant, Institute of Marine Science, University of
Alaska, Fairbanks, Alaska, and CEPEX (Controlled Ecosystem Pollution
EXperiment) Project, Patricia Bay Laboratory, Sidney, B.C., Canada

**PUBLICATIONS**

**Refereed Research Papers**

1.  Hard, J. J., D. G. Elliott, R. G. Pascho, D. M. Chase, L. K.
    Park, J. R. Winton, and D. E. Campton. Genetic variation in disease
    resistance of chinook salmon (*Oncorhynchus tshawytscha*): are
    phenotypic indicators of *Renibacterium* infection correlated with
    disease resistance of their progeny? [Canadian Journal of Fisheries
    and Aquatic Sciences]{.underline}. *Accepted, in revision*.

2.  Ford, M. J., H. Fuss, B. Boelts, E. LaHood, J. J. Hard, and J. M.
    Miller. Changes in run timing and natural smolt production in a
    naturally spawning coho salmon (*Oncorhynchus kisutch*) population
    after 60 years of intensive hatchery supplementation. [Canadian
    Journal of Fisheries and Aquatic Sciences]{.underline}. *Accepted,
    in revision.*

3.  Hard, J. J., L. S. Mills, and J. M. Peek. *In press*. Genetic
    implications of reduced survival of male red deer under harvest.
    [Wildlife Biology]{.underline}.

4.  McClelland, E. K., J. M. Myers, J. J. Hard, L. K. Park, and K. A.
    Naish. 2005. Two generations of outbreeding in coho salmon
    (*Oncorhynchus kisutch*): effects on size and growth. [Canadian
    Journal of Fisheries and Aquatic Sciences]{.underline}
    62:2538-2547*.*

5.  Smoker, W. W., I. A. Wang, A. J. Gharrett, and J. J. Hard. 2004.
    Embryo survival and smolt to adult survival in second-generation
    outbred coho salmon. [Journal of Fish Biology]{.underline} 65
    (Suppl. A):254-262.

6.  Thrower, F. P., J. J. Hard, and J. E. Joyce. 2004. Genetic
    architecture of growth and early life history transitions in
    anadromous and derived freshwater populations of steelhead
    (*Oncorhynchus mykiss*). [Journal of Fish Biology]{.underline} 65
    (Suppl. A):286-307.

7.  Granath, K. L., W. W. Smoker, A. J. Gharrett, and J. J. Hard. 2004.
    Effects on embryo development time and survival of intercrossing
    three geographically separate populations of southeast Alaska coho
    salmon (Oncorhynchus kisutch). [Environmental Biology of
    Fishes]{.underline} 69:299-306.

8.  Waples, R. S., R. G. Gustafson, L. A. Weitkamp, J. M. Myers, O. W.
    Johnson, P. J. Busby, J. J. Hard, G. J. Bryant, F. W. Waknitz, K.
    Neely, D. Teel, W. S. Grant, G. A. Winans, S. Phelps, A. Marshall,
    and B. M. Baker. 2001. Characterizing diversity in Pacific salmon
    from the Pacific Northwest. [Journal of Fish Biology]{.underline}
    59(a):1-41.

9.  Hard, J. J., L. Connell, W. K. Hershberger, and L. W. Harrell. 2000.
    Genetic variation in mortality of chinook salmon (*Oncorhynchus
    tshawytscha*) during a bloom of the marine alga *Heterosigma
    akashiwo.* [Journal of Fish Biology]{.underline} 56:1387-1397.

10. Hard, J. J., B. A. Berejikian, E. P. Tezak, S. L. Schroder, C. M.
    Knudsen, and L. T. Parker. 2000. Evidence for morphometric
    differentiation of wild and captively reared coho salmon: a
    geometric analysis. [Environmental Biology of Fishes]{.underline}
    58(1):61-73.

11. Hard, J. J., G. A. Winans, and J. C. Richardson. 1999. Phenotypic
    and genetic architecture of juvenile morphometry in chinook salmon.
    [Journal of Heredity]{.underline} 90(6):597-606.

12. Hard, J. J., and W. R. Heard. 1999. Analysis of straying variation
    in Alaskan hatchery chinook salmon following transplantation.
    [Canadian Journal of Fisheries and Aquatic Sciences]{.underline}
    56:578-589.

13. Berejikian, B. A., E. P. Tezak, S. L. Schroder, C. M. Knudsen,
    and J. J. Hard. 1997. Reproductive behavioral interactions between
    wild and captively reared coho salmon (*Oncorhynchus kisutch*).
    [ICES Journal of Marine Science]{.underline} 54:1040-1050.

14. Bradshaw, W. E., C. M. Holzapfel, C. A. Kleckner, and J. J.
    Hard. 1997. Heritability of development time and protandry in the
    pitcher-plant mosquito, *Wyeomyia smithii*. [Ecology]{.underline}
    78:969-976.

15. Hard, J. J. 1995. Science, education, and the fisheries scientist.
    [Fisheries]{.underline} 20(3):10-16.

16. Hard, J. J., W. E. Bradshaw, and C. M. Holzapfel. 1993. Genetic
    coordination of demography and phenology in the pitcher-plant
    mosquito, *Wyeomyia smithii*. [Journal of Evolutionary
    Biology]{.underline} 6:707-723.

17. Hard, J. J., W. E. Bradshaw, and C. M. Holzapfel. 1993. The genetic
    basis of photoperiodism and its divergence among populations of the
    pitcher-plant mosquito, *Wyeomyia smithii*. [The American
    Naturalist]{.underline} 142(3):457-473.

18. Hard, J. J., and W. E. Bradshaw. 1993. Reproductive allocation in
    the western tree-hole mosquito, *Aedes sierrensis*.
    [Oikos]{.underline} 66(1):55-65.

19. Hard, J. J., W. E. Bradshaw, and C. M. Holzapfel. 1992. Epistasis
    and the divergence of photoperiodism between populations of the
    pitcher-plant mosquito, *Wyeomyia* *smithii*. [Genetics]{.underline}
    131:389-396.

20. Courtney, S. P., and J. J. Hard. 1990. Host acceptance and
    life-history traits in *Drosophila busckii*: tests of the
    hierarchy-threshold model*.* [Heredity]{.underline}
    64(1990):371-375.

21. Hard, J. J., W. E. Bradshaw, and D. J. Malarkey. 1989. Resource- and
    density-dependent development in tree-hole mosquitoes.
    [Oikos]{.underline} 54(2):137-144.

22. Hard, J. J., A. C. Wertheimer, and W. F. Johnson. 1989. Geographic
    variation in the occurrence of red- and white-fleshed chinook salmon
    (*Oncorhynchus tshawytscha*) in western North America. [Canadian
    Journal of Fisheries and Aquatic Sciences]{.underline}
    46(7):1107-1113.

23. Hard, J. J. 1986. Production and yield of juvenile chinook salmon in
    two Alaskan lakes. [Transactions of the American Fisheries
    Society]{.underline} 115(2):305-313.

24. Hard, J. J., A. C. Wertheimer, W. R. Heard, and R. M. Martin. 1985.
    Early male maturity in two stocks of chinook salmon (*Oncorhynchus
    tshawytscha*) transplanted to an experimental hatchery in
    southeastern Alaska. [Aquaculture]{.underline} 48(1985):351-359.

**Refereed Book Chapters and Review Papers**

1.  Kapuscinski, A., J. Hard, R. Neira, K. Paulson, A. Ponniah, W.
    Kamonrat, W. Mwanja, I. Fleming, J. Gallardo, R. Devlin, and J.
    Trisak. Approaches to assessing gene flow. Chapter 5 in
    Environmental Risk Assessment of Genetically Modified Organisms,
    Volume 3: Building Scientific Capacity for Transgenic Fish in
    Developing Countries, CABI Publishing, *accepted.*

2.  Senanan, W., J. Hard, A. Alcivar-Warren, J. Trisak, M.
    Zakaraia-Ismail, and M. L. Hernandez. Risk management: post-approval
    monitoring and remediation. Chapter 9 in Environmental Risk
    Assessment of Genetically Modified Organisms, Volume 3: Building
    Scientific Capacity for Transgenic Fish in Developing Countries,
    CABI Publishing, *accepted.*

3.  Hard, J. J. 2006. Case study of Pacific salmon. *In* U.
    Dieckmann, O. R. Godø, M. Heino, and J. Mork (editors),
    Fisheries-Induced Adaptive Change. Cambridge Studies in Adaptive
    Dynamics, Cambridge University Press, *in press*.

4.  Hard, J. J. 2004. Evolution of chinook salmon life history under
    size-selective harvest. *In* A. Hendry and S. Stearns (editors),
    Evolution Illuminated: Salmon and Their Relatives, p. 315-337,
    Oxford University Press.

5.  Wang, S., J. J. Hard, and F. Utter. 2002. Salmonid inbreeding: a
    review. [Reviews in Fish Biology and Fisheries]{.underline}
    11:301-319.

6.  Wang, S., J. J. Hard, and F. Utter. 2002. Genetic variation and
    fitness in salmonids. [Conservation Genetics]{.underline} 3:321-333.

7.  Hard, J. J., R. G. Kope, and W. S. Grant. 2000. Endangered Species
    Act review of the status of pink salmon from Washington, Oregon, and
    California. *In* E. E. Knudsen, C. R. Steward, D. D.
    MacDonald, J. E. Williams, and D. W. Reiser (editors), Sustainable
    Fisheries Management: Pacific Salmon, p. 103-110, Lewis Publishers,
    Boca Raton, FL.

8.  Hard, J. J. 1995. A quantitative genetic perspective on the
    conservation of intraspecific diversity. [American Fisheries Society
    Symposium]{.underline} 17:304-326.

9.  Hard, J. J. 1995. Genetic monitoring of life-history characters in
    salmon supplementation: problems and opportunities. [American
    Fisheries Society Symposium]{.underline} 15:212-225.

10. Hard, J. J. 1987. All the kings' colors. [Alaska Fish &
    Game]{.underline} 19(4):4-5.

**Conference Proceedings**

1.  Hard, J. J. 2002. Genetic risks of hatchery salmon production to
    wild salmon. *In* C. Orr, P. Gallaugher, and J. Penikett (editors),
    Hatcheries and the protection of wild salmon, p. 72-85. Speaking for
    the Salmon workshop proceedings, Centre for Coastal Studies and
    Continuing Studies in Science at Simon Fraser University, Burnaby,
    B.C.

2.  Hard, J. J. 2002. Update on the status of pink salmon in the Pacific
    Northwest. *In* K. Neely, O. W. Johnson, K. W. Myers, and J. J. Hard
    (rapporteurs), Proceedings 20^th^ Northeast Pacific Pink and Chum
    Workshop, Seattle, Washington.

3.  Neely, K., O. W. Johnson, K. W. Myers, and J. J. Hard
    (rapporteurs). 2002. Proceedings 20^th^ Northeast Pacific Pink and
    Chum Workshop, Seattle, Washington.

4.  Johnson, O. W., A. M. Garrett, W. S. Grant, K. Neely, M. H.
    Ruckelshaus, F. W. Waknitz, and J. J. Hard. 2001. Summary of status
    review of coastal cutthroat trout from the Pacific northwest. Pages
    255-257 in Brewin, M. K., A. J. Paul, and M. Monita (editors). Bull
    trout II conference proceedings. Trout Unlimited Canada, Calgary,
    Alberta.

5.  Hard, J. J. 2000. Genetic interactions between wild and cultured
    salmon. *In* P. Gallaugher and C. Orr (editors), Aquaculture and the
    protection of wild salmon, p. 8-18. Speaking for the Salmon workshop
    proceedings, Continuing Studies in Science at Simon Fraser
    University, Burnaby, B.C.

6.  Hard, J. J. 1999. Status review of pink salmon in the Pacific
    Northwest. *In* S. Hawkins (rapporteur), Proceedings of the 19^th^
    Northeast Pacific Pink and Chum Salmon Workshop, p. 166-173, Juneau,
    Alaska.

**Technical reports**

1.  Hard, J. J., J. M. Myers, F. W. Waknitz, and the members of the
    Biological Review Team. 2005. Status review update for Puget Sound
    steelhead. Report submitted to NOAA/NMFS Northwest Regional Office,
    Seattle, Washington, 26 July 2005, 70 p. + 4 appendices.

2.  Hard, J. J., and K. A. Naish. 2005. Evaluate effects of inbreeding
    and inbreeding depression. In B. A. Berejikian (editor), Research on
    captive broodstock programs for Pacific salmon, p. xx-xx. Annual
    Report to Bonneville Power Administration, Project No. 93-056-00
    (Contract No. 00005227).

3.  Hard, J. J., and K. A. Naish. 2004. Evaluate effects of inbreeding
    and inbreeding depression. In B. A. Berejikian (editor), Research on
    captive broodstock programs for Pacific salmon, p. 164-189. Annual
    Report to Bonneville Power Administration, Project No. 93-056-00
    (Contract No. 00005227).

4.  Peek, J. M., M. S. Boyce, E. O. Garton, J. J. Hard, and L. S.
    Mills. 2002. Risks involved in current management of elk in
    Washington. Final report to Washington Department of Fish and
    Wildlife and Washington Wildlife Commission, Olympia, WA, 99 p.
    October 2002.

5.  Hard, J. J. 2002. Inbreeding. In B. A. Berejikian (editor), Research
    on captive broodstock programs for Pacific salmon, p. 96-110. Annual
    Report to Bonneville Power Administration, Project No. 93-056-00
    (Contract No. 00005227).

6.  Fuss, H. J., C. Sharpe, M. J. Ford, J. J. Hard, and E. LaHood. 2001.
    Differences in natural production between hatchery and wild coho
    salmon in Minter Creek, Washington. Annual Report to the Hatchery
    Scientific Research Group, Seattle, WA, June 2001.

7.  Hard, J. J. 2001. Inbreeding. In B. Berejikian and C. Nash
    (editors), Research on captive broodstock programs for Pacific
    salmon, p. 86-94. Annual Report to Bonneville Power Administration,
    Project No. 93-056-00 (Contract No. 00005227).

8.  Hard, J. J. 2000. Inbreeding. Pages 111-116 *In* B. Berejikian
    (editor), Research on captive broodstock programs for Pacific
    salmon. Report to Bonneville Power Administration, Contract No.
    1999A117859, Project No. 199305600, 124 electronic pages (BPA Report
    DOE/BP-17859-1), June 2000. (Available
    http://www.efw.bpa.gov/Environment/EW/EWP/DOCS/REPORTS/HATCHERY/A17859-1.pdf)

9.  Ford, M. J., and J. J. Hard. 2000. Does traditional hatchery
    production help conserve wild salmon -- a comment on the Fall Creek
    coho hatchery controversy. Unpubl. manuscript, 9 p., available as an
    Adobe Acrobat file from
    [[http://www.nwfsc.noaa.gov/cbd/LannanResponse.pdf]{.underline}](http://www.nwfsc.noaa.gov/cbd/LannanResponse.pdf).

10. Johnson, O. J., M. H. Ruckelshaus, W. S. Grant, F. W. Waknitz, A. M.
    Garrett, G. J. Bryant, K. M. Neely, and J. J. Hard. 1999. Status
    review of coastal cutthroat trout from Washington, Oregon, and
    California. [U.S. Department of Commerce, NOAA Technical
    Memorandum]{.underline} NMFS-NWFSC-37, 292 p.

11. Hard, J. J. 1997. Genetic effects of salmon hatcheries: outbreeding
    depression, artificial selection, and fitness. *In* Proceedings of
    Outbreeding Depression: A Research Planning Workshop. University of
    Alaska, Juneau, 2-5 March.

12. Hard, J. J., R. G. Kope, W. S. Grant, F. W. Waknitz, L. T. Parker,
    and R. S. Waples. 1996. Status review of pink salmon from
    Washington, Oregon, and California. [U.S. Department of Commerce,
    NOAA Technical Memorandum]{.underline} NMFS-NWFSC-25, 131 p.

13. Hard, J. J., and W. K. Hershberger. 1996. Research on quantitative
    genetic consequences of captive broodstock programs for Pacific
    salmon populations. In P. Swanson and 8 coauthors: Research on
    captive broodstock programs for Pacific salmon, Part VIII, p. 8-1 to
    8-28. Annual report to Bonneville Power Administration, Contract No.
    AI79-93BP55064, Proj. No. 93-56.

14. Hard, J. J., and W. K. Hershberger. 1995. Quantitative genetic
    consequences of captive broodstock programs for anadromous Pacific
    salmon (*Oncorhynchus* spp.). *In* T. A. Flagg and C. V. W. Mahnken
    (editors), An assessment of the status of captive broodstock
    technology for Pacific salmon, p. 2-1 to 2-75. Final Report to
    Bonneville Power Administration, Project No. 93-56 (Contract No.
    DE-AI79-93BP55064), June 1995.

15. Hard, J. J. 1994. Genetics and salmon management: expanded summary
    of a panel discussion. *In* L. K. Park, P. Moran, and R. S. Waples
    (editors), Applications of DNA technology to the management of
    Pacific salmon: proceedings of the workshop, p. 151-163. [U.S.
    Department of Commerce, NOAA Technical Memorandum]{.underline}
    NMFS-NWFSC-17, 178 p.

16. Hard, J. J. 1994. Density dependence, ecological carrying capacity,
    and Pacific salmon: a summary report. Memorandum to NMFS Northwest
    Regional Office from NMFS Northwest Fisheries Science Center,
    Seattle, Washington.

17. National Marine Fisheries Service (NMFS). 1993. Interim policy on
    artificial propagation of Pacific salmon under the Endangered
    Species Act. [Federal Register]{.underline} \[Docket No.
    921186-2286; April 5, 1993\] 58(63): 17573-17576

18. Hard, J. J., R. P. Jones, Jr., M. R. Delarm, and R. S. Waples. 1992.
    Pacific salmon and artificial propagation under the Endangered
    Species Act. [U.S. Department of Commerce, NOAA Technical
    Memorandum]{.underline} NMFS-NWFSC-2, 56 p.

19. Hard, J. J. 1986. Flesh color variation in chinook salmon
    (*Oncorhynchus tshawytscha*) at Little Port Walter, southeastern
    Alaska. [U.S. Department of Commerce, NOAA Technical
    Memorandum]{.underline} NMFS F/NWC-109, 30 p.

**Manuscripts in preparation or review**

1.  Gustafson, R. G., R. S. Waples, J. M. Myers, J. J. Hard, G. J.
    Bryant, O. W. Johnson, and L. Weitkamp. Characterizing historical
    Pacific salmon population diversity in the Pacific Northwest. To be
    submitted to [Conservation Biology.]{.underline}

2.  Neely, K. G., K. D. Shearer, J. J. Hard, P. Swanson, and W.
    Dickhoff. Early development, growth, proximate composition, and
    digestibility comparisons between a domesticated stock of coho
    salmon and its parent stock. To be submitted to
    [Aquaculture]{.underline}.

3.  Hard, J. J., J. E. Joyce, F. P. Thrower, and A. C. Wertheimer.
    Analysis of age and size at maturity in southeastern Alaska chinook
    salmon. To be submitted to [Canadian Journal of Fisheries and
    Aquatic Sciences]{.underline}.

4.  Hard, J. J., K. A. Naish, W. K. Hershberger, L. T. Parker, K. Neely,
    and S. Wang. Consequences of inbreeding in chinook salmon,
    *Oncorhynchus tshawytscha*. To be submitted to
    [Aquaculture.]{.underline}

5.  Thrower, F.P., and J. J. Hard. Effects of single-event full sibling
    inbreeding on growth and survival in steelhead. To be submitted to
    [Conservation Genetics.]{.underline}

**PAPERS GIVEN AT CONFERENCES AND SYMPOSIA (since 2000)**

**Invited**

1.  2007 An evolutionary approach to evaluating the consequences of
    selective harvest for salmon life history. Sustainability of the
    Arctic-Yukon-Kuskokwim Salmon Fisheries: What Do We Know About
    Salmon Ecology, Management, and Fisheries? Anchorage, Alaska, 6-9
    February

2.  2006 Variation in growth, precocious maturation, smoltification, and
    marine survival in anadromous and derived freshwater forms of
    southeast Alaskan *Oncorhynchus mykiss*: implications for
    conservation of steelhead. 10^th^ West Coast Steelhead Meeting, Port
    Townsend, Washington, 7-9 March

3.  2005 Genetic variation in disease resistance of chinook salmon
    (*Oncorhynchus tshawytscha*) exposed to two bacterial pathogens.
    Symposium on Bacterial Kidney Disease - Challenge for the 21st
    Century, Seattle, Washington, 15-17 November

4.  2005 Differentiation of wild Alaskan anadromous and derived
    freshwater populations of *Oncorhynchus mykiss* in neutral and
    adaptive traits: implications for conservation and recovery of
    steelhead. Annual meeting of the American Fisheries Society,
    Anchorage, Alaska, 11-15 September

5.  2004 Evaluating the potential for fisheries-induced evolutionary
    change in salmonids. Annual meeting of the American Fisheries
    Society, Madison, Wisconsin, 21-26 August

6.  2004 Potential outbreeding depression between an anadromous
    population of steelhead (*Oncorhynchus mykiss*) and an outplanted
    population partially isolated for 70 years and subjected to founder
    effects. Annual meeting of the American Fisheries Society, Madison,
    Wisconsin, 21-26 August (with F. P. Thrower and J. E. Joyce)

7.  2004 Quantitative genetics and conservation: applying a proven tool
    to emerging problems. Plenary address, workshop on Conservation
    Genetics Workshop on Imperiled Freshwater Molluscs and Fishes,
    Freshwater Mollusc Conservation Society, Shepherdstown, West
    Virginia, June

8.  2002 Constraint and opportunity: the complex role genetic
    architecture plays in adaptation. Keynote address, symposium on
    Genetic Basis, Architecture, and Determinants of Fitness-related
    Traits in Fishes, Congress on Ecological and Evolutionary Ethology
    of Fishes, Quebec City, Quebec, Canada, 16-19 August

9.  2002 Inbreeding and its consequences: what do we know and need to
    know? Workshop on Captive Broodstocks for Recovery of Imperiled
    Salmonid Populations, Gig Harbor, Washington, 25-26 June

10. 2002 Evaluating benefits and risks of hatchery salmon production:
    why the controversy? Salmonid Restoration Conference, Ukiah,
    California, 2-3 March

11. 2001 Some current issues related to the culture of Pacific salmon
    for enhancement. Cultivation of Salmon II Symposium, Bergen, Norway,
    6-10 May (with K. D. Shearer, M. Strom, and W. Fairgreave)

12. 2001 Update on the status of pink salmon in the Pacific Northwest.
    20^th^ Northeast Pacific Pink and Chum Workshop, Seattle,
    Washington, 21-23 March

13. 2001 Evolution of salmon life history under size-selective harvest.
    Symposium on Fisheries-Induced Adaptive Change, International
    Institute for Applied Systems Analysis (IIASA), Laxenburg, Austria,
    14-17 March

14. 2000 Defining units of salmon conservation: the challenge of
    interpreting biological diversity. Symposium on Evolutionary Ecology
    of Pacific Salmon (S-8), Society for Conservation Biology, Missoula,
    Montana, 5-9 June (with P. J. Busby, R. G. Gustafson, O. W.
    Johnson, J. M. Myers, and L. A. Weitkamp)

15. 2000 Consequences of inbreeding for survival and growth of captively
    reared chinook salmon: preliminary results. Fish and Fisheries in
    the Columbia River Basin: The Science Behind the Decisions, Annual
    Meeting of Oregon Chapter, American Fisheries Society, Eugene,
    Oregon, 16-18 February

**Contributed**

16. 2006 Potential trade-offs in resistance of chinook salmon
    (*Oncorhynchus tshawytscha*) to two bacterial pathogens resulting
    from selection of broodstock based on antigen level. International
    Association for Genetics in Aquaculture, Montpellier, France, 26-30
    June (talk accepted)

17. 2004 Heritability of precocious maturation, smolting and growth in
    anadromous and derived freshwater populations of steelhead
    (*Oncorhynchus mykiss*). Annual international conference of the
    Fisheries Society of the British Isles, London, 19-23 July
    (with F. P. Thrower and J. E. Joyce)

18. 2004 No effect on embryo survival or smolt to adult survival at sea
    in second generation of outbreeding of coho salmon. Annual
    international conference of the Fisheries Society of the British
    Isles, London, 19-23 July (with W. W. Smoker, I. A. Wang, and A. J.
    Gharrett)

19. 2004 Quantitative genetic analysis of chinook salmon resistance to
    two bacterial pathogens. Coastwide Genetics Workshop, Newport,
    Oregon, 17-18 June (poster with L. Park, R. Pascho, D. Elliott, D.
    Chase, and D. Campton)

20. 2003 Inheritance of age at maturity in Alaskan chinook salmon under
    size-selective harvest. Annual meeting of the American Fisheries
    Society, Quebec City, Quebec, Canada, 10-14 August

21. 2002 Consequences of size-selective fishing for evolution of chinook
    salmon life history. 20^th^ Lowell Wakefield Fisheries Symposium on
    Genetics of Subpolar fish and Invertebrates, Juneau, Alaska, 29-31
    May

22. 2002 Effects on embryo development time and survival of
    intercrossing three geographically separate stocks of southeast
    Alaska coho salmon (*Oncorhynchus kisutch*). 20^th^ Lowell Wakefield
    Fisheries Symposium on Genetics of Subpolar fish and Invertebrates,
    Juneau, Alaska, 29-31 May (with K. L. Granath, W. W. Smoker,
    and A. J. Gharrett)

23. 2002 Relative fitness of hatchery and natural coho salmon spawning
    in Minter Creek, Washington. 20^th^ Lowell Wakefield Fisheries
    Symposium on Genetics of Subpolar fish and Invertebrates, Juneau,
    Alaska, 29-31 May (with M. Ford, E. LaHood, B. Berejikian, H.
    Fuss, C. Sharpe, and P. Hulett)

**PAPERS GIVEN AT SEMINARS AND WORKSHOPS (since 2000)**

**Invited**

1.  2004 Genetic variation in life history in anadromous and derived
    freshwater forms of *Oncorhynchus mykiss*, Independent Scientific
    Advisory Board, Seattle, Washington, 8 December

2.  2004 Genetic variation in life history in anadromous and derived
    freshwater forms of *Oncorhynchus mykiss*, Recovery Science Review
    Panel, Santa Cruz, California, 1 December

3.  2004 An evolutionary look at size-selective harvest of chinook
    salmon, Independent Scientific Advisory Board Seattle, Washington,
    25 February

4.  2004 Consequences of interbreeding among coho salmon populations:
    preliminary results, Hatchery Scientific Research Group, Seattle,
    Washington, 9 February

5.  2004 An evolutionary look at size-selective harvest of chinook
    salmon, University of Washington, School of Fishery and Aquatic
    Sciences, Seattle, Washington, 5 February

6.  2003 Genetics issues in hatcheries and captive breeding programs,
    University of Washington, School of Fishery and Aquatic Sciences,
    Seattle, Washington, 26 November

7.  2003 Collaborative development of a CALFED proposal to reprogram
    California salmon production hatcheries for research, Sacramento,
    California, 27 March

8.  2002 Genetics issues in hatcheries and captive breeding programs,
    University of Washington, School of Fishery and Aquatic Sciences,
    Conservation Genetics 510 lecture, Seattle, Washington, 27 November

9.  2001 Genetic consequences of elk harvest. Elk Risk Assessment
    Panel - Progress Report. Washington Fish and Wildlife Commission
    Workshop, Olympia, Washington, 6 October

10. 2001 Genetic risks of hatchery salmon production to wild salmon,
    Hatcheries and the Protection of Wild Salmon , Simon Fraser
    University Centre for Coastal Studies, Burnaby, B.C., Canada, 6-8
    June

11. 2001 Quantitative genetics of salmon harvest, University of
    Washington, School of Fishery and Aquatic Sciences, Conservation
    Genetics 510 lecture, Seattle, Washington, 21 May

12. 2001 Evolutionary consequences of size-selective harvest for chinook
    salmon, University of Alaska Fairbanks, School of Fishery and Ocean
    Sciences, Juneau, Alaska, April

13. 2001 Defining units of salmon conservation: the challenge of
    interpreting biological diversity, University of Alaska Fairbanks,
    School of Fishery and Ocean Sciences, Juneau, Alaska, April

14. 2000 Workshop for Recovery and Restoration of East Coast Sturgeons
    in the Neuse and St. John's River Systems, National Ocean Service
    Charleston Laboratory, Ft. Johnson, South Carolina, 26-27 July

15. 2000 Sturgeon Culture Risk Assessment Workshop, Mote Marine
    Laboratory, Sarasota, Florida, 6-7 April

16. 2000 Genetic interactions between wild and cultured salmon,
    Aquaculture and the Protection of Wild Salmon , Simon Fraser
    University Continuing Studies in Science, Burnaby, B.C., Canada, 1-3
    March

**Contributed**

17. 2006 Genetic variation in resistance of chinook salmon to two
    bacterial pathogens. NWFSC Turk Lunch, 24 January

18. 2002 Hatchery Research in Conservation Biology Division.
    Presentation to NWFSC Accreditation Panel, Northwest Fisheries
    Science Center, Seattle, Washington, 16 September

19. 2002 Relative fitness of hatchery and natural coho salmon in Minter
    Creek - Progress report. Hatchery Scientific Research Group,
    Northwest Fisheries Science Center, Seattle, Washington, 16 January

**RESEARCH FUNDING**

2006- Determine how hatchery domestication reduces fitness of naturally
spawning salmon and identify ways to reduce these harmful effects (BiOp:
Strategies and Substrategies; Hatchery Strategy 2), NMFS Biological
Opinion Remand funding, \$1,477,500 awarded

2006- Selective Fishery Impacts on Yukon R. Chinook,
Arctic-Yukon-Kuskokwim Sustainable Salmon Initiative, \$183,005 awarded
(J.F. Bromaghin, USFWS, R. Nielson and L. McDonald, Western EcoSystems
Technology, Inc., co-P.I.s)

2000- Heritability of Disease Resistance and Immune Function in Chinook
Salmon, Bonneville Power Administration FY-2000 Initiative, \$200K
awarded (D. E. Campton, USFWS; D. G. Elliott and R. J. Pascho, USGS,
co-P.I.s)

2000- Differences in Natural Production Between Hatchery and Wild Coho:
Reproductive Competence as Influenced by Degree of Hatchery Ancestry,
Hatchery Scientific Review Group, Interagency Committee for Outdoor
Recreation, \$340K awarded (H. J. Fuss, P. L. Hulett, and C. S. Sharpe,
WDFW; K. P. Currens, NWIFC; and M. J. Ford, NWFSC, co-P.I.s)

1996- Outbreeding Depression in Pacific Salmon, NOAA/NMFS Recover
Protected Species Initiative, Project NWC-P18, \$1.35M awarded

1993- Quantitative Genetic Consequences of Captive Broodstock Programs
for Pacific Salmon, Bonneville Power Administration, Project 93-56,
\$1.65M awarded (K. A. Naish, University of Washington, co-P.I.)

**AWARDS, HONORS, AND FELLOWSHIPS**

2005 U.S. Department of Commerce Special Act Award

2000-02 U.S. Department of Commerce Special Act Awards

1996 U.S. Department of Commerce Group Bronze Medal, Endangered Species
Act Coastwide Status Reviews for Pacific Salmon

1996 U.S. Department of Commerce Group Bronze Medal, Genetics Project

1993 Finalist, Young Investigator\'s Prize, Society of American
Naturalists

1992 National Research Council Post-doctoral Fellowship (award offered)

1990-91 National Institutes of Health Genetics Training Grant
(pre-doctoral fellowship)

1987-88 Coca-Cola Scholarship, University of Oregon

1986-91 Graduate Fellowship, University of Oregon

1986 Finalist, American Fisheries Society Publications Awards for Best
Publication in the [Transactions of the American Fisheries
Society]{.underline}

1981-84 U.S. Department of Commerce Advanced Studies Program

1974-76 Alaska Stateroom Scholarship, University of Alaska

**PROFESSIONAL AFFILIATIONS AND CERTIFICATIONS**

2005- Mentor, NMFS - Sea Grant Joint Graduate Fellowship Program in
Population Dynamics and Marine Resource Economics

2000. National Research Council Post-doctoral Research Advisor

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1998. Affiliate Associate Professor of Fisheries, University of Alaska
      Fairbanks, School of Fisheries and Ocean Sciences

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1996. American Association for the Advancement of Science, Member

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1989. Sigma Xi: The Scientific Research Society, Associate

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1984. American Institute of Fishery Research Biologists, Member

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1982. American Fisheries Society (Certified Fisheries Scientist, 1986;
      President, Genetics Section, 2004-)

**PROFESSIONAL ACTIVITIES**

2006- Steering Committee (with R. Waples, R. Huey, J. Travis), Workshop
on Evolutionary consequences of anthropogenic changes on long-term
viability of Pacific salmon and steelhead, Seattle, Washington (planned,
December 2006)

2006- Co-chair (with O. Johnson, K. Neely, L. Weitkamp), 23^rd^
Northeast Pacific Pink and Chum Salmon Workshop, Seattle, Washington
(planned, February 2007)

2006 Member, Board of Examiners for P.V. Kalavathy, Ph.D. candidate at
Manonmaniam Sundaranar University, India (Dissertation entitled
\"Trophic polymorphism and adaptive plasticity in selected threatened
cyprinid fishes in streams and rivers of south India\")

2005 Invited participant, United Nations Environment Program Global
Environmental Facility, Scientific & Technical Advisory Panel Workshop
on the Environmental Risk Assessment of Transgenic Fish, October 2005,
Penang, Malaysia

2005 Co-lead, Endangered Species Act status review for Puget Sound
steelhead

2005- Northwest Fisheries Science Center Research Planning Team

2004-06 President, Genetics Section, American Fisheries Society

2004 Member, Scientific Panel, Battle Creek Steelhead Supplementation
Plan

2004 Advisory Group, Risk Assessment Modeling Project for Salmon
Hatcheries

2003. Associate Editor, *Transactions of the American Fisheries Society*

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2003. Member, Oregon Hatchery Research Center Science Team and Advisory
      Board

2002- Northwest Fisheries Science Center Promotion Management Advisory
Committee

2002-03 Northwest Fisheries Science Center Safety Committee

2001-04 Northwest Fisheries Science Center Internal Grants Program
Review Panel (co-chair, 2003-04)

2001-02 Member, Review Panel for Elk Harvest Strategies, Washington
Department of Fish and Wildlife and Washington Wildlife Commission

2001 Co-chair, 20^th^ Northeast Pacific Pink and Chum Salmon Workshop,
Seattle, Washington

2000 Panelist, Aquaculture and the Protection of Wild Salmon 'Think
Tank,' Simon Fraser University Continuing Studies in Science, Burnaby,
B.C., Canada

1997 Alaska Sea Grant Annual Review Panelist

1995 Genetics Panelist, Wild Salmon Supplementation Workshop, *Exxon
Valdez* Oil Spill Restoration Program

1994-96 Lead researcher, Endangered Species Act status review for pink
salmon

1994 Expert Panelist, Canadian Salmonid Enhancement Program Evaluation

1994- Chinook Salmon Captive Propagation Technical Oversight Committee

1991 Steering Committee, Twelfth Pacific Ecology Conference

1990-91 President\'s Task Force on Research and Graduate Education,
University of Oregon

**STUDENT SUPERVISION**

2005- Neala Kendall, University of Washington School of Aquatic and
Fishery Sciences (M.S. Advisory committee)

2005- Jessica Dales, Santa Clara University NOAA undergraduate intern,
Environmental Sciences major

2005- Tyler Dann, University of Alaska School of Fisheries and Ocean
Sciences (M.S. Advisory committee)

2005- Jon Drake, University of Washington School of Forest Sciences
(Ph.D. Advisory committee, Advanced Studies Program)

2004- Willy Eldridge, University of Washington School of Aquatic and
Fishery Sciences (Ph.D. Advisory committee); NMFS - Sea Grant Joint
Graduate Fellow in Population Dynamics and Marine Resource Economics

2003-05 Dr. Shaun Roark (Ph.D. Miami University, Ohio), NRC Postdoctoral
Associate

2002. 06 Kathleen Neely, University of Washington School of Aquatic and
      Fishery Sciences (M.S. Advisory committee, Advanced Studies
      Program)

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2002. 06 Stephanie Walden, University of Alaska School of Fisheries and
      Ocean Sciences (M.S. Advisory committee)

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2001. 05 Todd Seamons, University of Washington School of Fisheries
      (Ph.D. advisory committee)

1999-02 Brian Beckman, University of Washington School of Fisheries
(Ph.D. advisory committee)

1998-02 Karla Granath, University of Alaska School of Fisheries and
Ocean Sciences (M.S. Advisory committee)

1998-01 Dr. Kerry Naish (Ph.D. University of Wales, Swansea), NRC Senior
Postdoctoral Associate

1998-01 Shizhen Wang, University of Washington School of Fisheries
(Ph.D. advisory committee)

1996-97 Chris Richardson, University of Washington School of Fisheries
(undergraduate student; mentoring)

**OUTREACH**

2004 Graduate course led: Quantitative Genetics (FISH 510), University
of Washington School of Aquatic and Fishery Sciences, Seattle, Spring
Quarter, March-June

1999 Science Judge, Cedar River Elementary School Science Fair, Tahoma
School District, May

1994. University of Oregon Mentor Program

1987-88 Scientist-In-Residence, Eugene 4J School District, Oregon

**REVIEWS**

**Journals**

[American Fisheries Society Special Symposia]{.underline}, [American
Naturalist]{.underline}, [Canadian Journal of Fisheries and Aquatic
Sciences]{.underline}, [Canadian Journal of Zoology]{.underline},
[Conservation Biology,]{.underline} [Conservation Genetics]{.underline},
[Copeia]{.underline}, [Ecological Applications]{.underline},
[Ecology]{.underline}, [Ecology of Freshwater Fish]{.underline},
[Environmental Biology of Fishes]{.underline}, [Evolution]{.underline},
[Functional Ecology]{.underline}, [Genetics]{.underline},
[Heredity]{.underline}, [Journal of Fish Biology]{.underline}, [Journal
of Heredity]{.underline}, [North American Journal of Fisheries
Management]{.underline}, [Progressive Fish-Culturist]{.underline},
[Transactions of the American Fisheries Society]{.underline}

**Grant proposals and other documents**

Florida Fish and Wildlife Commission Finfish Genetics Policy, Washington
Sea Grant and Alaska Sea Grant research proposals, Hatchery Scientific
Research Group Operational Guidelines, Aquaculture Collaborative
Research and Development Program (Canada), U.S. Department of
Agriculture Small Business Innovation Research Program,
Saltonstall-Kennedy Grant research proposals, Aquanet (Canada), Natural
Sciences and Engineering Research Council of Canada (NSERC), USGS Gila
Trout Broodstock Management Plan, NWFSC Internal Grants Program

***References available upon request***